The Nubian wild ass (''Equus africanus africanus'') is the

nominate subspecies In biological classification, subspecies is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. Not all speci ...

African wild ass The African wild ass (''Equus africanus'') or African wild donkey is a wild member of the horse family, Equidae. This species is thought to be the ancestor of the domestic donkey (''Equus asinus''), which is sometimes placed within the same spec ...

, and one of the ancestors of the domestic donkey, which was domesticated about 6,000 years ago. It is presumed to be

extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...

, though two populations potentially survive on the

Caribbean The Caribbean (, ) ( es, El Caribe; french: la Caraïbe; ht, Karayib; nl, De Caraïben) is a region of the Americas that consists of the Caribbean Sea, its islands (some surrounded by the Caribbean Sea and some bordering both the Caribbean ...

island of

Bonaire Bonaire (; , ; pap, Boneiru, , almost pronounced ) is a Dutch island in the Leeward Antilles in the Caribbean Sea. Its capital is the port of Kralendijk, on the west ( leeward) coast of the island. Aruba, Bonaire and Curaçao form the ABC ...

and in

Gebel Elba Gabal Elba ( ar, جَبَل علْبَة ' , ), or Elba Mountain, is a peak and, in general, includes the associated mountainous area in the Halaib Triangle of Northeast Africa. Despite being claimed by both Egypt and Sudan, the area is currently ...

Description

According to Groves & Chubb, Nubian wild asses had longer ears than the

Somali wild ass The Somali wild ass (''Equus africanus somaliensis'') is a subspecies of the African wild ass. It is found in Somalia, the Southern Red Sea region of Eritrea, and the Afar Region of Ethiopia. The legs of the Somali wild ass are striped, resembl ...

, ranging from 182-245mm in length. These animals also possessed the "cross-pattern" famous in domestic donkeys but absent in the Somali subspecies, and lacked the "zebra-stripe-pattern" on the legs as found in Somali specimens.

Habitat and distribution

Nubian wild asses lived in the

Nubian Desert The Nubian Desert ( ar, صحراء النوبة, ''Şaḩrā’ an Nūbyah'') is in the eastern region of the Sahara Desert, spanning approximately 400,000 km2 of northeastern Sudan and northern Eritrea, between the Nile and the Red Sea ...

of northeastern

Sudan Sudan ( or ; ar, السودان, as-Sūdān, officially the Republic of the Sudan ( ar, جمهورية السودان, link=no, Jumhūriyyat as-Sūdān), is a country in Northeast Africa. It shares borders with the Central African Republic t ...

, from east of the

Nile River The Nile, , Bohairic , lg, Kiira , Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered the longest riv ...

to the

Red Sea The Red Sea ( ar, البحر الأحمر - بحر القلزم, translit=Modern: al-Baḥr al-ʾAḥmar, Medieval: Baḥr al-Qulzum; or ; Coptic: ⲫⲓⲟⲙ ⲛ̀ϩⲁϩ ''Phiom Enhah'' or ⲫⲓⲟⲙ ⲛ̀ϣⲁⲣⲓ ''Phiom ǹšari''; ...

and south to the

Atbarah River The Atbarah River ( ar, نهر عطبرة; transliterated: Nahr 'Atbarah), also known as the Black Nile, is a river in northeast Africa. It rises in northwest Ethiopia, approximately 50 km north of Lake Tana and 30 km west of Gondar. ...

and northern

Eritrea Eritrea ( ; ti, ኤርትራ, Ertra, ; ar, إرتريا, ʾIritriyā), officially the State of Eritrea, is a country in the Horn of Africa region of Eastern Africa, with its capital and largest city at Asmara. It is bordered by Ethiopi ...

Extinction and potential survival

Bonaire's Critically-Endangered Nubian Wild Ass

The Nubian wild ass has not been seen since the 1950s-70s and is presumed

. The last sighting of wild asses in the animal's native range were in the 1970s, in Eritrea's Barka Valley and the border between Eritrea and Sudan, during aerial flights. Similar to the decline of the Somali wild ass, the Nubian wild ass's extinction can largely be attributed to hunting, competition with

livestock Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to ani ...

for limited desert resources, and

hybridization Hybridization (or hybridisation) may refer to: *Hybridization (biology), the process of combining different varieties of organisms to create a hybrid *Orbital hybridization, in chemistry, the mixing of atomic orbitals into new hybrid orbitals *Nu ...

with the Domestic donkey. In 2014–2015, a mitochondrial DNA analysis was conducted on a population of feral donkeys on the Caribbean island of

. The study included DNA samples from the Bonaire animals, museum specimens of Nubian wild asses, sequences from four Somali wild asses, and a sequence from one Domestic donkey, used as a control. The results found that the Bonaire animals were very close to known pure Nubian wild asses and very different from other asses. However, research published in 2010 found that modern donkeys are descended from two lineages of wild ass, one being Nubian and the other being of unknown ancestry. This being the case, the fact that only one donkey was used in the test suggests that the findings are rather inconclusive. For more conclusive results to be found, more testing must be completed with a larger group of donkeys. Additionally, the analysis was a result of an outcry over the government of Bonaire planning a

cull In biology, culling is the process of segregating organisms from a group according to desired or undesired characteristics. In animal breeding, it is the process of removing or segregating animals from a breeding stock based on a specific tr ...

of the feral animals, leading to the possibility of the results being slightly biased towards shutting down the cull. The Gebel Elba National Park, near the border of

Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a List of transcontinental countries, transcontinental country spanning the North Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via a land bridg ...

and Sudan, is claimed to be home to a wild population of Nubian wild asses. However, the purity of the animals in question has yet to be studied.

References

{{Taxonbar, from=Q583920 African wild ass Mammals described in 1866 Herbivorous mammals Mammals of North Africa Mammals of Sudan Taxa named by Theodor von Heuglin