Phaethornis Longirostris
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The long-billed hermit (''Phaethornis longirostris'') is a
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the family
Trochilidae Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
, the
hummingbird Hummingbirds are birds native to the Americas and comprise the Family (biology), biological family Trochilidae. With approximately 366 species and 113 genus, genera, they occur from Alaska to Tierra del Fuego, but most species are found in Cen ...
s. It is found from central
Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...
south through
Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...
,
Colombia Colombia, officially the Republic of Colombia, is a country primarily located in South America with Insular region of Colombia, insular regions in North America. The Colombian mainland is bordered by the Caribbean Sea to the north, Venezuel ...
and
Ecuador Ecuador, officially the Republic of Ecuador, is a country in northwestern South America, bordered by Colombia on the north, Peru on the east and south, and the Pacific Ocean on the west. It also includes the Galápagos Province which contain ...
into
Peru Peru, officially the Republic of Peru, is a country in western South America. It is bordered in the north by Ecuador and Colombia, in the east by Brazil, in the southeast by Bolivia, in the south by Chile, and in the south and west by the Pac ...
.


Taxonomy and systematics

It has often been considered
conspecific Biological specificity is the tendency of a characteristic such as a behavior or a biochemical variation to occur in a particular species. Biochemist Linus Pauling stated that "Biological specificity is the set of characteristics of living organism ...
with what is now the long-tailed hermit, ''P. superciliosus'', which is found east of the
Andes The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the List of longest mountain chains on Earth, longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range ...
. The two populations were called the western and eastern long-tailed hermits; the "eastern" was dropped after the split into two species and the renaming of the western population.Remsen, J. V., Jr., J. I. Areta, E. Bonaccorso, S. Claramunt, A. Jaramillo, D. F. Lane, J. F. Pacheco, M. B. Robbins, F. G. Stiles, and K. J. Zimmer. Version 24 August 2021. A classification of the bird species of South America. American Ornithological Society. https://www.museum.lsu.edu/~Remsen/SACCBaseline.htm retrieved August 24, 2021 The South American Classification Committee (SACC) of the
American Ornithological Society The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
, the
International Ornithological Committee The International Ornithologists' Union (IOU) is an international organization for the promotion of ornithology. It links basic and applied research and nurtures education and outreach activities. Specifically, the IOU organizes and funds global co ...
(IOC), and the
Clements taxonomy ''The Clements Checklist of Birds of the World'' is a book by Jim Clements which presents a list of the bird species of the world. The most recent printed version is the sixth edition (2007), but has been updated yearly, the last version in 202 ...
assign these four subspecies to the long-billed hermit:Clements, J. F., T. S. Schulenberg, M. J. Iliff, S. M. Billerman, T. A. Fredericks, J. A. Gerbracht, D. Lepage, B. L. Sullivan, and C. L. Wood. 2021. The eBird/Clements checklist of Birds of the World: v2021. Downloaded from https://www.birds.cornell.edu/clementschecklist/download/ Retrieved August 25, 2021 *''P. l. longirostris'' Delattre (1843) *''P. l. cephalus'' Bourcier & Mulsant (1848) *''P. l. susurrus''
Bangs Bang, bang!, or bangs may refer to: Products * M1922 Bang rifle, a US semi-automatic rifle designed by Søren Hansen Bang * Bang, a List of model car brands, model car brand * Bang (beverage), an energy drink Geography * Bang, Central African ...
(1901)
*''P. l. baroni''
Hartert Ernst Johann Otto Hartert (29 October 1859 – 11 November 1933) was a widely published German people, German ornithologist. Life and career Hartert was born in the Free and Hanseatic City of Hamburg on 29 October 1859. In July 1891, he married ...
(1897)
However,
BirdLife International BirdLife International is a global partnership of non-governmental organizations that strives to conserve birds and their habitats. BirdLife International's priorities include preventing extinction of bird species, identifying and safeguarding i ...
's ''
Handbook of the Birds of the World The ''Handbook of the Birds of the World'' (HBW) is a multi-volume series produced by the Spanish publishing house Lynx Edicions in partnership with BirdLife International. It is the first handbook to cover every known living species of bird. ...
'' (HBW) treats ''P. l. baroni'' as a separate species, "Ecuadorian hermit".HBW and BirdLife International (2020) ''Handbook of the Birds of the World and BirdLife International digital checklist of the birds of the world'' Version 5. Available at: http://datazone.birdlife.org/userfiles/file/Species/Taxonomy/HBW-BirdLife_Checklist_v5_Dec20.zip xls zipped 1 MBretrieved May 27, 2021 The SACC in 2006 had rejected that treatment. Other subspecies within ''longirostrus'' and ''cephalus'' have been proposed but have not been accepted.del Hoyo, J., C. Hinkelmann, N. Collar, P. F. D. Boesman, and G. M. Kirwan (2020). Long-billed Hermit (''Phaethornis longirostris''), version 1.0. In Birds of the World (S. M. Billerman, B. K. Keeney, P. G. Rodewald, and T. S. Schulenberg, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.lobher.01 retrieved December 9, 2021


Description

The long-billed hermit is long. Males weigh and females . The nominate subspecies ''P. l. longirostris'' has greenish-brown upperparts and brownish to grayish underparts. Its uppertail
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
have dark and light ochre bands and the central tail feathers are long with long white tips. It has a long decurved bill, with the female's being shorter but more curved than the male's. ''P. l. cephalus'' is similar but its underparts become more ochraceous towards the southern part of its range. ''P. l. susurrus'' is larger than ''cephalus'' but similarly colored. ''P. l. baroni''s upperparts are dull metallic green and its belly is whiter than those of the other subspecies.


Distribution and habitat

The subspecies of long-billed hermit are found thus: *''P. l. longirostris'', from Mexico's
Oaxaca Oaxaca, officially the Free and Sovereign State of Oaxaca, is one of the 32 states that compose the political divisions of Mexico, Federative Entities of the Mexico, United Mexican States. It is divided into municipalities of Oaxaca, 570 munici ...
and
Chiapas Chiapas, officially the Free and Sovereign State of Chiapas, is one of the states that make up the Political divisions of Mexico, 32 federal entities of Mexico. It comprises Municipalities of Chiapas, 124 municipalities and its capital and large ...
states south through
Guatemala Guatemala, officially the Republic of Guatemala, is a country in Central America. It is bordered to the north and west by Mexico, to the northeast by Belize, to the east by Honduras, and to the southeast by El Salvador. It is hydrologically b ...
into northern
Honduras Honduras, officially the Republic of Honduras, is a country in Central America. It is bordered to the west by Guatemala, to the southwest by El Salvador, to the southeast by Nicaragua, to the south by the Pacific Ocean at the Gulf of Fonseca, ...
*''P. l. cephalus'', from eastern Honduras through
Nicaragua Nicaragua, officially the Republic of Nicaragua, is the geographically largest Sovereign state, country in Central America, comprising . With a population of 7,142,529 as of 2024, it is the third-most populous country in Central America aft ...
,
Costa Rica Costa Rica, officially the Republic of Costa Rica, is a country in Central America. It borders Nicaragua to the north, the Caribbean Sea to the northeast, Panama to the southeast, and the Pacific Ocean to the southwest, as well as Maritime bo ...
, and
Panama Panama, officially the Republic of Panama, is a country in Latin America at the southern end of Central America, bordering South America. It is bordered by Costa Rica to the west, Colombia to the southeast, the Caribbean Sea to the north, and ...
into northwestern Colombia as far south as
Santander Department Santander () is a department of Colombia. Santander inherited the name of one of the nine original states of the United States of Colombia. It is located in the central northern part of the country, borders the Magdalena River to the east, Bo ...
*''P. l. susurrus'', the isolated
Sierra Nevada de Santa Marta The Sierra Nevada de Santa Marta (English: ''Snow-Covered Mountain Range of Saint Martha'') is an isolated mountain range in northern Colombia, separate from the Andes range that runs through the north of the country. Reaching an elevation of ...
in northeastern Colombia *''P. l. baroni'', from Ecuador's western
Esmeraldas Province Esmeraldas () is a northwestern coastal province of Ecuador. The capital and largest city is Esmeraldas. Esmeraldas is one of the three provinces of Ecuador that borders Colombia, and it is the most northern province in the country. The provin ...
south into Peru's Tumbes and
Piura Piura is a city in northwestern Peru, located north of the Sechura Desert along the Piura River. It is the capital of the Piura Region and the Piura Province. Its population was 484,475 as of 2017 and it is the 7th most populous city in Peru. ...
departments. The species inhabits the understory and edges of a variety of landscapes including
rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...
, tall
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
, humid
semi-deciduous Semi-deciduous or semi-evergreen is a botanical term which refers to plants that lose their foliage for a very short period, when old leaves fall off and new foliage growth is starting. This phenomenon occurs in tropical and sub-tropical wood ...
forest,
cloudforest A cloud forest, also called a water forest, primas forest, or tropical montane cloud forest, is a generally tropical or subtropical, evergreen, montane, moist forest characterized by a persistent, frequent or seasonal low-level cloud cover, ...
, and
gallery forest A gallery forest is one formed as a corridor along rivers or wetlands, projecting into landscapes that are otherwise only sparsely treed such as savannas, grasslands, or deserts. The gallery forest maintains a more temperate microclimate above th ...
. It has been recorded from sea level to in Costa Rica, to in eastern Mexico, to in northern Colombia, to in Peru, and to at least and possibly to in Ecuador.


Behavior


Movement

The long-billed hermit is not known to make large-scale movements but is thought to wander short distances.


Feeding

The long-billed hermit is a " trap-line" feeder like other hermit hummingbirds, visiting a circuit of a variety of flowering plants for nectar. Examples include ''
Heliconia ''Heliconia'' is a genus of flowering plants in the monotypic family Heliconiaceae. Most of the 194 known species are native to the tropical Americas, but a few are indigenous to certain islands of the western Pacific and Maluku (province), ...
'', '' Costus'', '' Aphelandra'', and ''
Passiflora ''Passiflora'', known also as the passion flowers or passion vines, is a genus of about 550 species of flowering plants, the type genus of the family Passifloraceae. ''Passiflora'' species are widely cultivated for their striking flowers, fla ...
''. It also consumes small
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s.


Breeding

The long-billed hermit's breeding seasons vary considerably throughout its range, from April to July in Mexico, May to September in Panama, and January to April or May in Colombia. The nest is a cone-shaped cup made of plant fibers and spider silk suspended from the underside of a drooping leaf. The clutch size is two eggs; The incubation period is 17 to 18 days with
fledging Fledging is the stage in a flying animal's life between hatching or birth and becoming capable of flight. This term is most frequently applied to birds, but is also used for bats. For altricial birds, those that spend more time in vulnerable c ...
22 to 23 days after hatch. During the breeding season, male long-billed hermits sing in communal leks of up to 25 birds, and also wiggle their long tails in display. Competitive lek singing can occupy half of the daylight hours to attract females. In addition, mature males have a longer
bill Bill(s) may refer to: Common meanings * Banknote, paper cash (especially in the United States) * Bill (law), a proposed law put before a legislature * Invoice, commercial document issued by a seller to a buyer * Bill, a bird or animal's beak Pl ...
than females, and appear to use its
dagger A dagger is a fighting knife with a very sharp point and usually one or two sharp edges, typically designed or capable of being used as a cutting or stabbing, thrusting weapon.State v. Martin, 633 S.W.2d 80 (Mo. 1982): This is the dictionary or ...
-like tip as a secondary sexual trait to defend against other males at the lek.


Vocalization

The songs of the three northern subspecies of long-billed hermit are "a continuous series of single, piercing, usually upslurred 'sweeup' notes". Their call is "a sharp, explosive 'week!'" that is often given in flight. The song of ''P. l. baroni'' is "a continuous series of single, sparrow-like, chipping notes...'tchee..tchee..tchee..'." Its call is "a thin 'seep'" also given in flight.


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
follows HBW taxonomy, and so has separately assessed the long-billed hermit and "Ecuadorian hermit" as being of Least Concern. The population sizes and trends of both are unknown. ''P. l. susurrus'' has a small range that potentially places it at risk. ''P. l. cephalus'' occurs in many protected areas in Costa Rica. ''P. l. baroni'' is considered fairly common throughout its range and occurs in some protected areas.


References


External links


Article
AOCR


Further reading

* Hilty, '' Birds of Venezuela'', * Stiles and Skutch, ''A guide to the birds of Costa Rica'' {{Taxonbar, from=Q1055821
long-billed hermit The long-billed hermit (''Phaethornis longirostris'') is a bird in the family Trochilidae, the hummingbirds. It is found from central Mexico south through Central America, Colombia and Ecuador into Peru. Taxonomy and systematics It has often ...
Hummingbird species of Central America Birds of Mexico Birds of Belize Birds of Guatemala Birds of Honduras Birds of Nicaragua Birds of Costa Rica Birds of Panama Birds of Colombia Hummingbird species of South America long-billed hermit'