The Percidae are a

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

ray-finned fish Actinopterygii (; ), members of which are known as ray-finned fish or actinopterygians, is a class of bony fish that comprise over 50% of living vertebrate species. They are so called because of their lightly built fins made of webbings of sk ...

, part of the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Perciformes Perciformes (), also called the Acanthopteri, is an order or superorder of ray-finned fish in the clade Percomorpha. ''Perciformes'' means " perch-like". Among the well-known members of this group are perches and darters ( Percidae), and als ...

, which are found in fresh and

brackish Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...

waters of the

Northern Hemisphere The Northern Hemisphere is the half of Earth that is north of the equator. For other planets in the Solar System, north is defined by humans as being in the same celestial sphere, celestial hemisphere relative to the invariable plane of the Solar ...

. The majority are

Nearctic The Nearctic realm is one of the eight biogeographic realms constituting the Earth's land surface. The Nearctic realm covers most of North America, including Greenland, Central Florida, and the highlands of Mexico. The parts of North America ...

, but there are also

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

species. The family contains nearly 250 species in 12 genera. The perches and their relatives are in this family; well-known species include the

walleye The walleye (''Sander vitreus'', Synonym (taxonomy), synonym ''Stizostedion vitreum''), also called the walleyed pike, yellow pike, yellow pikeperch or yellow pickerel, is a freshwater perciform fish native to most of Canada and to the Northern ...

, sauger, ruffe, and three species of

perch Perch is a common name for freshwater fish from the genus ''Perca'', which belongs to the family Percidae of the large order Perciformes. The name comes from , meaning the type species of this genus, the European perch (''P. fluviatilis'') ...

. However, small fish known as darters are also a part of this family. Despite the widespread and speciose nature of this family, the vast majority of its species diversity is within the darters, which are restricted to eastern North America, and the majority of which have small ranges. Excluding the darters, this family contains only 20 species. In Europe, percids were likely a more ecologically dominant group in the past than now, with the immigration of

cyprinids Cyprinidae is a family of freshwater fish commonly called the carp or minnow family, including the carps, the true minnows, and their relatives the barbs and barbels, among others. Cyprinidae is the largest and most diverse fish family, and t ...

to Europe during the

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

causing a decline in their importance.

Characteristics

The family is characterised by having the

dorsal fin A dorsal fin is a fin on the back of most marine and freshwater vertebrates. Dorsal fins have evolved independently several times through convergent evolution adapting to marine environments, so the fins are not all homologous. They are found ...

split into two which are normally separated or have a narrow connection, although this is wider in the genus ''Zingel'', the front section contains the spines and the rear section contains the soft rays. The

anal fin Fins are moving appendages protruding from the body of fish that interact with water to generate thrust and help the fish swim. Apart from the tail or caudal fin, fish fins have no direct connection with the back bone and are supported o ...

contains 1 or 2 spines, if there is a second spine it is typically weak. The

pelvic fins Pelvic fins or ventral fins are paired fins located on the ventral (belly) surface of fish, and are the lower of the only two sets of paired fins (the other being the laterally positioned pectoral fins). The pelvic fins are homologous to the hi ...

are placed on the

thorax The thorax (: thoraces or thoraxes) or chest is a part of the anatomy of mammals and other tetrapod animals located between the neck and the abdomen. In insects, crustaceans, and the extinct trilobites, the thorax is one of the three main di ...

and have a single spine and 5 soft rays. They also have skeletal

synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ...

. The maximum size attained is in the

zander The zander (''Sander lucioperca''), sander or pikeperch, is a species of ray-finned fish from the Family (biology), family Percidae, which also includes perch, Gymnocephalus, ruffe and Darter (fish), darter. It is found in freshwater and brackis ...

(''Sander lucioperca'') but most of the species in the family are much smaller. Their scales are ctenoid and their bodies are normally somewhat elongate.

Systematics

The following classification is based on ''

Eschmeyer's Catalog of Fishes Catalog of Fishes is a comprehensive on-line database and reference work on the scientific names of fish species and genera. It is global in its scope and is hosted by the California Academy of Sciences. It has been compiled and is continuously up ...

'' (2025): * Subfamily Percinae Rafinesque, 1815 ** Genus '' Gymnocephalus''

Bloch Bloch is a surname of German origin. Notable people with this surname include: A *Adele Bloch-Bauer (1881–1925), Austrian entrepreneur *Albert Bloch (1882–1961), American painter *Alexandre Bloch (1857–1919), French painter *Alfred Bloch ( ...

, 1793 ** Genus '' Perca''

Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

, 1758 ** Genus ''

Percarina ''Percarina'' is a genus of ray-finned fish in the family Percidae found in eastern Europe. They are characterised by having the first dorsal fin, having 9–11 spines and being widely separated from the second dorsal fin. Previously, they were ...

'' Nordmann, 1840 * Subfamily Luciopercinae

Jordan Jordan, officially the Hashemite Kingdom of Jordan, is a country in the Southern Levant region of West Asia. Jordan is bordered by Syria to the north, Iraq to the east, Saudi Arabia to the south, and Israel and the occupied Palestinian ter ...

& Evermann, 1896 ** Tribe Luciopercini Jordan & Evermann 1896 *** Genus ''

Sander A sander is a power tool used to smooth surfaces by abrasion with sandpaper. Sanders have a means to attach the sandpaper and a mechanism to move it rapidly contained within a housing with means to handhold it or fix it to a workbench. Woodw ...

'' Oken, 1817 *** Genus †'' Leobergia'' Schtylko, 1934 (

Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...

of eastern Europe) ** Tribe Romanichthyini Dumitrescu, Bănărescu & Stoica 1957 *** Genus '' Romanichthys'' Dumitrescu, Bănărescu & Stoica 1957 *** Genus '' Zingel'' Cloquet, 1817 * Subfamily Etheostomatinae

Agassiz Jean Louis Rodolphe Agassiz ( ; ) FRS (For) FRSE (May 28, 1807 – December 14, 1873) was a Swiss-born American biologist and geologist who is recognized as a scholar of Earth's natural history. Spending his early life in Switzerland, he recei ...

, 1850 ** Genus '' Ammocrypta'' Jordan, 1877 ** Genus '' Etheostoma'' Rafinesque, 1817 ** Genus '' Nothonotus'' Putnam, 1863 ** Genus '' Percina'' Haldeman, 1842 Under a former treatment by the 5th Edition of ''

Fishes of the World ''Fishes of the World'' is a standard reference for the systematics of fishes. It was first written in 1976 by the American ichthyologist Joseph S. Nelson (1937–2011). Now in its fifth edition (2016), the work is a comprehensive overview of t ...

'', ''Gymnocephalus'' was placed in its own subfamily, Acerinae, as was ''Percarina'' in Percarininae. Although the family likely underwent an evolutionary radiation during the

Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...

, the earliest definitive fossil remains of percids are of modern ''

'' from the

Middle Miocene The Middle Miocene is a sub-epoch of the Miocene epoch (geology), epoch made up of two Stage (stratigraphy), stages: the Langhian and Serravallian stages. The Middle Miocene is preceded by the Early Miocene. The sub-epoch lasted from 15.97 ± 0. ...

of Canada. †'' Vixperca'' Peña Zarzuelo, 1991 from the

Middle Eocene The Eocene ( ) is a geological epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period in the modern Cenozoic Era. The name ''Eocene'' comes from the Ancient Greek (''Ēṓs'', ' Dawn') a ...

of Spain has been suggested to be the earliest known percid, but its remains are fragmentary and show closer similarities to lates perches than to true perches.

References

{{Authority control Perciformes families Taxa named by Constantine Samuel Rafinesque