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The Orleanian age is a period of
geologic time The geologic time scale, or geological time scale, (GTS) is a representation of time based on the rock record of Earth. It is a system of chronological dating that uses chronostratigraphy (the process of relating strata to time) and geochron ...
( MN 3–5, (mya)), within the
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
and used more specifically with
European Land Mammal Ages The European Land Mammal Mega Zones (abbreviation: ELMMZ, more commonly known as European land mammal ages or ELMA) are zones in rock layers that have a specific assemblage of fossils (biozones) based on occurrences of fossil assemblages of Europe ...
. It precedes the
Astaracian The Astaracian age is a period of geologic time (), equivalent with the Middle Miocene and used more specifically with European Land Mammal Ages. It precedes the Vallesian age and follows the Orleanian age. The Astaracian overlaps the Langhian and ...
age and follows the
Agenian The Agenian age is a period of geologic time (23.8–20 Ma) within the Miocene used more specifically with European Land Mammal Ages. It follows the Orleanian age and overlaps the Aquitanian and Burdigalian The Burdigalian is, in the geologic ...
age. ; Agenian-Early Orleanian migration At the time for
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but the ...
-
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recent" ...
boundary (23.8 mya), the Tethys Seaway, a deep through between the
Arabian The Arabian Peninsula, (; ar, شِبْهُ الْجَزِيرَةِ الْعَرَبِيَّة, , "Arabian Peninsula" or , , "Island of the Arabs") or Arabia, is a peninsula of Western Asia, situated northeast of Africa on the Arabian Plate ...
and
Iranian Plate The Iranian Plate is a small tectonic plate thought to underlie the Persian plateau, covering the modern-day countries of Iran and Afghanistan, and parts of Iraq and Pakistan. It is compressed between the Arabian Plate to the southwest, the E ...
s, isolated Africa from Eurasia; an isolation that lasted until the gradual closure of the Tethys at the end of MN 3 (19-18 mya), and America was connected to Asia by the Beringian landbridge. The horse-like
equid Equidae (sometimes known as the horse family) is the taxonomic family of horses and related animals, including the extant horses, asses, and zebras, and many other species known only from fossils. All extant species are in the genus '' Equus'', w ...
''
Anchitherium ''Anchitherium'' (meaning ''near beast'') was a fossil horse with a three- toed hoof. ''Anchitherium'' was a browsing (leaf eating) horse that originated in the early Miocene of North America and subsequently dispersed to Europe and Asia ...
'' dispersed into Asia over the latter and is known on the Iberian Peninsula from MN 3 and from Asia Minor in MN 6. It has not been found on the Balkan Peninsula, which suggests that the Balkans were isolated from Asia Minor at that time. In Western Europe, the pig-like suids '' Hyotherium'' appears in MN 1 and '' Xenohyus'' in MN 2. The hippo-like anthracothere '' Brachyodus'' appears in Western Europe in MN 3 and, while the MN 1-3 fauna is rare in the Eastern Mediterranean, also in Greece from around MN 3-4. The spalacid (rodent) '' Debruijnia'' is known from Asia Minor from MN 3 and from
Aliveri Aliveri ( el, Αλιβέρι) is a town and a community in the island of Evvoia, Greece. It is part of the municipal unit Tamyneoi, and the seat of the municipality Kymi-Aliveri. Aliveri is situated in the western part of the island, on the S ...
, Greece, from early MN 4, suggesting the presence of some kind of early connection between Asia Minor and the Balkans. ; Middle Orleanian migration The connection between Eurasia and Africa, known as the " Gomphotherium land bridge", was re-established at the end of MN 3. However, temporary landbridges appears to have connected Africa to Eurasia before the final closure of the Tethys Seaway, allowing some mammals to emigrate between the landmasses. With the Balkans solidly connected to Asia Minor, the seaways to the
Paratethys The Paratethys sea, Paratethys ocean, Paratethys realm or just Paratethys was a large shallow inland sea that stretched from the region north of the Alps over Central Europe to the Aral Sea in Central Asia. Paratethys was peculiar due to its p ...
were closed, which transformed the shallow sea into an isolated basin with its own endemic fauna. The ''Gomphotherium''-landbridge provided the African hyrocoid '' Pliohyrax'' with the opportunity to emigrate to India where it is known from the end of MN 3. It has been found in Turkey from MN 6 and, as the primitive deinothere (elephant-like) ''
Prodeinotherium ''Prodeinotherium'' is an extinct representative of the family Deinotheriidae that lived in Africa, Europe, and Asia in the early and middle Miocene. ''Prodeinotherium'', meaning "before terrible beast", was first named in 1930, but soon after, t ...
'' is known from
Lesbos Lesbos or Lesvos ( el, Λέσβος, Lésvos ) is a Greek island located in the northeastern Aegean Sea. It has an area of with approximately of coastline, making it the third largest island in Greece. It is separated from Asia Minor by the na ...
, Greece, it seems likely there was a connection between the regions around 18.4 mya. Emigration occurred in both directions across the landbridge around 20.7 mya. The MN 3 fauna of
Negev The Negev or Negeb (; he, הַנֶּגֶב, hanNegév; ar, ٱلنَّقَب, an-Naqab) is a desert and semidesert region of southern Israel. The region's largest city and administrative capital is Beersheba (pop. ), in the north. At its sout ...
, Israel, (Abstract) provides a mixture of African and Eurasian
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
representative for this interchange: the elephant-like ''Prodeinotherium'' and ''Gomphotherium'', the carnivorous '' Anasinopa'', the small ungulate '' Dorcatherium'', the early pika '' Kenyalagomys'', and rodent '' Megapedetes'' from Africa; and the bovid ''
Eotragus ''Eotragus'' is an extinct genus of early bovid. Species belonging to the genus inhabited Europe, Africa, and Asia during the Miocene some 20-18 million years ago. It is related to the modern nilgai and four-horned antelope. It was small and ...
'', the suid ''
Listriodon ''Listriodon'' is an extinct genus of pig-like animals that lived in Eurasia during the Miocene. Description ''Listriodon'' species were generally small in size. In morphology, they show many similarities with peccaries rather than modern pigs. ...
'', and ''
Rhinocerotidae A rhinoceros (; ; ), commonly abbreviated to rhino, is a member of any of the five extant species (or numerous extinct species) of odd-toed ungulates in the family Rhinocerotidae. (It can also refer to a member of any of the extinct species ...
'' from Asia. The first to emigrate from Africa were the
proboscidea The Proboscidea (; , ) are a taxonomic order of afrotherian mammals containing one living family (Elephantidae) and several extinct families. First described by J. Illiger in 1811, it encompasses the elephants and their close relatives. Fr ...
ns (elephant-like) '' Gomphoteres'' and '' Deinotheres'' who eventually formed part of the
Bugti Bugti ( bal, بُگٹی ) is a Baloch tribe found in eastern Balochistan, Pakistan. , it was estimated to comprise over 180,000 people, mostly living in the Dera Bugti region of Pakistan. They are in turn divided into the Rahija,Marhita, Perozan ...
fauna in India around 18.3 mya and are known from the eastern Mediterranean around 18.4 mya and the Iberian Peninsula at the beginning of MN 4, 18 mya. Emigrants from Asia, included the bovid ''Eotragus'', the suid ''
Bunolistriodon ''Bunolistriodon'' is an extinct genus of pig-like animals from Eurasia and Africa during the Miocene. The exact position of ''Bunolistriodon'' has been subject to debate. It has been considered a synonym of ''Listriodon ''Listriodon'' is an ...
'', the
nimravid Nimravidae is an extinct family of carnivorans, sometimes known as false saber-toothed cats, whose fossils are found in North America and Eurasia. Not considered to belong to the true cats (family Felidae), the nimravids are generally considered ...
(cat) '' Prosansanosmilus'', and rodents such as '' Megacricetodon'', '' Democricetodon'', '' Cricetodon'', and '' Eumyarion''. The subfamily ''
Democricetodontinae The Democricetodontinae are a subfamily of fossil rodents from Miocene epoch. The Democricetodontini were named as a paraphyletic taxon within the Cricetidae by Lindsay in 1987. It was reranked as a subfamily by Theocharopoulos in 2000. See al ...
'' appeared in Asia Minor during MN 1-2, in Africa and Asia during MN 3, in America during MN 3-4 and finally in the Balkans and Western Europe during MN 4.


See also

*
Langhian The Langhian is, in the ICS geologic timescale, an age or stage in the middle Miocene Epoch/Series. It spans the time between 15.97 ± 0.05 Ma and 13.65 ± 0.05 Ma (million years ago) during the Middle Miocene.GeoWhen (2007) The Langhian was ...


References

{{Neogene Footer Miocene Miocene geochronology