A stem is one of two main structural axes of a

vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...

, the other being the

root In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the su ...

. It supports

leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...

flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...

s and

fruit In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering. Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...

s, transports water and dissolved substances between the roots and the shoots in the xylem and phloem, stores nutrients, and produces new living tissue. The stem can also be called halm or haulm. The stem is normally divided into nodes and internodes: * The nodes hold one or more leaves, as well as buds which can grow into branches (with

conifer cone A conifer cone (in formal botanical usage: strobilus, plural strobili) is a seed-bearing organ on gymnosperm plants. It is usually woody, ovoid to globular, including scales and bracts arranged around a central axis, especially in conifers ...

s, or

flowers A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanism ...

). Adventitious roots may also be produced from the nodes. * The internodes distance one node from another. The term " shoots" is often confused with "stems"; "shoots" generally refers to new fresh plant growth including both stems and other structures like leaves or flowers. In most plants stems are located above the soil surface but some plants have

underground stem Underground stems are modified plant parts that derive from stem tissue but exist under the soil surface. They function as storage tissues for food and nutrients, in propagation of new clones, and in perennation (survival from one growing season to ...

s. Stems have four main functions which are: * Support for and the elevation of leaves,

s, and

s. The stems keep the leaves in the light and provide a place for the plant to keep its flowers and fruits. * Transport of fluids between the roots and the shoots in the xylem and phloem (see below). * Storage of nutrients. * Production of new living tissue. The normal lifespan of plant cells is one to three years. Stems have cells called meristems that annually generate new living tissue. Stems have two pipe-like tissues called xylem and phloem. The xylem tissue transports water by the action of transpiration pull, capillary action, and

root pressure Root pressure is the transverse osmotic pressure within the cells of a root system that causes sap to rise through a plant stem to the leaves. Root pressure occurs in the xylem of some vascular plants when the soil moisture level is high eithe ...

. The phloem tissue consists of sieve tubes and their companion cells. The two tissues are separated by cambium, a tissue that divides to form xylem or phloem cells.

Specialized terms

Stems are often specialized for storage, asexual reproduction, protection, or

photosynthesis Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...

, including the following: Senecio angulatus stems

* Acaulescent: used to describe stems in plants that appear to be stemless. Actually these stems are just extremely short, the leaves appearing to rise directly out of the ground, e.g. some ''Viola'' species. * Arborescent: tree with woody stems normally with a single trunk. * Axillary bud: a bud which grows at the point of attachment of an older leaf with the stem. It potentially gives rise to a shoot. *

Branch A branch, sometimes called a ramus in botany, is a woody structural member connected to the central trunk of a tree (or sometimes a shrub). Large branches are known as boughs and small branches are known as twigs. The term '' twig'' usuall ...

ed: aerial stems are described as being branched or unbranched. * Bud: an embryonic shoot with immature stem tip. *

Bulb In botany, a bulb is structurally a short stem with fleshy leaves or leaf basesBell, A.D. 1997. ''Plant form: an illustrated guide to flowering plant morphology''. Oxford University Press, Oxford, U.K. that function as food storage organs dur ...

: a short vertical underground stem with fleshy storage leaves attached, e.g.

onion An onion (''Allium cepa'' L., from Latin ''cepa'' meaning "onion"), also known as the bulb onion or common onion, is a vegetable that is the most widely cultivated species of the genus '' Allium''. The shallot is a botanical variety of the on ...

daffodil ''Narcissus'' is a genus of predominantly spring flowering perennial plants of the amaryllis family, Amaryllidaceae. Various common names including daffodil,The word "daffodil" is also applied to related genera such as '' Sternbergia'', ''Is ...

, and tulip. Bulbs often function in reproduction by splitting to form new bulbs or producing small new bulbs termed bulblets. Bulbs are a combination of stem and leaves so may better be considered as leaves because the leaves make up the greater part. * Caespitose: when stems grow in a tangled mass or clump or in low growing mats. * Cladode (including phylloclade): a flattened stem that appears more-or-less leaf like and is specialized for photosynthesis, e.g.

cactus A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Gree ...

pads. *

Climbing Climbing is the activity of using one's hands, feet, or any other part of the body to ascend a steep topographical object that can range from the world's tallest mountains (e.g. the eight thousanders), to small boulders. Climbing is done ...

: stems that cling or wrap around other plants or structures. *

Corm A corm, bulbo-tuber, or bulbotuber is a short, vertical, swollen underground plant stem that serves as a storage organ that some plants use to survive winter or other adverse conditions such as summer drought and heat (perennation). The word ' ...

: a short enlarged underground, storage stem, e.g.

taro Taro () (''Colocasia esculenta)'' is a root vegetable. It is the most widely cultivated species of several plants in the family Araceae that are used as vegetables for their corms, leaves, and petioles. Taro corms are a food staple in Afri ...

crocus ''Crocus'' (; plural: crocuses or croci) is a genus of seasonal flowering plants in the family Iridaceae (iris family) comprising about 100 species of perennials growing from corms. They are low growing plants, whose flower stems remain under ...

, gladiolus. * Decumbent: stems that lie flat on the ground and turn upwards at the ends. * Fruticose: stems that grow shrublike with woody like habit. *

Herbaceous Herbaceous plants are vascular plants that have no persistent woody stems above ground. This broad category of plants includes many perennials, and nearly all annuals and biennials. Definitions of "herb" and "herbaceous" The fourth edition o ...

: non woody, they die at the end of the growing season. * Internode: an interval between two successive nodes. It possesses the ability to elongate, either from its base or from its extremity depending on the species. * Node: a point of attachment of a leaf or a

twig A twig is a thin, often short, branch of a tree or bush. The buds on the twig are an important diagnostic characteristic, as are the abscission scars where the leaves have fallen away. The color, texture, and patterning of the twig bark ar ...

on the stem in seed plants. A node is a very small growth zone. * Pedicel: stems that serve as the stalk of an individual flower in an

inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...

or infrutescence. * Peduncle: a stem that supports an

. * Prickle: a sharpened extension of the stem's outer layers, e.g.

rose A rose is either a woody perennial flowering plant of the genus ''Rosa'' (), in the family Rosaceae (), or the flower it bears. There are over three hundred species and tens of thousands of cultivars. They form a group of plants that can be ...

s. * Pseudostem: a false stem made of the rolled bases of leaves, which may be tall, as in

banana A banana is an elongated, edible fruit – botanically a berry – produced by several kinds of large herbaceous flowering plants in the genus ''Musa''. In some countries, bananas used for cooking may be called "plantains", disting ...

. *

Rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...

: a horizontal underground stem that functions mainly in reproduction but also in storage, e.g. most

fern A fern (Polypodiopsida or Polypodiophyta ) is a member of a group of vascular plants (plants with xylem and phloem) that reproduce via spores and have neither seeds nor flowers. The polypodiophytes include all living pteridophytes exce ...

s, iris. *

Runner Running is a method of terrestrial locomotion allowing humans and other animals to move rapidly on foot. Running is a type of gait characterized by an aerial phase in which all feet are above the ground (though there are exceptions). This is ...

: a type of stolon, horizontally growing on top of the ground and rooting at the nodes, aids in reproduction. e.g. garden strawberry, ''

Chlorophytum comosum ''Chlorophytum comosum'', usually called spider plant or common spider plant due to its spider-like look, also known as spider ivy, ribbon plant (a name it shares with '' Dracaena sanderiana''), and hen and chickens is a species of evergreen pere ...

''. * Scape: a stem that holds flowers that comes out of the ground and has no normal leaves. Hosta,

lily ''Lilium'' () is a genus of herbaceous flowering plants growing from bulbs, all with large prominent flowers. They are the true lilies. Lilies are a group of flowering plants which are important in culture and literature in much of the world. M ...

, iris,

garlic Garlic (''Allium sativum'') is a species of bulbous flowering plant in the genus '' Allium''. Its close relatives include the onion, shallot, leek, chive, Welsh onion and Chinese onion. It is native to South Asia, Central Asia and northeas ...

. *

Stolon In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...

: a horizontal stem that produces rooted plantlets at its nodes and ends, forming near the surface of the ground. *

Thorn Thorn(s) or The Thorn(s) may refer to: Botany * Thorns, spines, and prickles, sharp structures on plants * ''Crataegus monogyna'', or common hawthorn, a plant species Comics and literature * Rose and Thorn, the two personalities of two DC Comic ...

: a modified stem with a sharpened point. *

Tuber Tubers are a type of enlarged structure used as storage organs for nutrients in some plants. They are used for the plant's perennation (survival of the winter or dry months), to provide energy and nutrients for regrowth during the next growing ...

: a swollen, underground storage stem adapted for storage and reproduction, e.g.

potato The potato is a starchy food, a tuber of the plant ''Solanum tuberosum'' and is a root vegetable native to the Americas. The plant is a perennial in the nightshade family Solanaceae. Wild potato species can be found from the southern Uni ...

. * Woody: hard textured stems with secondary xylem.

Stem structure

Stem usually consist of three tissues, dermal tissue, ground tissue and vascular tissue. The dermal tissue covers the outer surface of the stem and usually functions to waterproof, protect and control gas exchange. The ground tissue usually consists mainly of parenchyma cells and fills in around the vascular tissue. It sometimes functions in photosynthesis. Vascular tissue provides long distance transport and structural support. Most or all ground tissue may be lost in woody stems. The dermal tissue of aquatic plants stems may lack the waterproofing found in aerial stems. The arrangement of the vascular tissues varies widely among plant species.

Dicot stems

Dicot The dicotyledons, also known as dicots (or, more rarely, dicotyls), are one of the two groups into which all the flowering plants (angiosperms) were formerly divided. The name refers to one of the typical characteristics of the group: namely, ...

stems with primary growth have pith in the center, with vascular bundles forming a distinct ring visible when the stem is viewed in cross section. The outside of the stem is covered with an epidermis, which is covered by a waterproof cuticle. The epidermis also may contain

stomata In botany, a stoma (from Greek ''στόμα'', "mouth", plural "stomata"), also called a stomate (plural "stomates"), is a pore found in the epidermis of leaves, stems, and other organs, that controls the rate of gas exchange. The pore is b ...

for gas exchange and multicellular stem hairs called trichomes. A cortex consisting of hypodermis (collenchyma cells) and

endodermis The endodermis is the central, innermost layer of cortex in land plants. It is a cylinder of compact living cells, the radial walls of which are impregnated with hydrophobic substances (Casparian strip) to restrict apoplastic flow of water to th ...

(starch containing cells) is present above the pericycle and vascular bundles. Woody dicots and many nonwoody dicots have secondary growth originating from their lateral or secondary meristems: the

vascular cambium The vascular cambium is the main growth tissue in the stems and roots of many plants, specifically in dicots such as buttercups and oak trees, gymnosperms such as pine trees, as well as in certain other vascular plants. It produces secondary xy ...

and the cork cambium or phellogen. The vascular cambium forms between the xylem and phloem in the vascular bundles and connects to form a continuous cylinder. The vascular cambium cells divide to produce

secondary xylem Xylem is one of the two types of transport tissue in vascular plants, the other being phloem. The basic function of xylem is to transport water from roots to stems and leaves, but it also transports nutrients. The word ''xylem'' is derived from ...

to the inside and

secondary phloem Phloem (, ) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is ...

to the outside. As the stem increases in diameter due to production of secondary xylem and secondary phloem, the cortex and epidermis are eventually destroyed. Before the cortex is destroyed, a cork cambium develops there. The cork cambium divides to produce waterproof cork cells externally and sometimes phelloderm cells internally. Those three tissues form the periderm, which replaces the epidermis in function. Areas of loosely packed cells in the periderm that function in gas exchange are called lenticels. Secondary xylem is commercially important as

wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...

. The seasonal variation in growth from the

is what creates yearly tree rings in temperate climates. Tree rings are the basis of

dendrochronology Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and atm ...

, which dates wooden objects and associated artifacts. Dendroclimatology is the use of tree rings as a record of past climates. The aerial stem of an adult

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...

is called a

trunk Trunk may refer to: Biology * Trunk (anatomy), synonym for torso * Trunk (botany), a tree's central superstructure * Trunk of corpus callosum, in neuroanatomy * Elephant trunk, the proboscis of an elephant Computing * Trunk (software), in rev ...

. The dead, usually darker inner wood of a large diameter trunk is termed the heartwood and is the result of tylosis. The outer, living wood is termed the sapwood.

Monocot stems

Vascular bundles are present throughout the

monocot Monocotyledons (), commonly referred to as monocots, (Lilianae '' sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one ...

stem, although concentrated towards the outside. This differs from the dicot stem that has a ring of vascular bundles and often none in the center. The shoot apex in monocot stems is more elongated. Leaf sheathes grow up around it, protecting it. This is true to some extent of almost all monocots. Monocots rarely produce secondary growth and are therefore seldom woody, with palms and

bamboo Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...

being notable exceptions. However, many monocot stems increase in diameter via anomalous secondary growth.

Gymnosperm stems

All

gymnosperm The gymnosperms ( lit. revealed seeds) are a group of seed-producing plants that includes conifers, cycads, '' Ginkgo'', and gnetophytes, forming the clade Gymnospermae. The term ''gymnosperm'' comes from the composite word in el, γυμν ...

s are woody plants. Their stems are similar in structure to woody dicots except that most gymnosperms produce only tracheids in their xylem, not the vessels found in dicots. Gymnosperm wood also often contains

resin In polymer chemistry and materials science, resin is a solid or highly viscous substance of plant or synthetic origin that is typically convertible into polymers. Resins are usually mixtures of organic compounds. This article focuses on nat ...

ducts. Woody dicots are called hardwoods, e.g. oak,

maple ''Acer'' () is a genus of trees and shrubs commonly known as maples. The genus is placed in the family Sapindaceae.Stevens, P. F. (2001 onwards). Angiosperm Phylogeny Website. Version 9, June 2008 nd more or less continuously updated since h ...

and walnut. In contrast, softwoods are gymnosperms, such as

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...

spruce A spruce is a tree of the genus ''Picea'' (), a genus of about 35 species of coniferous evergreen trees in the family Pinaceae, found in the northern temperate and boreal (taiga) regions of the Earth. ''Picea'' is the sole genus in the subfam ...

and fir. Image:Redwood M D Vaden.jpg, The trunk of this

redwood Sequoioideae, popularly known as redwoods, is a subfamily of coniferous trees within the family Cupressaceae. It includes the largest and tallest trees in the world. Description The three redwood subfamily genera are '' Sequoia'' from coasta ...

tree is its stem.

Fern stems

Most

s have

rhizome In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...

s with no vertical stem. The exception is tree ferns, with vertical stems up to about 20 metres. The stem anatomy of ferns is more complicated than that of dicots because fern stems often have one or more leaf gaps in cross section. A leaf gap is where the vascular tissue branches off to a frond. In cross section, the vascular tissue does not form a complete cylinder where a leaf gap occurs. Fern stems may have solenosteles or dictyosteles or variations of them. Many fern stems have phloem tissue on both sides of the xylem in cross-section. File:Ferns.jpg, Tasmanian tree fern

Relation to xenobiotics

Foreign chemicals such as air pollutants,C. Michael Hogan. 2010
"Abiotic factor"
''Encyclopedia of Earth''. Emily Monosson and C. Cleveland, eds. National Council for Science and the Environment . Washington, D.C. herbicides and pesticides can damage stem structures.

Economic importance

There are thousands of species whose stems have economic uses. Stems provide a few major staple crops such as

and

Sugarcane Sugarcane or sugar cane is a species of (often hybrid) tall, perennial grass (in the genus '' Saccharum'', tribe Andropogoneae) that is used for sugar production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fibrous stalk ...

stems are a major source of sugar.

Maple sugar Maple sugar is a traditional sweetener in Canada and the northeastern United States, prepared from the sap of the maple tree (" maple sap"). Sources Three species of maple trees in the genus '' Acer'' are predominantly used to produce mapl ...

is obtained from trunks of

trees.

Vegetable Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is applied to plants collectively to refer to all edible plant matter, including the edible flower, flowers, ...

s from stems are asparagus, bamboo shoots, cactus pads or nopalitos, kohlrabi, and

water chestnut Water chestnut may refer to either of two plants (both sometimes used in Chinese cuisine): * The Chinese water chestnut ('' Eleocharis dulcis''), eaten for its crisp corm * The water caltrop The water caltrop is any of three extant species of th ...

. The spice,

cinnamon Cinnamon is a spice obtained from the inner bark of several tree species from the genus '' Cinnamomum''. Cinnamon is used mainly as an aromatic condiment and flavouring additive in a wide variety of cuisines, sweet and savoury dishes, breakf ...

is bark from a tree trunk. Gum arabic is an important food additive obtained from the trunks of ''

Acacia senegal ''Senegalia senegal'' (also known as ''Acacia senegal'') is a small thorny deciduous tree from the genus ''Senegalia'', which is known by several common names, including gum acacia, gum arabic tree, Sudan gum and Sudan gum arabic. In parts of In ...

'' trees.

Chicle Chicle () is a natural gum traditionally used in making chewing gum and other products. It is collected from several species of Mesoamerican trees in the genus ''Manilkara'', including '' M. zapota'', '' M. chicle'', '' M. staminodella'', and '' ...

, the main ingredient in chewing gum, is obtained from trunks of the chicle tree. Medicines obtained from stems include

quinine Quinine is a medication used to treat malaria and babesiosis. This includes the treatment of malaria due to '' Plasmodium falciparum'' that is resistant to chloroquine when artesunate is not available. While sometimes used for nocturnal leg ...

from the bark of

cinchona ''Cinchona'' (pronounced or ) is a genus of flowering plants in the family Rubiaceae containing at least 23 species of trees and shrubs. All are native to the tropical Andean forests of western South America. A few species are reportedly nat ...

trees,

camphor Camphor () is a waxy, colorless solid with a strong aroma. It is classified as a terpenoid and a cyclic ketone. It is found in the wood of the camphor laurel (''Cinnamomum camphora''), a large evergreen tree found in East Asia; and in the k ...

distilled from wood of a tree in the same genus that provides cinnamon, and the muscle relaxant curare from the bark of tropical vines.

Wood Wood is a porous and fibrous structural tissue found in the stems and roots of trees and other woody plants. It is an organic materiala natural composite of cellulose fibers that are strong in tension and embedded in a matrix of lignin ...

is used in thousands of ways, e.g.

buildings A building, or edifice, is an enclosed structure with a roof and walls standing more or less permanently in one place, such as a house or factory (although there's also portable buildings). Buildings come in a variety of sizes, shapes, and func ...

furniture Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating ( tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks) ...

boat A boat is a watercraft of a large range of types and sizes, but generally smaller than a ship, which is distinguished by its larger size, shape, cargo or passenger capacity, or its ability to carry boats. Small boats are typically found on i ...

airplane An airplane or aeroplane (informally plane) is a fixed-wing aircraft that is propelled forward by thrust from a jet engine, propeller, or rocket engine. Airplanes come in a variety of sizes, shapes, and wing configurations. The broad ...

s, wagons, car parts,

musical instrument A musical instrument is a device created or adapted to make musical sounds. In principle, any object that produces sound can be considered a musical instrument—it is through purpose that the object becomes a musical instrument. A person who pl ...

sports equipment Sports equipment, sporting equipment, also called sporting goods, are the tools, materials, apparel, and gear used to compete in a sport and varies depending on the sport. The equipment ranges from balls, nets, and protective gear like helmets. ...

railroad tie A railroad tie, crosstie (American English), railway tie ( Canadian English) or railway sleeper ( Australian and British English) is a rectangular support for the rails in railroad tracks. Generally laid perpendicular to the rails, ties tran ...

utility pole A utility pole is a column or post typically made out of wood used to support overhead power lines and various other public utilities, such as electrical cable, fiber optic cable, and related equipment such as transformers and street lights. I ...

s, fence posts, pilings,

toothpick A toothpick is a small thin stick of wood, plastic, bamboo, metal, bone or other substance with at least one and sometimes two pointed ends to insert between teeth to remove detritus, usually after a meal. Toothpicks are also used for festiv ...

match A match is a tool for starting a fire. Typically, matches are made of small wooden sticks or stiff paper. One end is coated with a material that can be ignited by friction generated by striking the match against a suitable surface. Wooden mat ...

es,

plywood Plywood is a material manufactured from thin layers or "plies" of wood veneer that are glued together with adjacent layers having their wood grain rotated up to 90 degrees to one another. It is an engineered wood from the family of manufactured ...

coffin A coffin is a funerary box used for viewing or keeping a corpse, either for burial or cremation. Sometimes referred to as a casket, any box in which the dead are buried is a coffin, and while a casket was originally regarded as a box for j ...

shingles Shingles, also known as zoster or herpes zoster, is a viral disease characterized by a painful skin rash with blisters in a localized area. Typically the rash occurs in a single, wide mark either on the left or right side of the body or fac ...

barrel A barrel or cask is a hollow cylindrical container with a bulging center, longer than it is wide. They are traditionally made of wooden staves and bound by wooden or metal hoops. The word vat is often used for large containers for liquids, ...

staves, toys,

tool A tool is an object that can extend an individual's ability to modify features of the surrounding environment or help them accomplish a particular task. Although many animals use simple tools, only human beings, whose use of stone tools dates b ...

handles, picture frames, veneer,

charcoal Charcoal is a lightweight black carbon residue produced by strongly heating wood (or other animal and plant materials) in minimal oxygen to remove all water and volatile constituents. In the traditional version of this pyrolysis process, ...

and

firewood Firewood is any wooden material that is gathered and used for fuel. Generally, firewood is not highly processed and is in some sort of recognizable log or branch form, compared to other forms of wood fuel like pellets or chips. Firewood ca ...

Wood pulp Pulp is a lignocellulosic fibrous material prepared by chemically or mechanically separating cellulose fibers from wood, fiber crops, waste paper, or rags. Mixed with water and other chemical or plant-based additives, pulp is the major raw ...

is widely used to make

paper Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distribu ...

paperboard Paperboard is a thick paper-based material. While there is no rigid differentiation between paper and paperboard, paperboard is generally thicker (usually over 0.30 mm, 0.012 in, or 12 points) than paper and has certain superior attributes ...

cellulose Cellulose is an organic compound with the formula , a polysaccharide consisting of a linear chain of several hundred to many thousands of β(1→4) linked D-glucose units. Cellulose is an important structural component of the primary cell wa ...

sponges, cellophane and some important

plastic Plastics are a wide range of synthetic or semi-synthetic materials that use polymers as a main ingredient. Their plasticity makes it possible for plastics to be moulded, extruded or pressed into solid objects of various shapes. This adapta ...

s and

textile Textile is an Hyponymy and hypernymy, umbrella term that includes various Fiber, fiber-based materials, including fibers, yarns, Staple (textiles)#Filament fiber, filaments, Thread (yarn), threads, different #Fabric, fabric types, etc. At f ...

s, such as cellulose acetate and

rayon Rayon is a semi-synthetic fiber, made from natural sources of regenerated cellulose, such as wood and related agricultural products. It has the same molecular structure as cellulose. It is also called viscose. Many types and grades of viscose ...

Bamboo Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...

stems also have hundreds of uses, including paper, buildings, furniture, boats, musical instruments,

fishing pole A fishing rod is a long, thin rod used by anglers to catch fish by manipulating a line ending in a hook (formerly known as an ''angle'', hence the term "angling"). At its most basic form, a fishing rod is a straight rigid stick/pole with a ...

water pipe Plumbing is any system that conveys fluids for a wide range of applications. Plumbing uses pipes, valves, plumbing fixtures, tanks, and other apparatuses to convey fluids. Heating and cooling (HVAC), waste removal, and potable water delive ...

s, plant stakes, and scaffolding. Trunks of

palm trees Palm most commonly refers to: * Palm of the hand, the central region of the front of the hand * Palm plants, of family Arecaceae **List of Arecaceae genera * Several other plants known as "palm" Palm or Palms may also refer to: Music * Palm (ba ...

and tree ferns are often used for building. Stems of reed are an important building material for use in

thatching Thatching is the craft of building a roof with dry vegetation such as straw, water reed, sedge (''Cladium mariscus''), rushes, heather, or palm branches, layering the vegetation so as to shed water away from the inner roof. Since the bulk of ...

in some areas.

Tannin Tannins (or tannoids) are a class of astringent, polyphenolic biomolecules that bind to and precipitate proteins and various other organic compounds including amino acids and alkaloids. The term ''tannin'' (from Anglo-Norman ''tanner'' ...

s used for tanning

leather Leather is a strong, flexible and durable material obtained from the tanning, or chemical treatment, of animal skins and hides to prevent decay. The most common leathers come from cattle, sheep, goats, equine animals, buffalo, pigs and hog ...

are obtained from the wood of certain trees, such as quebracho.

Cork Cork or CORK may refer to: Materials * Cork (material), an impermeable buoyant plant product ** Cork (plug), a cylindrical or conical object used to seal a container ***Wine cork Places Ireland * Cork (city) ** Metropolitan Cork, also known as G ...

is obtained from the bark of the cork oak.

Rubber Rubber, also called India rubber, latex, Amazonian rubber, ''caucho'', or ''caoutchouc'', as initially produced, consists of polymers of the organic compound isoprene, with minor impurities of other organic compounds. Thailand, Malaysia, and ...

is obtained from the trunks of ''

Hevea brasiliensis ''Hevea brasiliensis'', the Pará rubber tree, ''sharinga'' tree, seringueira, or most commonly, rubber tree or rubber plant, is a flowering plant belonging to the spurge family Euphorbiaceae originally native to the Amazon basin, but is now pa ...

.''

Rattan Rattan, also spelled ratan, is the name for roughly 600 species of Old World climbing palms belonging to subfamily Calamoideae. The greatest diversity of rattan palm species and genera are in the closed- canopy old-growth tropical fores ...

, used for furniture and baskets, is made from the stems of tropical vining palms. Bast fibers for textiles and rope are obtained from stems include

flax Flax, also known as common flax or linseed, is a flowering plant, ''Linum usitatissimum'', in the family Linaceae. It is cultivated as a food and fiber crop in regions of the world with temperate climates. Textiles made from flax are known in ...

hemp Hemp, or industrial hemp, is a botanical class of '' Cannabis sativa'' cultivars grown specifically for industrial or medicinal use. It can be used to make a wide range of products. Along with bamboo, hemp is among the fastest growing plants ...

jute Jute is a long, soft, shiny bast fiber that can be spun into coarse, strong threads. It is produced from flowering plants in the genus ''Corchorus'', which is in the mallow family Malvaceae. The primary source of the fiber is '' Corchorus ol ...

and ramie. The earliest paper was obtained from the stems of

papyrus Papyrus ( ) is a material similar to thick paper that was used in ancient times as a writing surface. It was made from the pith of the papyrus plant, '' Cyperus papyrus'', a wetland sedge. ''Papyrus'' (plural: ''papyri'') can also refer to ...

by the ancient Egyptians.

Amber Amber is fossilized tree resin that has been appreciated for its color and natural beauty since Neolithic times. Much valued from antiquity to the present as a gemstone, amber is made into a variety of decorative objects."Amber" (2004). In M ...

is fossilized sap from tree trunks; it is used for

jewelry Jewellery ( UK) or jewelry ( U.S.) consists of decorative items worn for personal adornment, such as brooches, rings, necklaces, earrings, pendants, bracelets, and cufflinks. Jewellery may be attached to the body or the clothes. From a w ...

and may contain ancient animals. Resins from conifer wood are used to produce

turpentine Turpentine (which is also called spirit of turpentine, oil of turpentine, terebenthene, terebinthine and (colloquially) turps) is a fluid obtained by the distillation of resin harvested from living trees, mainly pines. Mainly used as a special ...

and rosin. Tree bark is often used as a mulch and in growing media for container plants. It also can become the natural

habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...

lichen A lichen ( , ) is a composite organism that arises from algae or cyanobacteria living among filaments of multiple fungi species in a mutualistic relationship.paper birch * Twisted branches of corkscrew willow and Harry Lauder's walking stick (''Corylus avellana'' 'Contorta') * Red, peeling bark of

paperbark maple ''Acer griseum'', the paperbark maple or blood-bark maple, is a species of flowering plant in the family Sapindaceae, native to central China.Flora of China (draft)''Acer griseum''/ref> ''Acer griseum'' is found in the Chinese provinces of Gansu ...

References

External links

* {{Authority control

Stem Stem or STEM may refer to: Plant structures * Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang * Stipe (botany), a stalk to support some other structure * Stipe (mycology), the stem of a mushro ...