Oenothera lamarckiana
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''Oenothera glazioviana'' is a species of flowering plant in the evening primrose family known by the common names large-flowered evening-primrose and redsepal evening primrose. ''Oenothera lamarckiana'' was formerly believed to be a different species, but is now regarded as a synonym of ''Oe. glazioviana''. The plant can be found in scattered locations worldwide, mostly as an
introduced species An introduced species, alien species, exotic species, adventive species, immigrant species, foreign species, non-indigenous species, or non-native species is a species living outside its native distributional range, but which has arrived ther ...
. It originated in
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
. It has long been cultivated as an
ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...
. In some locations it has become an
invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...
.


Description

''Oenothera glazioviana'' is generally a biennial herb producing an erect stem approaching in maximum height. It is roughly hairy in texture, the hairs with reddish blistering or glandular bases. The crinkly leaves are up to 15 centimeters long. The
inflorescence In botany, an inflorescence is a group or cluster of flowers arranged on a plant's Plant stem, stem that is composed of a main branch or a system of branches. An inflorescence is categorized on the basis of the arrangement of flowers on a mai ...
is a showy spike of many large flowers. When in bud the long red
sepal A sepal () is a part of the flower of angiosperms (flowering plants). Usually green, sepals typically function as protection for the flower in bud, and often as support for the petals when in bloom., p. 106 Etymology The term ''sepalum'' ...
s are visible. When in bloom each flower has four bright yellow petals up to 5 centimeters long which fade orange to red with age. The fruit is a lance-shaped capsule 2 or 3 centimeters long.


Taxonomy

''Oenothera glazioviana'' was first described by Marc Micheli in 1875. Originally native to
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
, it has become naturalized in many countries, and has acquired a large number of
synonyms A synonym is a word, morpheme, or phrase that means precisely or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
.


''Oenothera lamarckiana''

About a century ago, it was believed that there was a different species, either native to some obscure and unknown place in North America, from which it had quickly spread across the world, or more likely a new species which had very recently evolved in the last few decades, possibly in Europe from a hybrid of two other species, and thence had become a common weed. These theories stemmed from the fact that although the species was now a common species, and while an obviously striking species unlikely to be overlooked by botanists, it had only been recorded in recent times, and never in a truly wild state. At the time this
taxon In biology, a taxon (back-formation from ''taxonomy''; : taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular name and ...
was important for the brand new study of
genetics Genetics is the study of genes, genetic variation, and heredity in organisms.Hartl D, Jones E (2005) It is an important branch in biology because heredity is vital to organisms' evolution. Gregor Mendel, a Moravian Augustinians, Augustinian ...
, the debate about the cause of evolution, whether that was
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the Heredity, heritable traits characteristic of a population over generation ...
or one of the alternatives such as
mutationism Mutationism is one of several alternatives to evolution by natural selection that have existed both before and after the publication of Charles Darwin's 1859 book ''On the Origin of Species''. In the theory, mutation was the source of novelty, cr ...
, and particularly to the discovery of
polyploidy Polyploidy is a condition in which the cells of an organism have more than two paired sets of ( homologous) chromosomes. Most species whose cells have nuclei (eukaryotes) are diploid, meaning they have two complete sets of chromosomes, one fro ...
. It was later discovered that it had, in fact, already been discovered and described by a botanist in Brazil only a few decades beforehand, in 1875, as ''Oenothera glazioviana'', and had likely spread across the world from there since then, but this had apparently somehow been overlooked.


Distribution

''Oenothera glazioviana'' is native to
Brazil Brazil, officially the Federative Republic of Brazil, is the largest country in South America. It is the world's List of countries and dependencies by area, fifth-largest country by area and the List of countries and dependencies by population ...
. It is cultivated as an ornamental plant, and has become naturalized in many countries around the world, like Britain and Ireland, where it is the most common species of its genus.


Ecology

Under the synonym ''Oenothera lamarckiana'', it is described as a very adaptable plant: however it needs full sun, average moisture, and well-drained
soil Soil, also commonly referred to as earth, is a mixture of organic matter, minerals, gases, water, and organisms that together support the life of plants and soil organisms. Some scientific definitions distinguish dirt from ''soil'' by re ...
s. It is easily grown from seed. It began being grown in European gardens during the 1800s. File:Oenothera erythrosepala - Große Nachtkerze - Grote teunisbloem - l'Onagre à sépales striés de rouge - Evening Primrose.JPG, ''Oenothera glazioviana'' File:Oenothera erythrosepala - Große Nachtkerze - Grote teunisbloem - l'Onagre à sépales striés de rouge - Evening Primrose - ganze Blume.JPG, ''Oenothera glazioviana'' File:Nachtkerze (Oenothera glazioviana).webm, Video of blooming in real-time


References


Further reading

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External links


Jepson Manual Treatment''Oenothera glazioviana'' — U.C. Photo gallery
{{Taxonbar, from=Q1043961 glazioviana Night-blooming plants Garden plants of Europe Articles containing video clips Flora of Brazil