Ocellated Antbird
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The ocellated antbird (''Phaenostictus mcleannani'') is a species of antbird in subfamily Thamnophilinae of family Thamnophilidae, the "typical antbirds". It is found in Colombia, Costa Rica, Ecuador, Honduras, Nicaragua, and Panama.


Taxonomy and systematics

The ocellated antbird was formally described in 1861 by the American amateur ornithologist
George Newbold Lawrence George Newbold Lawrence (October 20, 1806 – January 17, 1895) was an American businessman and amateur ornithologist. Early life Lawrence was born in the city of New York on October 20, 1806. From his youth, Lawrence was a lover of birds and sp ...
and given the
binomial name In taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin grammatical forms, altho ...
''Phlogopsis mcleannani'' (misspelled as Meleannani). The ocellated antbird is now placed in the genus ''Phaenostictus'' that was erected in 1909 by the American ornithologist
Robert Ridgway Robert Ridgway (July 2, 1850 – March 25, 1929) was an American ornithologist specializing in systematics. He was appointed in 1880 by Spencer Fullerton Baird, secretary of the Smithsonian Institution, to be the first full-time curator of birds ...
. The genus name is derived from the
Ancient Greek Ancient Greek (, ; ) includes the forms of the Greek language used in ancient Greece and the classical antiquity, ancient world from around 1500 BC to 300 BC. It is often roughly divided into the following periods: Mycenaean Greek (), Greek ...
''phainō'' meaning "to display" and ''stiktos'' for "spotted". The
specific epithet In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...
honors James McLeannan, a railway engineer on the Panama Canal Railway, who had collected the
type specimen In biology, a type is a particular wikt:en:specimen, specimen (or in some cases a group of specimens) of an organism to which the scientific name of that organism is formally associated. In other words, a type is an example that serves to ancho ...
in Panama. Ridgway considered that the species was related to the genus '' Phlegopsis'' (the bare-eyes) but that it differed in having a longer tail, rounded nostrils and a few other characters. Molecular phylogenetic studies of the antbird family, Thamnophilidae, have found that the ocellated antbird sits in the tribe Pithyini and its closest relatives are in genus '' Pithys''. The ocellated antbird is the only member of its genus and has three subspecies, the nominate ''P. m. mcleannani'' ( Lawrence, 1860), ''P. m. saturatus'' (
Richmond Richmond most often refers to: * Richmond, British Columbia, a city in Canada * Richmond, California, a city in the United States * Richmond, London, a town in the London Borough of Richmond upon Thames, England * Richmond, North Yorkshire, a town ...
, 1896), and ''P. m. pacificus'' ( Hellmayr, 1924).


Description

The ocellated antbird is a largish antbird with a long bill and tail. It is long and weighs about .Batcheller, H. (2020). Ocellated Antbird (''Phaenostictus mcleannani''), version 1.0. In Birds of the World (T. S. Schulenberg, Editor). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.oceant1.01 (subscription required) retrieved May 24, 2024 Females tend to be slightly smaller than males and weigh slightly less. The species conforms with
Bergmann's rule Bergmann's rule is an ecogeographical rule that states that, within a broadly distributed taxonomic clade, populations and species of larger size are found in colder environments, while populations and species of smaller size are found in warmer ...
, with birds closer to the Equator having smaller wings and bills than those further away. Except for their slight size differences, the sexes are otherwise the same, and the subspecies have very little variation. Adults have a brown-gray crown, a rufous-brown nape, bright blue bare skin around the eye, and black cheeks and throat. Their back feathers and wing
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
are black with narrow rufous-brown edges that give a scaly appearance. Their rump is plain greenish brown, their flight feathers plain dark brown, and their tail black. Their breast is plain rufous-brown like their nape. Their belly feathers are black with wider rufous-brown edges than the back feathers and their vent area is reddish brown. Immature birds are similar to adults but with a darker cap and a less crisp pattern on their back and belly.


Distribution and habitat

The ocellated antbird ranges from Honduras to Ecuador. Subspecies ''P. m. saturatus'' is the northernmost, found from northern and eastern Honduras south through Nicaragua and the Caribbean slope of Costa Rica into
Bocas del Toro Province Bocas del Toro (; meaning "Mouths of the Bull") is a Provinces of Panama, province of Panama. Its area is 4,643.9 square kilometers, comprising the mainland and nine main islands. The province consists of the Bocas del Toro Archipelago, Bahía Al ...
in extreme western Panama. The nominate subspecies is found from central and eastern Panama into northwestern and western Colombia as far as Valle del Cauca Department. ''P. m. pacificus'' is found from extreme southwestern Colombia into northwestern Ecuador's
Esmeraldas Province Esmeraldas () is a northwestern coastal province of Ecuador. The capital and largest city is Esmeraldas. Esmeraldas is one of the three provinces of Ecuador that borders Colombia, and it is the most northern province in the country. The provin ...
. The ocellated antbird inhabits lowland and foothill
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, live oak, acacias, magnolia, and banksia in more temperate zo ...
, where it favors old-growth
primary forest An old-growth forest or primary forest is a forest that has developed over a long period of time without Disturbance (ecology), disturbance. Due to this, old-growth forests exhibit unique ecological features. The Food and Agriculture Organizati ...
but occurs in mature
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
as well. It almost entirely remains in the forest undergrowth. In elevation it reaches in much of Central America though only to in Costa Rica. In Colombia it occurs below and in Ecuador mostly below but locally reaches .


Behavior


Movement

The ocellated antbird is a year-round resident throughout its range.


Feeding

The ocellated antbird is an obligate ant follower that feeds on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s that flee foraging
army ant The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limited ...
swarms such as those of ''
Eciton burchellii ''Eciton burchellii'' is a species of New World army ant in the genus '' Eciton''. This species performs expansive, organized swarm raids that give it the informal name, ''Eciton'' army ant. This species displays a high degree of worker polymor ...
''. It typically forages in pairs or family groups, perching within about of the ground, often near the front of the ant swarm, and sallying after prey. Though ocellated antbirds do not join conventional
mixed-species foraging flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock (birds), flock of usually insectivorous birds of different species that join each other and move together while fora ...
s, often several species of birds attend the same ant swarms, and the ocellated antbird is often the dominant species. It seldom enters or crosses open areas unless the ants that they are following do so.


Breeding

The ocellated antbird's social biology is unusual for the antbird family. The breeding pair form the nucleus of a group or clan that includes their male offspring and their mates. These clans work together to defend nesting territories against rivals. Their nest is a cup made of twigs, bits of vine, and leaves held together with fungal
rhizomorph Mycelial cords are linear aggregations of parallel-oriented hyphae. The mature cords are composed of wide, empty vessel hyphae surrounded by narrower sheathing hyphae. Cords may look similar to plant roots, and also frequently have similar functi ...
s, and is typically placed on the ground between buttresses of a tree. The clutch is typically two eggs. Both parents incubate the clutch and provision the young. The incubation period, time to fledging, and other details of parental care are not known.


Vocalization

The ocellated antbird's primary song is a loud "high, thin series of whistles that rises then falls, with latter notes gaining in harsh quality" that has been written as "peee-peee-pee-peepee-ee-ee-ee-ee-ee-ee-ee-eer-eer". It also has a fainter version of the song with more notes but without the harsh ending and another version with "chirring" notes interspersed. At ant swarms it often gives a nasal "dzurrr" or "dzeerr".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the ocellated antbird as being of Least concern. It has a very large range; its population size is not known and is believed to be decreasing. No immediate threats have been identified. "Ocellated Antbirds are demonstrated to be very sensitive to forest fragmentation nd itslarge feeding range requirements also contribute to their sensitivity."


References

{{Taxonbar, from=Q1082851 ocellated antbird Birds of Honduras Birds of Nicaragua Birds of Costa Rica Birds of Panama Birds of Tumbes-Chocó-Magdalena ocellated antbird Taxonomy articles created by Polbot