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''Myrtus'' (commonly called myrtle) is a
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
of
flowering plant Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s in the family
Myrtaceae Myrtaceae, the myrtle family, is a family of dicotyledonous plants placed within the order Myrtales. Myrtle, pōhutukawa, bay rum tree, clove, guava, acca (feijoa), allspice, and eucalyptus are some notable members of this group. All sp ...
. It was first described by Swedish botanist Linnaeus in 1753.Tropicos, ''Myrtus'' L.
/ref> Over 600 names have been proposed in the genus, but nearly all have either been moved to other genera or been regarded as synonyms. The genus ''Myrtus'' has three
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
recognised today: *'' Myrtus communis'' – Common myrtle; native to the Mediterranean region in southern
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
*''Myrtus nivellei'' – Saharan myrtle; native to
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
*'' Myrtus phyllireaefolia''


Description


Common myrtle

'' Myrtus communis'', the "common myrtle", is native across the Mediterranean region, Macaronesia, western Asia, and the Indian subcontinent. It is also cultivated. The plant is an
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
shrub or small tree, growing to tall. The
leaf A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ...
is entire, 3–5 cm long, with a fragrant
essential oil An essential oil is a concentrated hydrophobic liquid containing volatile (easily evaporated at normal temperatures) chemical compounds from plants. Essential oils are also known as volatile oils, ethereal oils, aetheroleum, or simply as the o ...
. The star-like
flower A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
has five petals and sepals, and numerous
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
s. Petals usually are white. The flower is pollinated by
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three pa ...
s. The fruit is a round berry containing several
seed A seed is an embryonic plant enclosed in a protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, including the gymnosperm and angiosper ...
s, most commonly blue-black in colour. A variety with yellow-amber berries is also present. The seeds are dispersed by
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s that eat the berries.


Saharan myrtle

'' Myrtus nivellei'', the Saharan myrtle, ( Tuareg language: ''tefeltest''), is
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the mountains of the central Sahara Desert.Uicnmed.org: ''Myrtus nivellei'' - Batt & Trab. - Myrtaceae
. accessed 1.10.2014.
It is found in a restricted range in the
Tassili n'Ajjer Tassili n'Ajjer ( Berber: ''Tassili n Ajjer'', ar, طاسيلي ناجر; "Plateau of rivers") is a national park in the Sahara desert, located on a vast plateau in southeastern Algeria. Having one of the most important groupings of prehistoric ...
Mountains in southern
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
, and the Tibesti Mountains in northern
Chad Chad (; ar, تشاد , ; french: Tchad, ), officially the Republic of Chad, '; ) is a landlocked country at the crossroads of North and Central Africa. It is bordered by Libya to the north, Sudan to the east, the Central African Republic ...
. It occurs in small areas of sparse relict woodland at montane elevations above the central Saharan desert plains. It is a traditional medicinal plant for the Tuareg people.


Fossil record

250
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
seeds of †''Myrtus palaeocommunis'' have been described from middle Miocene strata of the Fasterholt area near Silkeborg in Central
Jutland Jutland ( da, Jylland ; german: Jütland ; ang, Ēota land ), known anciently as the Cimbric or Cimbrian Peninsula ( la, Cimbricus Chersonesus; da, den Kimbriske Halvø, links=no or ; german: Kimbrische Halbinsel, links=no), is a peninsula of ...
,
Denmark ) , song = ( en, "King Christian stood by the lofty mast") , song_type = National and royal anthem , image_map = EU-Denmark.svg , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of Denmark , establish ...
.


Uses


Gardening

''Myrtus communis'' is widely cultivated as an
ornamental plant Ornamental plants or garden plants are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars that ...
for use as a shrub in
garden A garden is a planned space, usually outdoors, set aside for the cultivation, display, and enjoyment of plants and other forms of nature. The single feature identifying even the wildest wild garden is ''control''. The garden can incorporate bot ...
s and
park A park is an area of natural, semi-natural or planted space set aside for human enjoyment and recreation or for the protection of wildlife or natural habitats. Urban parks are green spaces set aside for recreation inside towns and cities. ...
s. It is often used as a
hedge A hedge or hedgerow is a line of closely spaced shrubs and sometimes trees, planted and trained to form a barrier or to mark the boundary of an area, such as between neighbouring properties. Hedges that are used to separate a road from adjoi ...
plant, with its small leaves shearing cleanly. When trimmed less frequently, it has numerous flowers in late summer. It requires a long hot summer to produce its flowers, and protection from winter frosts. The species and the subspecies ''M. communis'' subsp. ''tarentina'' have gained the
Royal Horticultural Society The Royal Horticultural Society (RHS), founded in 1804 as the Horticultural Society of London, is the UK's leading gardening charity. The RHS promotes horticulture through its five gardens at Wisley (Surrey), Hyde Hall (Essex), Harlow Carr (Nor ...
's
Award of Garden Merit The Award of Garden Merit (AGM) is a long-established annual award for plants by the British Royal Horticultural Society (RHS). It is based on assessment of the plants' performance under UK growing conditions. History The Award of Garden Merit ...
.


Culinary

''Myrtus communis'' is used in the islands of
Sardinia Sardinia ( ; it, Sardegna, label=Italian, Corsican and Tabarchino ; sc, Sardigna , sdc, Sardhigna; french: Sardaigne; sdn, Saldigna; ca, Sardenya, label= Algherese and Catalan) is the second-largest island in the Mediterranean Sea, aft ...
and
Corsica Corsica ( , Upper , Southern ; it, Corsica; ; french: Corse ; lij, Còrsega; sc, Còssiga) is an island in the Mediterranean Sea and one of the 18 regions of France. It is the fourth-largest island in the Mediterranean and lies southeast of ...
to produce an aromatic liqueur called ''Mirto'' by macerating it in alcohol. ''Mirto'' is one of the most typical drinks of Sardinia and comes in two varieties: ''mirto rosso'' (red) produced by macerating the berries, and ''mirto bianco'' (white) produced from the less common yellow berries and sometimes the leaves. Many Mediterranean pork dishes include myrtle berries, and roasted piglet is often stuffed with myrtle sprigs in the belly cavity, to impart an aromatic flavour to the meat. The berries, whole or ground, have been used as a pepper substitute. They contribute to the distinctive flavor of some versions of Italian Mortadella sausage and the related American
Bologna sausage Bologna sausage, informally baloney ( ), is a sausage derived from the Italian mortadella, a similar-looking, finely ground pork sausage, originally from the city of Bologna (). Typical seasonings for bologna include black pepper, nutmeg, ...
. In Calabria, a myrtle branch is threaded through dried figs and then baked. The figs acquire a pleasant taste from the essential oils of the herb. They are then enjoyed through the winter months.


Medicinal

Myrtle, along with willow tree bark, occupies a minor place in the writings of
Hippocrates Hippocrates of Kos (; grc-gre, Ἱπποκράτης ὁ Κῷος, Hippokrátēs ho Kôios; ), also known as Hippocrates II, was a Greek physician of the classical period who is considered one of the most outstanding figures in the history o ...
, Pliny,
Dioscorides Pedanius Dioscorides ( grc-gre, Πεδάνιος Διοσκουρίδης, ; 40–90 AD), “the father of pharmacognosy”, was a Greek physician, pharmacologist, botanist, and author of '' De materia medica'' (, On Medical Material) —a 5-vo ...
, Galen, and the Arabian writers. Celsus, for instance, suggested that 'soda in vinegar, or ladanum in myrtle oil and wine' could be used to treat various ailments of the scalp. It is possible that Myrtle's effect was due to high levels of salicylic acid. In several countries, particularly in Europe and China, there has been a tradition for prescribing this substance for sinus infections. A systematic review of herbal medicines used for the treatment of rhinosinusitis concluded that the evidence that any herbal medicines are beneficial in the treatment of rhinosinusitis is limited, and that for ''Myrtus'' there is insufficient data to verify the significance of clinical results.


In myth and ritual


Classical

In
Greek mythology A major branch of classical mythology, Greek mythology is the body of myths originally told by the ancient Greeks, and a genre of Ancient Greek folklore. These stories concern the origin and nature of the world, the lives and activities o ...
and ritual the myrtle was sacred to the goddesses
Aphrodite Aphrodite ( ; grc-gre, Ἀφροδίτη, Aphrodítē; , , ) is an ancient Greek goddess associated with love, lust, beauty, pleasure, passion, and procreation. She was syncretized with the Roman goddess . Aphrodite's major symbols incl ...
and also
Demeter In ancient Greek religion and Greek mythology, mythology, Demeter (; Attic Greek, Attic: ''Dēmḗtēr'' ; Doric Greek, Doric: ''Dāmā́tēr'') is the Twelve Olympians, Olympian goddess of the harvest and agriculture, presiding over crops, ...
:
Artemidorus Artemidorus Daldianus ( grc-gre, Ἀρτεμίδωρος ὁ Δαλδιανός) or Ephesius was a professional diviner who lived in the 2nd century AD. He is known from an extant five-volume Greek work, the ''Oneirocritica'' or ''Oneirokritikon' ...
asserts that in interpreting dreams "a myrtle garland signifies the same as an olive garland, except that it is especially auspicious for farmers because of Demeter and for women because of Aphrodite. For the plant is sacred to both goddesses." Pausanias explains that one of the Graces in the sanctuary at Elis holds a myrtle branch because "the rose and the myrtle are sacred to Aphrodite and connected with the story of Adonis, while the Graces are of all deities the nearest related to Aphrodite." Myrtle is the garland of Iacchus, according to
Aristophanes Aristophanes (; grc, Ἀριστοφάνης, ; c. 446 – c. 386 BC), son of Philippus, of the deme Kydathenaion ( la, Cydathenaeum), was a comic playwright or comedy-writer of ancient Athens and a poet of Old Attic Comedy. Eleven of his ...
, and of the victors at the Theban ''Iolaea'', held in honour of the Theban hero
Iolaus In Greek mythology, Iolaus (; Ancient Greek: Ἰόλαος ''Iólaos'') was a Theban divine hero. He was famed for being Heracles' nephew and for helping with some of his Labors, and also for being one of the Argonauts. Family Iolaus was ...
. In Rome, Virgil explains that "the poplar is most dear to Alcides, the vine to Bacchus, the myrtle to lovely Venus, and his own laurel to Phoebus." At the Veneralia, women bathed wearing crowns woven of myrtle branches, and myrtle was used in wedding rituals. In the
Aeneid The ''Aeneid'' ( ; la, Aenē̆is or ) is a Latin epic poem, written by Virgil between 29 and 19 BC, that tells the legendary story of Aeneas, a Trojan who fled the fall of Troy and travelled to Italy, where he became the ancestor of ...
, myrtle marks the grave of the murdered Polydorus in
Thrace Thrace (; el, Θράκη, Thráki; bg, Тракия, Trakiya; tr, Trakya) or Thrake is a geographical and historical region in Southeast Europe, now split among Bulgaria, Greece, and Turkey, which is bounded by the Balkan Mountains to ...
.
Aeneas In Greco-Roman mythology, Aeneas (, ; from ) was a Trojan hero, the son of the Trojan prince Anchises and the Greek goddess Aphrodite (equivalent to the Roman Venus). His father was a first cousin of King Priam of Troy (both being grandsons ...
' attempts to uproot the shrub cause the ground to bleed, and the voice of the dead Polydorus warns him to leave. The spears which impaled Polydorus have been magically transformed into the myrtle which marks his grave.


Jewish

In Jewish liturgy, the myrtle is one of the four sacred plants ( Four Species) of
Sukkot or ("Booths, Tabernacles") , observedby = Jews, Samaritans, a few Protestant denominations, Messianic Jews, Semitic Neopagans , type = Jewish, Samaritan , begins = 15th day of Tishrei , ends = 21st day of Tis ...
, the Feast of Tabernacles representing the different types of personality making up the community. The myrtle having fragrance but not pleasant taste, represents those who have good deeds to their credit despite not having knowledge from
Torah The Torah (; hbo, ''Tōrā'', "Instruction", "Teaching" or "Law") is the compilation of the first five books of the Hebrew Bible, namely the books of Genesis, Exodus, Leviticus, Numbers and Deuteronomy. In that sense, Torah means the ...
study. The three branches are lashed or braided together by the worshipers a palm leaf, a willow bough, and a myrtle branch. The
etrog Etrog ( he, אֶתְרוֹג, plural: '; Ashkenazi Hebrew: ', plural: ') is the yellow citron or ''Citrus medica'' used by Jews during the week-long holiday of Sukkot as one of the four species. Together with the ''lulav'', ''hadass'', and '' a ...
or citron is the fruit held in the other hand as part of the
lulav ''Lulav'' (; he, לולב) is a closed frond of the date palm tree. It is one of the Four Species used during the Jewish holiday of Sukkot. The other Species are the ''hadass'' (Myrtus, myrtle), '' aravah'' ( willow), and ''etrog'' (citron). ...
wave ritual. In Jewish mysticism, the myrtle represents the phallic, masculine force at work in the universe. For this reason myrtle branches were sometimes given the bridegroom as he entered the nuptial chamber after a wedding (Tos. Sotah 15:8; Ketubot 17a). Myrtles are both the symbol and scent of Eden (BhM II: 52; Sefer ha-Hezyonot 17). The
Hechalot The Hekhalot literature (sometimes transliterated Heichalot) from the Hebrew word for "Palaces", relating to visions of ascents into heavenly palaces. The genre overlaps with ''Merkabah'' or "Chariot" literature, concerning Ezekiel's chariot, so t ...
text Merkavah Rabbah requires one to suck on a myrtle leaves as an element of a theurgic ritual. Kabbalists link myrtle to the sefirah of Tiferet and use sprigs in their Shabbat (especially Havdalah) rites to draw down its harmonizing power as the week is initiated (Shab. 33a; Zohar Chadash, SoS, 64d; Sha’ar ha-Kavvanot, 2, pp. 73–76). Myrtle leaves were added to the water in the last (7th) rinsing of the head in the traditional Sephardic tahara manual (teaching the ritual for washing the dead). Myrtles are often used to recite a blessing over a fragrant plant during the
Havdalah Havdalah ( he, הַבְדָּלָה, "separation") is a Jewish religious ceremony that marks the symbolic end of Shabbat and ushers in the new week. The ritual involves lighting a special havdalah candle with several wicks, blessing a cup of ...
ceremony, as well as before Kiddush is some Sefardic and
Hasidic Hasidism, sometimes spelled Chassidism, and also known as Hasidic Judaism ( Ashkenazi Hebrew: חסידות ''Ḥăsīdus'', ; originally, "piety"), is a Jewish religious group that arose as a spiritual revival movement in the territory of conte ...
traditions.


Mandaean

In the Mandaean religion, myrtle wreaths ('' klila'') are used by priests in important religious rituals and ceremonies, such as
baptism Baptism (from grc-x-koine, βάπτισμα, váptisma) is a form of ritual purification—a characteristic of many religions throughout time and geography. In Christianity, it is a Christian sacrament of initiation and adoption, almost ...
and death masses ('' masiqta''). Myrtle wreaths also form part of the ''
darfash The drabshaDrower, Ethel Stefana (1937). ''The Mandaeans of Iraq and Iran''. Oxford at the Clarendon Press. ( myz, ࡃࡓࡀࡁࡔࡀ) or darfash ( ar, درفش) is the symbol of the Mandaean faith. It is typically translated as 'banner'. Etymo ...
'', the official symbol of
Mandaeism Mandaeism (Classical Mandaic: ࡌࡀࡍࡃࡀࡉࡉࡀ ; Arabic: المندائيّة ), sometimes also known as Nasoraeanism or Sabianism, is a Gnostic, monotheistic and ethnic religion. Its adherents, the Mandaeans, revere Adam, Abe ...
consisting of an olive wooden cross covered with a white silk cloth.


Contemporary

In neo-pagan and wicca rituals, myrtle, though not indigenous beyond the Mediterranean Basin, is now commonly associated with and sacred to Beltane ( May Day). Myrtle in a wedding bouquet is a general European custom. A sprig of myrtle from
Queen Victoria Victoria (Alexandrina Victoria; 24 May 1819 – 22 January 1901) was Queen of the United Kingdom of Great Britain and Ireland from 20 June 1837 until her death in 1901. Her reign of 63 years and 216 days was longer than that of any previ ...
's wedding bouquet was planted as a slip, and sprigs from it have continually been included in royal wedding bouquets.


Garden history


Rome

Because of its elegance of habit, appealing odour, and amenity to clipping by the ''
topiarius Topiary is the horticultural practice of training perennial plants by clipping the foliage and twigs of trees, shrubs and subshrubs to develop and maintain clearly defined shapes, whether geometric or fanciful. The term also refers to plants w ...
'', as much as for sacred associations, the myrtle was an indispensable feature of Roman gardens. As a reminder of home, it will have been introduced wherever Roman elites were settled, even in areas of the Mediterranean Basin where it was not already endemic: "the Romans... must surely have attempted to establish a shrub so closely associated with their mythology and tradition," observes Alice Coats. In
Gaul Gaul ( la, Gallia) was a region of Western Europe first described by the Romans. It was inhabited by Celtic and Aquitani tribes, encompassing present-day France, Belgium, Luxembourg, most of Switzerland, parts of Northern Italy (only during ...
and
Britannia Britannia () is the national personification of Britain as a helmeted female warrior holding a trident and shield. An image first used in classical antiquity, the Latin ''Britannia'' was the name variously applied to the British Isles, Grea ...
it will not have proved hardy.


England

In England it was reintroduced in the 16th century, traditionally with the return from Spain in 1585 of Sir Walter Raleigh, who also brought with him the first orange trees seen in England. ''Myrtus communis'' will have needed similar protection from winter cold and wet. Alice Coats notes an earlier testimony: in 1562
Queen Elizabeth I Elizabeth I (7 September 153324 March 1603) was Queen of England and Ireland from 17 November 1558 until her death in 1603. Elizabeth was the last of the five House of Tudor monarchs and is sometimes referred to as the "Virgin Queen". Eli ...
's great minister Lord Burghley wrote to Mr Windebank in Paris to ask him for a lemon, a pomegranate and a myrtle, with instructions for their culture—which suggests that the myrtle, like the others, was not yet familiar. By 1597
John Gerard John Gerard (also John Gerarde, c. 1545–1612) was an English herbalist with a large garden in Holborn, now part of London. His 1,484-page illustrated ''Herball, or Generall Historie of Plantes'', first published in 1597, became a popular gar ...
lists six varieties being grown in southern England,Gerard, ''The Herball'', 1597. and by 1640 John Parkinson noted a double-flowering one. Alice Coats suggests that this was the very same double that the diarist and gardener
John Evelyn John Evelyn (31 October 162027 February 1706) was an English writer, landowner, gardener, courtier and minor government official, who is now best known as a diarist. He was a founding Fellow of the Royal Society. John Evelyn's diary, or m ...
noted "was first discovered by the incomparable
Nicolas-Claude Fabri de Peiresc Nicolas-Claude Fabri de Peiresc (1 December 1580 – 24 June 1637), often known simply as Peiresc, or by the Latin form of his name, Peirescius, was a French astronomer, antiquary and savant, who maintained a wide correspondence with scientis ...
, which a mule had cropt from a wild shrub." In the late 17th and early 18th centuries myrtles in cases, pots and tubs were brought out to summer in the garden and wintered with other tender greens in an orangery. Fairchild, ''The City Gardener'' (1722) notes their temporary use, rented from a nurseryman annually to fill an empty fireplace in the warm months. With the influx to England of more dramatic tender plants and shrubs from Japan or Peru in the 19th century, it was more difficult to find room for the common myrtle of borderline hardiness.


Related plants

Many other related plants native to
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
,
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
and elsewhere, previously classified in a wider interpretation of the genus ''Myrtus'', are now species within other genera, including: '' Eugenia,
Lophomyrtus ''Lophomyrtus'' is a genus of the myrtle family described as a genus in 1941. The entire genus is endemic to New Zealand. It consists of evergreen shrubs or trees, noted for their colorful leaves, which are purple, chocolate, red or bronze-gree ...
, Luma,
Rhodomyrtus ''Rhodomyrtus'' is a group of shrubs and trees in the family Myrtaceae described as a genus in 1841. The genus is native to southern China, the Indian Subcontinent, Southeast Asia, Melanesia, and Australia. Its greatest levels of diversity are ...
, Syzygium, Ugni'', and at least a dozen other genera. The name "myrtle" is also used in common names (vernacular names) of unrelated plants in several other genera, such as: "Crepe myrtle" (''
Lagerstroemia ''Lagerstroemia'' (), commonly known as crape myrtle (also spelled crepe myrtle or crêpe myrtle), is a genus of around 50 species of deciduous and evergreen trees and shrubs native to the Indian subcontinent, southeast Asia, northern Aust ...
'' species and hybrids, Lythraceae); "Wax myrtle" ('' Morella'' species, Myricaceae); and "Creeping myrtle" ('' Vinca'' species,
Apocynaceae Apocynaceae (from '' Apocynum'', Greek for "dog-away") is a family of flowering plants that includes trees, shrubs, herbs, stem succulents, and vines, commonly known as the dogbane family, because some taxa were used as dog poison Members ...
).


References


External links


Myrtle (''Myrtus communis'' L.)
from Gernot Katzer's Spice Pages
www.myrtus-communis.de
(German)
Myrtus in Flora Europaea
{{Taxonbar, from=Q165152 Myrtaceae genera Matorral shrubland Medicinal plants Plants used in traditional African medicine Garden plants of Europe Garden plants of Africa Drought-tolerant plants Bird food plants Shrubs Taxa named by Carl Linnaeus