Phengaris Atroguttata And Phengaris Xiushani Habitat
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''Phengaris'' is a
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of
gossamer-winged butterflies Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfl ...
in the
subfamily In biological classification, a subfamily (Latin: ', plural ') is an auxiliary (intermediate) taxonomic rank, next below family but more inclusive than genus. Standard nomenclature rules end botanical subfamily names with "-oideae", and zo ...
Polyommatinae Polyommatinae, the blues, are a diverse subfamily of gossamer-winged butterflies (family Lycaenidae). This subfamily of butterfly was discovered in 1775. These butterflies have been considered one of the poorest studied groups within the family Ly ...
. Commonly, these butterflies are called large blues, which if referring to a particular
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
is '' P. arion'', a species resident in Europe and some parts of Asia. ''Phengaris'' is currently defined to include the genus ''Maculinea''. The
type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...
of the latter was the
Alcon blue ''Phengaris alcon'', the Alcon blue or Alcon large blue, is a butterfly of the family Lycaenidae and is found in Europe and across the Palearctic to Siberia and Mongolia. Description from Seitz ''L. alcon'' Schiff. (= areas Esp., ''euphemus' ...
(''"M." alcon''). This species was found to be less closely related to most other supposed members of ''Maculinea'' than the traditional ''Phengaris'' species, and hence the two genera were merged to form a
monophyletic In biological cladistics for the classification of organisms, monophyly is the condition of a taxonomic grouping being a clade – that is, a grouping of organisms which meets these criteria: # the grouping contains its own most recent co ...
group. Alternatively, ''Maculinea'' could be restricted to the one or two species of "Alcon" blues, and the rest of it be separated as a new genus. But this would create two very small genera, which is generally avoided by modern
taxonomist In biology, taxonomy () is the science, scientific study of naming, defining (Circumscription (taxonomy), circumscribing) and classifying groups of biological organisms based on shared characteristics. Organisms are grouped into taxon, taxa (si ...
s. As ''Phengaris'' is the older name, it thus replaces ''Maculinea''.Brower (2008) ''Maculinea'' was alternatively considered to be a
subgenus In biology, a subgenus ( subgenera) is a taxonomic rank directly below genus. In the International Code of Zoological Nomenclature, a subgeneric name can be used independently or included in a species name, in parentheses, placed between the ge ...
of '' Glaucopsyche''. But while ''Glaucopsyche'' and ''Phengaris'' (including ''Maculinea'') are certainly close relatives, they are unlikely to be that close. The most recently discovered species in the genus is '' P. xiushani'', first reported in 2010 and found in undisturbed forested mountains, unlike all the other members which live in
grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...
s over their entire range of distribution.


Species

The species, listed in the presumed
phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...
sequence from the most ancestral to the most
apomorphic In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to hav ...
, are:See Brower (2008) and the references in Savela (2006, 2008) Basal group (''Maculinea
sensu stricto ''Sensu'' is a Latin word meaning "in the sense of". It is used in a number of fields including biology, geology, linguistics, semiotics, and law. Commonly it refers to how strictly or loosely an expression is used in describing any particular c ...
'') * ''
Phengaris alcon ''Phengaris alcon'', the Alcon blue or Alcon large blue, is a butterfly of the family Lycaenidae and is found in Europe and across the Palearctic to Siberia and Mongolia. Description from Seitz ''L. alcon'' Schiff. (= areas Esp., ''euphemus' ...
'' – Alcon blue, Alcon large blue ** †''
Phengaris alcon arenaria The Dutch alcon blue (''Phengaris alcon arenaria'') was a subspecies of the alcon blue butterfly ('' Phengaris alcon''). There is not much known about this subspecies, but it has always been very rare. It was endemic to the Netherlands, where two ...
'' – Dutch Alcon blue * ''
Phengaris rebeli ''Phengaris rebeli'' (formerly ''Maculinea rebeli''), common name mountain Alcon blue, is a species of butterfly in the family Lycaenidae. It was first found and described in Styria, Austria, on Mount Hochschwab around 1700. Although it was init ...
'' – mountain Alcon blue (may belong in ''P. alcon'') ''Phengaris sensu stricto'' * ''
Phengaris albida ''Phengaris albida'' is a species of butterfly of the family Lycaenidae. It is only known from western China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population excee ...
'' (Leech, 1893) * ''
Phengaris atroguttata ''Phengaris atroguttata'', the great spotted blue, Retrieved April 23, 2018. is a small butterfly found in India, Myanmar, and China that belongs to the lycaenids or blues family. Taxonomy The butterfly was earlier known as ''Polyommatus atro ...
'' – great spotted blue * ''
Phengaris daitozana ''Phengaris'' is a genus of gossamer-winged butterflies in the subfamily Polyommatinae. Commonly, these butterflies are called large blues, which if referring to a particular species is '' P. arion'', a species resident in Europe and some parts o ...
'' (Wileman, 1908) * ''
Phengaris xiushani ''Phengaris xiushani'' is a species of butterfly of the family Lycaenidae. It is only known from Yunnan in China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population e ...
'' Wang & Settele, 2010 – Xiushan's large blue Other ''"Maculinea"'' * ''
Phengaris arion The large blue (''Phengaris arion'') is a species of butterfly in the family Lycaenidae. The species was first defined in 1758 and first recorded in Britain in 1795. In 1979 the species became mostly extinct in Britain but has been successfully ...
'' – large blue * ''
Phengaris arionides The greater large blue (''Phengaris arionides'') is a species of butterfly in the family Lycaenidae. It is found in Northeast China, Japan, and the Russian Far East The Russian Far East ( rus, Дальний Восток России, p ...
'' – greater large blue * ''
Phengaris cyanecula ''Phengaris'' is a genus of gossamer-winged butterflies in the subfamily Polyommatinae. Commonly, these butterflies are called large blues, which if referring to a particular species is ''Large blue, P. arion'', a species resident in Europe and s ...
'' (Eversmann, 1848) ** ''Phengaris cyanecula cyanecula''
Mongolia Mongolia is a landlocked country in East Asia, bordered by Russia to the north and China to the south and southeast. It covers an area of , with a population of 3.5 million, making it the world's List of countries and dependencies by po ...
** ''Phengaris cyanecula obscurior'' (Staudinger, 1901)
Tian-Shan The Tian Shan, also known as the Tengri Tagh or Tengir-Too, meaning the "Mountains of God/Heaven", is a large system of mountain ranges in Central Asia. The highest peak is Jengish Chokusu at high and located in Kyrgyzstan. Its lowest point is ...
** ''Phengaris cyanecula taras'' (Fruhstorfer, 1915)
Alay Mountains The Alay or Alai Range (; ) is a mountain range that extends from the Tien Shan mountain range in Kyrgyzstan west into Tajikistan. It is part of the Pamir-Alay mountain system. The range runs approximately east to west. Its highest summit is Pik ...
** ''Phengaris cyanecula ussuriensis'' (Sheljuzhko, 1928)
Amur Oblast Amur Oblast () is a federal subject of Russia (an oblast), located on the banks of the Amur and Zeya rivers in the Russian Far East. The oblast borders Heilongjiang province of the People's Republic of China (PRC) to the south. The administrati ...
,
Ussuri The Ussuri ( ; ) or Wusuli ( ) is a river that runs through Khabarovsk and Primorsky Krais, Russia and the southeast region of Northeast China in the province of Heilongjiang. It rises in the Sikhote-Alin mountain range, flowing north and formi ...
* '' Phengaris kurentzovi'' (Sibatani, Saigusa & Hirowatari, 1994) ** ''Phengaris kurentzovi kurentzovi'' northern
China China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
and
Korea Korea is a peninsular region in East Asia consisting of the Korean Peninsula, Jeju Island, and smaller islands. Since the end of World War II in 1945, it has been politically Division of Korea, divided at or near the 38th parallel north, 3 ...
** ''Phengaris kurentzovi daurica'' Dubatolov, 1999
Transbaikalia Transbaikal, Trans-Baikal, Transbaikalia ( rus, Забайка́лье, r=Zabaykal'ye, p=zəbɐjˈkalʲjɪ), or Dauria (, ''Dauriya'') is a mountainous region to the east of or "beyond" (trans-) Lake Baikal at the south side of the eastern Si ...
* '' Phengaris ligurica'' (Wagner, 1904) (may belong in ''P. arion'') * '' Phengaris nausithous'' – dusky large blue * '' Phengaris ogumae'' (Matsumura, 1910) (tentatively placed here) ** ''Phengaris ogumae ogumae''
Japan Japan is an island country in East Asia. Located in the Pacific Ocean off the northeast coast of the Asia, Asian mainland, it is bordered on the west by the Sea of Japan and extends from the Sea of Okhotsk in the north to the East China Sea ...
** ''Phengaris ogumae doii'' (Matsumura, 1928)
Kuriles The Kuril Islands or Kurile Islands are a volcanic archipelago administered as part of Sakhalin Oblast in the Russian Far East. The islands stretch approximately northeast from Hokkaido in Japan to Kamchatka Peninsula in Russia, separating the ...
* '' Phengaris takamukui'' * '' Phengaris teleius'' – scarce large blue ''
Incertae sedis or is a term used for a taxonomy (biology), taxonomic group where its broader relationships are unknown or undefined. Alternatively, such groups are frequently referred to as "enigmatic taxa". In the system of open nomenclature, uncertainty ...
'' * '' Phengaris xiaheana'' (Murayama, 1919)


Footnotes


References

* (2008): Tree of Life Web Project &ndash
''Phengaris''
Version of 2008-JAN-06. Retrieved 2009-OCT-04. * (2006): Markku Savela's Lepidoptera and some other life forms &ndash

Version of 2006-MAR-12. Retrieved 2009-OCT-04. * (2008): Markku Savela's Lepidoptera and some other life forms &ndash

Version of 2008-JUL-05. Retrieved 2009-OCT-04.


External links


Images representing ''Phengaris''
at
Consortium for the Barcode of Life The Consortium for the Barcode of Life (CBOL) was an international initiative dedicated to supporting the development of DNA barcoding as a global standard for species identification. CBOL's Secretariat Office is hosted by the National Museum of ...
{{Taxonbar, from=Q48406 Lycaenidae genera Taxa named by William Doherty