The North American cougar (''Puma concolor couguar'') is a

cougar The cougar (''Puma concolor'') (, ''Help:Pronunciation respelling key, KOO-gər''), also called puma, mountain lion, catamount and panther is a large small cat native to the Americas. It inhabits North America, North, Central America, Cent ...

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

. It is the biggest cat in North America (

North American jaguar The jaguar (''Panthera onca'') is a large cat species and the only living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat species in the Americas an ...

s are fairly small), and the second largest cat in the

New World The term "New World" is used to describe the majority of lands of Earth's Western Hemisphere, particularly the Americas, and sometimes Oceania."America." ''The Oxford Companion to the English Language'' (). McArthur, Tom, ed., 1992. New York: ...

. It was once common in eastern North America and is still prevalent in the western half of the continent. This subspecies includes populations in

western Canada Western Canada, also referred to as the Western provinces, Canadian West, or Western provinces of Canada, and commonly known within Canada as the West, is a list of regions of Canada, Canadian region that includes the four western provinces and t ...

, the

western United States The Western United States (also called the American West, the Western States, the Far West, the Western territories, and the West) is List of regions of the United States, census regions United States Census Bureau. As American settlement i ...

Florida Florida ( ; ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders the Gulf of Mexico to the west, Alabama to the northwest, Georgia (U.S. state), Georgia to the north, the Atlantic ...

Mexico Mexico, officially the United Mexican States, is a country in North America. It is the northernmost country in Latin America, and borders the United States to the north, and Guatemala and Belize to the southeast; while having maritime boundar ...

and

Central America Central America is a subregion of North America. Its political boundaries are defined as bordering Mexico to the north, Colombia to the southeast, the Caribbean to the east, and the Pacific Ocean to the southwest. Central America is usually ...

, and possibly

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

northwest of the

Andes Mountains The Andes ( ), Andes Mountains or Andean Mountain Range (; ) are the longest continental mountain range in the world, forming a continuous highland along the western edge of South America. The range is long and wide (widest between 18°S ...

. It thus includes the

extirpated Local extinction, also extirpation, is the termination of a species (or other taxon) in a chosen geographic area of study, though it still exists elsewhere. Local extinctions are contrasted with global extinctions. Local extinctions mark a chan ...

eastern cougar and extant

Florida panther The Florida panther is a North American cougar (''P. c. couguar'') population in South Florida. It lives in pinelands, tropical hardwood hammocks and mixed freshwater swamp forests. Its range includes the Big Cypress National Preserve, Everglade ...

population Population is a set of humans or other organisms in a given region or area. Governments conduct a census to quantify the resident population size within a given jurisdiction. The term is also applied to non-human animals, microorganisms, and pl ...

Taxonomic history

As of 2017, ''P. c. cougar'' was recognised as being valid by the Cat Classification Taskforce of the Cat Specialist Group. ''P. c. costaricensis'' had been regarded as a subspecies in Central America.

Description

The North American cougar has a solid tawny-colored coat without spots, though the color can vary from buff to cinnamon-brown and juveniles may have mild leftover spotting. It weighs . Females average , about the same as a

jaguar The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...

in the Chamela-Cuixmala Biosphere Reserve on the Mexican Pacific coast.

Habitat and distribution

The North American cougar lives in various places and

habitat In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...

s. Several populations still exist and are thriving in the

, Southern Florida, and

, but the North American cougar was once commonly found in eastern portions of the United States. It was believed to be extirpated there in the early 1900s. In Michigan, it was thought to have been killed off and extinct in the early 1900s. Today there is evidence to support that cougars could be on the rise in Mexico and could have a substantial population in years to come. Some mainstream scientists believe that small

relict A relict is a surviving remnant of a natural phenomenon. Biology A relict (or relic) is an organism that at an earlier time was abundant in a large area but now occurs at only one or a few small areas. Geology and geomorphology In geology, a r ...

populations may exist (around 50 individuals), especially in the

Appalachian Mountains The Appalachian Mountains, often called the Appalachians, are a mountain range in eastern to northeastern North America. The term "Appalachian" refers to several different regions associated with the mountain range, and its surrounding terrain ...

and eastern Canada. Recent scientific findings in hair traps in Fundy National Park in New Brunswick have confirmed the existence of at least three cougars in New Brunswick. The Ontario Puma Foundation estimates that there are currently 850 cougars in

Ontario Ontario is the southernmost Provinces and territories of Canada, province of Canada. Located in Central Canada, Ontario is the Population of Canada by province and territory, country's most populous province. As of the 2021 Canadian census, it ...

. The

Quebec Quebec is Canada's List of Canadian provinces and territories by area, largest province by area. Located in Central Canada, the province shares borders with the provinces of Ontario to the west, Newfoundland and Labrador to the northeast, ...

wildlife services also considers cougars to be present in the province as a threatened species after multiple DNA tests confirmed cougar hair in lynx mating sites. The only unequivocally known eastern population is the critically endangered

. There have been unconfirmed sightings in Elliotsville Plantation, Maine (north of Monson) and as early as 1997 in

New Hampshire New Hampshire ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Massachusetts to the south, Vermont to the west, Maine and the Gulf of Maine to the east, and the Canadian province of Quebec t ...

Sightings in the United States

Reported sightings of cougars in the United States continue today, including in locations of their former range where they are considered extirpated. *

California California () is a U.S. state, state in the Western United States that lies on the West Coast of the United States, Pacific Coast. It borders Oregon to the north, Nevada and Arizona to the east, and shares Mexico–United States border, an ...

** The California Department of Fish and Game has roughly estimated 4,000 to 6,000 cougars present in California since 1972 ** In 2021, a cougar was sighted walking through a San Francisco neighborhood ** In 2022, a cougar was sighted at a school in Daly City *

Minnesota Minnesota ( ) is a U.S. state, state in the Upper Midwestern region of the United States. It is bordered by the Canadian provinces of Manitoba and Ontario to the north and east and by the U.S. states of Wisconsin to the east, Iowa to the so ...

** In Southern Minnesota there have been a few sightings by deer hunters; the Department of Natural Resources has verified 14 cougar sightings since 2007. *

Connecticut Connecticut ( ) is a U.S. state, state in the New England region of the Northeastern United States. It borders Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the south. ...

** In 2011, a cougar was sighted in

Greenwich, Connecticut Greenwich ( ) is a New England town, town in southwestern Fairfield County, Connecticut, United States. As of the 2020 United States census, 2020 census, it had a population of 63,518. It is the largest town on Gold Coast (Connecticut), Connectic ...

, and later killed by an SUV in Milford after allegedly travelling from South Dakota. *

Illinois Illinois ( ) is a U.S. state, state in the Midwestern United States, Midwestern United States. It borders on Lake Michigan to its northeast, the Mississippi River to its west, and the Wabash River, Wabash and Ohio River, Ohio rivers to its ...

** On April 14, 2008, a cougar triggered a flurry of reports before being cornered and killed in the Chicago neighborhood of Roscoe Village while officers tried to contain it. The cougar was the first sighted in the city limits of Chicago since the city was founded in 1833. :*On November 22, 2013, a cougar was found on a farm near

Morrison Morrison may refer to: People * Morrison (surname), people with the Scottish surname Morrison * Morrison Heady (1829–1915), American poet * Morrison Mann MacBride (1877–1938), Canadian merchant Places in the United States * Morrison, Colorad ...

in Whiteside County, Illinois. An Illinois Department of Natural Resources officer subsequently shot and killed the cougar after determining it posed a risk to the public. *

Michigan Michigan ( ) is a peninsular U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest, Upper Midwestern United States. It shares water and land boundaries with Minnesota to the northwest, Wisconsin to the west, ...

** According to the Michigan Department of Natural Resources, there were fifteen confirmed cougar sightings in the state in 2020 and, as of October, there had been ten confirmed sightings in the

Upper Peninsula The Upper Peninsula of Michigan—also known as Upper Michigan or colloquially the U.P. or Yoop—is the northern and more elevated of the two major landmasses that make up the U.S. state of Michigan; it is separated from the Lower Peninsula b ...

in 2021. The Department said that the steep increase in sightings may be attributable to the proliferation of

trail camera A remote camera, also known as a trail camera or game camera, is a camera placed by a photographer in areas where the photographer generally cannot be at the camera to snap the shutter. This includes areas with limited access, tight spaces where ...

s. *

Tennessee Tennessee (, ), officially the State of Tennessee, is a landlocked U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders Kentucky to the north, Virginia to the northeast, North Carolina t ...

** On September 26, 2015, a hair sample was submitted by a hunter in

Carroll County, Tennessee Carroll County is a county located in the western division of the U.S. state of Tennessee. As of the 2020 census, the population was 28,440. Its county seat is Huntingdon. The county was established by the Tennessee General Assembly on Novembe ...

; DNA analysis indicated it was a female with genetics similar to cougars in South Dakota.

Bobcat The bobcat (''Lynx rufus''), also known as the wildcat, bay lynx, or red lynx, is one of the four extant species within the medium-sized wild cat genus '' Lynx''. Native to North America, it ranges from southern Canada through most of the c ...

s in this state currently reside in regions that were once roamed by cougars. *

Wisconsin Wisconsin ( ) is a U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest of the United States. It borders Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michig ...

** Genetic analysis of DNA from a cougar sighting in Wisconsin in 2008 indicated that a cougar was in Wisconsin and that it was not a

captive animal Animal captivity is the confinement of domestic and wild animals. More specifically, animals that are held by humans and prevented from escaping are said to be in captivity. The term animal captivity is usually applied to wild animals that are he ...

. The cougar is thought to have migrated from a native population in the Black Hills of

South Dakota South Dakota (; Sioux language, Sioux: , ) is a U.S. state, state in the West North Central states, North Central region of the United States. It is also part of the Great Plains. South Dakota is named after the Dakota people, Dakota Sioux ...

; however, the genetic analysis could not affirm that hypothesis. Whether other, perhaps breeding, cougars are present is also uncertain. A second sighting was reported and tracks were documented in a nearby Wisconsin community. Unfortunately, a genetic analysis could not be done and a determination could not be made. This cougar later made its way south into the northern

Chicago Chicago is the List of municipalities in Illinois, most populous city in the U.S. state of Illinois and in the Midwestern United States. With a population of 2,746,388, as of the 2020 United States census, 2020 census, it is the List of Unite ...

suburb of Wilmette. ** On June 3, 2013, a verified sighting was made in Florence County, Wisconsin. The cougar was photographed by an automatic trail camera, and confirmed by DNR biologists in October, 2013. ** In December, 2020, two sightings, one verified, were made in Dane County, in and around Stoughton, Wisconsin. The cougar was photographed by an individual, and confirmed by the Wisconsin DNR. ** In November 2021, a DNR representative told WDJT-TV that the Department confirms about 15 cougar sightings per year in the state. While the origins of these animals are unknown, some cougar experts believe some are captive animals that have been released or escaped.

Ecology

The North American Cougar is a

carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they ar ...

and its main sources of prey are deer, elk, mountain goats, moose and bighorn sheep. Despite being a large predator, the North American Cougar can also be the prey of larger predators like

wolves The wolf (''Canis lupus''; : wolves), also known as the grey wolf or gray wolf, is a canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, including the dog and dingo, though gr ...

and bears. The North American cougar usually hunts at night and sometimes travels long distances in search of food. They are short distance sprinters and can remain hidden for hours to surprise unsuspecting prey and pounce when they least expect it. They use their strong jaws and large canines to puncture the neck of their prey, breaking the neck and efficiently killing their prey. They also grab their prey by the throat to suffocate it. It is fast and can maneuver quite easily and skillfully. Depending on the abundance of

prey Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not ki ...

, such as

deer A deer (: deer) or true deer is a hoofed ruminant ungulate of the family Cervidae (informally the deer family). Cervidae is divided into subfamilies Cervinae (which includes, among others, muntjac, elk (wapiti), red deer, and fallow deer) ...

, it shares the same prey as the

in Central or North America. Other

sympatric In biology, two closely related species or populations are considered sympatric when they exist in the same geographic area and thus frequently encounter each other. An initially interbreeding population that splits into two or more distinct spe ...

predators include the

grizzly bear The grizzly bear (''Ursus arctos horribilis''), also known as the North American brown bear or simply grizzly, is a population or subspecies of the brown bear inhabiting North America. In addition to the mainland grizzly (''Ursus arctos horr ...

and

American black bear The American black bear (''Ursus americanus''), or simply black bear, is a species of medium-sized bear which is Endemism, endemic to North America. It is the continent's smallest and most widely distributed bear species. It is an omnivore, with ...

s. Cougars are known to prey on

bear Bears are carnivoran mammals of the family (biology), family Ursidae (). They are classified as caniforms, or doglike carnivorans. Although only eight species of bears are extant, they are widespread, appearing in a wide variety of habitats ...

cubs. Cougars in the

Great Basin The Great Basin () is the largest area of contiguous endorheic watersheds, those with no outlets to the ocean, in North America. It spans nearly all of Nevada, much of Utah, and portions of California, Idaho, Oregon, Wyoming, and Baja Californi ...

have been recorded to prey on feral horses, as well as feral donkeys in the Sonoran and

Mojave Desert The Mojave Desert (; ; ) is a desert in the rain shadow of the southern Sierra Nevada mountains and Transverse Ranges in the Southwestern United States. Named for the Indigenous peoples of the Americas, indigenous Mohave people, it is located pr ...

s. Rivalry between the cougar and grizzly bear was a popular topic in North America. Fights between them were staged, and those in the

wilderness Wilderness or wildlands (usually in the plurale tantum, plural) are Earth, Earth's natural environments that have not been significantly modified by human impact on the environment, human activity, or any urbanization, nonurbanized land not u ...

were recorded by people, including native peoples. The North American Cougar plays an important role in regulating ecosystems as a large predator. The presence of the cougar as a predator prevents the overpopulation of herbivorous prey, like deer, in an ecosystem. Overpopulation of prey can result in the destruction of vegetation and biodiversity in an ecosystem.

Reproduction

Adult male cougars can breed with multiple female cougars any time of the year, however the peak breeding season is in the months of January and August. When cougars are 2–3 years old, they reach the level of sexual maturity. The breeding process does not last a long time, with the male accompanying the female in heat for up to a week after which they separate. After mating, the male cougar plays no further role except driving off male intruders and the female cougar bears the full responsibility of raising her young. The average litter size is three cubs and each of the babies weigh a little over a pound (500 grams). Cougars have a 90–96 day

gestation Gestation is the period of development during the carrying of an embryo, and later fetus, inside viviparous animals (the embryo develops within the parent). It is typical for mammals, but also occurs for some non-mammals. Mammals during pregn ...

period, allowing the breeding process to continue throughout the year.

Threats and conservation

The primary causes of the declining population of cougars is due to hunting and loss of habitat.

Trophy hunting Trophy hunting is a form of hunting for field sports, sport in which parts of the hunted wild animals are kept and displayed as trophies. The animal being targeted, known as the "game (hunting), game", is typically a mature male specimen from a p ...

and loss of territory reflect the most significant threats upon the cougar extinction status. Most of the cougars’ prey is found near humans. Whether it be through sport hunting or through the protection of livestock, humans frequently kill cougars intentionally. Though indirect killings through vehicle collisions do occur, the intentional human impact is far more drastic. Humans continue to affect the declining cougar population through the occupation of their habitats. Cougars tend to occupy areas that are prime for development and expansion. From mountains to deserts, humans utilize the cougar territory to build new sites and structures for their purposes. As a consequence, cougars lose their habitats which leads to conflict with humans. Even though conservation efforts of the cougar have decreased against the "more appealing" jaguar, it is hunted less frequently because it has no spots, and is thus less desirable to hunters. Despite the declining population of cougars, the potential extinction of the North American Cougar is not seen as a large concern. In

Oregon Oregon ( , ) is a U.S. state, state in the Pacific Northwest region of the United States. It is a part of the Western U.S., with the Columbia River delineating much of Oregon's northern boundary with Washington (state), Washington, while t ...

, a population of 5,000 individuals was reported in 2006, exceeding a target of 3,000. California has actively sought to protect the cat and has an estimated population of 4,000 to 6,000. With the increase of human development and infrastructure growth in California, the cougar population in the state is becoming more isolated from one another. The Bay Area Puma Project aims to obtain information on cougar populations in the San Francisco Bay area and the animals' interactions with habitat, prey, humans, and residential communities. A study on wildlife ecologists showed that urban cougar populations exist around the

Los Angeles metropolitan area Greater Los Angeles is the most populous metropolitan area in the U.S. state of California, encompassing five counties in Southern California extending from Ventura County in the west to San Bernardino County and Riverside County in the east, ...

, with individuals of these populations having the smallest home ranges recorded for any cougars studied, and being primarily nocturnal and not crepuscular, most likely adaptations to avoid humans in high-density areas.

Communication and behavior

Cougars rely on scent, noises, and posture to communicate with each other to exchange messages. Each message depends on how the cougar delivers the sound. Growling, snarling and hissing is understood as a threat by other animals. Mother cougars chirp to communicate with their kittens. The caterwaul is a screeching sound made by female cougars during the mating season when competing males are present. Cougars use various methods to signal and communicate with each other. When cougars perceive a looming threat or danger nearby, they flatten their ears and either maintain eye contact with the threat or retreat to a less visible location in preparation to attack.

References

;Sources * Wright, Bruce S. The Eastern Panther: A Question of Survival. Toronto: Clarke, Irwin and Company, 1972.

External links

Eastern Cougar Foundation

National Heritage Information Centre: General Element Report: ''Puma concolor''

* ttps://www.iucnredlist.org/species/18868/97216466 The IUCN Red List of Threatened Species
Photograph of a black or dark cougar in Costa Rica

{{Taxonbar, from=Q2044439, from2=Q3235287 Cougar Felids of Central America Mammals described in 1792 Mammals of Canada Mammals of the United States Mammals of Mexico Taxa named by Robert Kerr (writer) Fur trade Fauna of California