''Noteosuchus'' is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of basal

rhynchosaur Rhynchosaurs are a group of extinct herbivorous Triassic archosauromorph reptiles, belonging to the order Rhynchosauria. Members of the group are distinguished by their triangular skulls and elongated, beak like premaxillary bones. Rhynchosaurs ...

known from the earliest

Triassic The Triassic ( ; sometimes symbolized 🝈) is a geologic period and system which spans 50.5 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.4 Mya. The Triassic is t ...

deposits of Eastern

Cape Province The Province of the Cape of Good Hope (), commonly referred to as the Cape Province () and colloquially as The Cape (), was a province in the Union of South Africa and subsequently the Republic of South Africa. It encompassed the old Cape Co ...

South Africa South Africa, officially the Republic of South Africa (RSA), is the Southern Africa, southernmost country in Africa. Its Provinces of South Africa, nine provinces are bounded to the south by of coastline that stretches along the Atlantic O ...

. It was first named by David Meredith Seares Watson in

1912 This year is notable for Sinking of the Titanic, the sinking of the ''Titanic'', which occurred on April 15. In Albania, this leap year runs with only 353 days as the country achieved switching from the Julian to Gregorian Calendar by skippin ...

and the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

is ''Eosuchus colletti''. The generic name ''Eosuchus'' is preoccupied by the generic name of '' Eosuchus lerichei'' Dollo, 1907, a

gavialoid Gavialoidea is one of three superfamilies of crocodylians, the other two being Alligatoroidea and Crocodyloidea. Although many extinct species are known, only the gharial ''Gavialis gangeticus'' and the false gharial ''Tomistoma schlegelii'' are ...

crocodilia Crocodilia () is an order of semiaquatic, predatory reptiles that are known as crocodilians. They first appeared during the Late Cretaceous and are the closest living relatives of birds. Crocodilians are a type of crocodylomorph pseudosuchia ...

n known from northern

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

. Thus, an alternative generic name, ''Noteosuchus'', was proposed by

Robert Broom Robert Broom Fellow of the Royal Society, FRS FRSE (30 November 1866 6 April 1951) was a British- South African medical doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University ...

1925 Events January * January 1 – The Syrian Federation is officially dissolved, the State of Aleppo and the State of Damascus having been replaced by the State of Syria (1925–1930), State of Syria. * January 3 – Benito Mussolini m ...

. The generic name erected by Broom (1925) is a

compound Compound may refer to: Architecture and built environments * Compound (enclosure), a cluster of buildings having a shared purpose, usually inside a fence or wall ** Compound (fortification), a version of the above fortified with defensive struc ...

, meaning "Not ''Eosuchus''", while "Eosuchus" is derived from the name of

Eos In ancient Greek mythology and Ancient Greek religion, religion, Eos (; Ionic Greek, Ionic and Homeric Greek ''Ēṓs'', Attic Greek, Attic ''Héōs'', "dawn", or ; Aeolic Greek, Aeolic ''Aúōs'', Doric Greek, Doric ''Āṓs'') is the go ...

, the goddess of the dawn in

Greek mythology Greek mythology is the body of myths originally told by the Ancient Greece, ancient Greeks, and a genre of ancient Greek folklore, today absorbed alongside Roman mythology into the broader designation of classical mythology. These stories conc ...

, and ''suchus'',

Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...

ized from the

Greek Greek may refer to: Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group *Greek language, a branch of the Indo-European language family **Proto-Greek language, the assumed last common ancestor of all kno ...

''souchos'', an Egyptian crocodile god, thus meaning "dawn crocodile". The

specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...

, ''colletti'', honors Mr. Collett for the discovery of the

holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...

and only known specimen.

Discovery

''Noteosuchus'' is known solely from the

AM 3591, a well-preserved partial

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

skeleton A skeleton is the structural frame that supports the body of most animals. There are several types of skeletons, including the exoskeleton, which is a rigid outer shell that holds up an organism's shape; the endoskeleton, a rigid internal fra ...

, missing the head, which is currently housed at the

Albany Museum The Albany Museum, South Africa, situated in Makhanda, South Africa, is affiliated to Rhodes University and dates back to 1855,Chinsamy, Anusuya. (1997). "Albany Museum, Grahamstown, South Africa." ''Encyclopedia of Dinosaurs''. Edited by Philli ...

in South Africa. It was collected from a hill slope at Grassy Ridge locality, from the

Katberg Formation The Katberg Formation is a geological formation that is found in the Beaufort Group, a major geological group that forms part of the greater Karoo Supergroup in South Africa. The Katberg Formation is the lowermost geological formation of the Tar ...

of the

Beaufort Group The Beaufort Group is the third of the main subdivisions of the Karoo Supergroup in South Africa. It is composed of a lower Adelaide Subgroup and an upper Tarkastad Subgroup. It follows conformably after the Ecca Group and unconformably under ...

. This horizon belongs to the ''Lystrosaurus'' Assemblage Zone, dating to the early

Induan The Induan is the first age of the Early Triassic epoch in the geologic timescale, or the lowest stage of the Lower Triassic series in chronostratigraphy. It spans the time between 251.9 Ma and 249.9 Ma (million years ago). The Induan is so ...

stage of the Early Triassic

period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...

. Thus, ''Noteosuchus'' represents the geologically oldest known

species.

Classification

David M. Dilkes (1998) suggested that ''Noteosuchus colletti'' may be a

junior synonym In taxonomy, the scientific classification of living organisms, a synonym is an alternative scientific name for the accepted scientific name of a taxon. The botanical and zoological codes of nomenclature treat the concept of synonymy differently. ...

of ''

Mesosuchus browni ''Mesosuchus'' ("middle crocodile") is an extinct genus of basal rhynchosaur from early Middle Triassic (early Anisian stage) deposits of Eastern Cape, South Africa. It is known from the holotype SAM 5882, a partial skeleton, and from the para ...

'', and thus this species has been widely ignored since. However, Ezcurra, Scheyer and Butler (2014) noted that this synonymy was based only on generalized rhynchosaur

plesiomorphic In phylogenetics, a plesiomorphy ("near form") and symplesiomorphy are synonyms for an ancestral character shared by all members of a clade, which does not distinguish the clade from other clades. Plesiomorphy, symplesiomorphy, apomorphy, an ...

similarities rather than

autapomorphies In phylogenetics, an autapomorphy is a distinctive feature, known as a derived trait, that is unique to a given taxon. That is, it is found only in one taxon, but not found in any others or outgroup taxa, not even those most closely related to ...

. They found this to be unlikely because the temporal gap between the two species spans most of the

Early Triassic The Early Triassic is the first of three epochs of the Triassic Period of the geologic timescale. It spans the time between 251.9 Ma and Ma (million years ago). Rocks from this epoch are collectively known as the Lower Triassic Series, which ...

, as ''Mesosuchus'' is otherwise known only from the

Middle Triassic In the geologic timescale, the Middle Triassic is the second of three epoch (geology), epochs of the Triassic period (geology), period or the middle of three series (stratigraphy), series in which the Triassic system (stratigraphy), system is di ...

. Additionally, their

phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...

recovered ''Noteosuchus'' and ''Mesosuchus'' in a

polytomy An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree ...

with ''

Howesia browni ''Howesia'' is an extinct genus of basal rhynchosaur from early Middle Triassic (early Anisian stage) deposits of Eastern Cape, South Africa. It is known from the holotype SAM 5884, a partial skeleton with palate and partial lower jaws and from ...

'', a result which is inconsistent with such synonymy. ''Noteosuchus'' in a

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

based on Ezcurra ''et al.'' (2016):

References

{{Taxonbar, from=Q15975802 Rhynchosauria Early Triassic reptiles of Africa Induan genera Fossils of South Africa Fossil taxa described in 1912 Fossil taxa described in 1925 Prehistoric reptile genera