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__NOTOC__ Mrs. Ples is the popular nickname for the most complete skull of an ''
Australopithecus africanus ''Australopithecus africanus'' is an extinct species of australopithecine which lived between about 3.3 and 2.1 million years ago in the Late Pliocene to Early Pleistocene of South Africa. The species has been recovered from Taung, Sterkfontei ...
'' ever found in South Africa. Many ''Australopithecus'' fossils have been found near
Sterkfontein Sterkfontein (Afrikaans for ''Strong Spring'') is a set of limestone caves of special interest to paleo-anthropologists located in Gauteng province, about northwest of Johannesburg, South Africa in the Muldersdrift area close to the town of ...
, about northwest of Johannesburg, in a region of
Gauteng Gauteng ( ) is one of the nine provinces of South Africa. The name in Sotho-Tswana languages means 'place of gold'. Situated on the Highveld, Gauteng is the smallest province by land area in South Africa. Although Gauteng accounts for only ...
(part of the old Transvaal) now designated as the
Cradle of Humankind The Cradle of Humankind is a paleoanthropological site and is located about northwest of Johannesburg, South Africa, in the Gauteng province. Declared a World Heritage Site by UNESCO in 1999, the site is home to the largest concentration of h ...
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for ...
. Mrs. Ples was discovered by
Robert Broom Robert Broom FRS FRSE (30 November 1866 6 April 1951) was a British- South African doctor and palaeontologist. He qualified as a medical practitioner in 1895 and received his DSc in 1905 from the University of Glasgow. From 1903 to 1910, he ...
and John T. Robinson on April 18, 1947. Because of Broom's use of dynamite and pickaxe while excavating, Mrs. Ples's skull was blown into pieces and some fragments are missing. Nonetheless, Mrs./Mr. Ples is one of the most "perfect" pre-human skulls ever found. The skull is currently held at the
Ditsong National Museum of Natural History The Ditsong National Museum of Natural History, formerly the Transvaal Museum, is a natural history museum situated in Pretoria, South Africa. It is located on Paul Kruger Street, between Visagie and Minnaar Streets, opposite the Pretoria City Ha ...
in
Pretoria Pretoria () is South Africa's administrative capital, serving as the seat of the executive branch of government, and as the host to all foreign embassies to South Africa. Pretoria straddles the Apies River and extends eastward into the foothi ...
. The nickname "Mrs. Ples" was coined by Broom's young co-workers. It derives from the scientific name ''Plesianthropus transvaalensis'' (near-man from the Transvaal), that Broom initially gave the skull, later subsumed ( synonymized) into the species ''Australopithecus africanus.'' In scientific publications the specimen is referred to by its catalogue number, STS 5. The genus ''
Australopithecus ''Australopithecus'' (, ; ) is a genus of early hominins that existed in Africa during the Late Pliocene and Early Pleistocene. The genus ''Homo'' (which includes modern humans) emerged within ''Australopithecus'', as sister to e.g. ''Australo ...
'', of which there are several species, is considered the likely precursor of the genus ''
Homo ''Homo'' () is the genus that emerged in the (otherwise extinct) genus '' Australopithecus'' that encompasses the extant species ''Homo sapiens'' ( modern humans), plus several extinct species classified as either ancestral to or closely rela ...
'', to which all humanity belongs. Though its
cranium The skull is a bone protective cavity for the brain. The skull is composed of four types of bone i.e., cranial bones, facial bones, ear ossicles and hyoid bone. However two parts are more prominent: the cranium and the mandible. In humans, th ...
is comparable to a
chimpanzee The chimpanzee (''Pan troglodytes''), also known as simply the chimp, is a species of great ape native to the forest and savannah of tropical Africa. It has four confirmed subspecies and a fifth proposed subspecies. When its close relative t ...
's, ''Australopithecus'' walked upright, as humans do. This was a surprise to anthropologists at the time, because it had been assumed that the big brain of ''Homo'' had preceded, or at least evolved in tandem with, our upright gait. Mrs. Ples, whose cranial capacity is only about , was one of the first fossils to reveal that upright walking had evolved well before any significant growth in brain size. The sex of the specimen is not completely certain and so ''Mrs. Ples'' may in fact be ''Mr. Ples''. Moreover, X-ray analysis of the specimen's teeth (''see below'') has suggested it was an adolescent. Hence a designation of ''
Miss Miss (pronounced ) is an English language honorific typically used for a girl, for an unmarried woman (when not using another title such as " Doctor" or "Dame"), or for a married woman retaining her maiden name. Originating in the 17th century, i ...
Ples'' or ''
Master Master or masters may refer to: Ranks or titles * Ascended master, a term used in the Theosophical religious tradition to refer to spiritually enlightened beings who in past incarnations were ordinary humans *Grandmaster (chess), National Master ...
Ples'' is also possible. The paleoanthropologist, Prof.
Frederick E. Grine Frederick Edward Grine is an American paleoanthropologist. He is a Professor of anthropology and anatomical sciences at the State University of New York at Stony Brook. He received his bachelor's degree from Washington & Jefferson College, and his ...
, has studied the dental morphology of Mrs./Mr. Ples with a view to finally establishing Mrs./Ms. Ples's sex. Using the Computed Tomography (CT) scans of STS 5 from the experiments of Weber et al., (2012) they compared them to CT scans of more recently discovered ''A. africanus'' skulls from Sterkfontein. These scans allowed Grine to reconstruct the roots of the teeth, in order to see how the molar and canine teeth developed. This study concluded that Mrs./Mr. Ples was indeed a middle-aged female. However, the question is not entirely settled, since other studies have come to the opposite conclusion. Some experts have suggested that a partial skeleton, known only by its catalogue number of STS 14, which was discovered in the same year, in the same geological deposit and in proximity to Mrs. Ples, may belong to this skull. If correct, this would make Mrs. Ples the South African counterpart to the famous
Lucy Lucy is an English feminine given name derived from the Latin masculine given name Lucius with the meaning ''as of light'' (''born at dawn or daylight'', maybe also ''shiny'', or ''of light complexion''). Alternative spellings are Luci, Luce, Lu ...
fossil. This skull, along with others discovered at
Taung Taung is a small town situated in the North West Province of South Africa. The name means ''place of the lion'' and was named after Tau, the King of the Barolong. ''Tau'' is the Tswana word for lion. Education High,Secondary and Middle Schools ...
, Sterkfontein, and
Makapansgat Makapansgat () (or Makapan Valley World Heritage Site) is an archaeological location within the Makapansgat and Zwartkrans Valleys, northeast of Mokopane in Limpopo province, South Africa. It is an important palaeontological site, with the local ...
offered compelling evidence in favour of Charles Darwin's hypothesis that humanity's origin lay in Africa. The fossil was originally dated by a combination of palaeomagnetism and uranium-lead techniques to 2.1-2.6 million years ago. However, as of 2022 the specimen is believed to be 3.4-3.7 million years old after surrounding rocks were dated In 2004, Mrs. Ples was voted 95th in the
SABC 3 SABC 3 (stylised as S3) is a South African free-to-air public television network owned by the South African Broadcasting Corporation (SABC). It carries programming in English and, few in other South African languages. It has a number of its own r ...
's and e.tv's '' Great South Africans'' Top 100 list.


See also

*
List of human evolution fossils The following tables give an overview of notable finds of hominin fossils and remains relating to human evolution, beginning with the formation of the tribe Hominini (the divergence of the human and chimpanzee lineages) in the late Miocene, roug ...
*
Taung Child The Taung Child (or Taung Baby) is the fossilised skull of a young '' Australopithecus africanus''. It was discovered in 1924 by quarrymen working for the Northern Lime Company in Taung, South Africa. Raymond Dart described it as a new specie ...


References


Further reading

*


External links


Maropeng - The Cradle of Humankind
official website
UNESCO - Fossil Hominid Sites of Sterkfontein, Swartkrans, Kromdraai, and EnvironsMetadata of STS 5 on NESPOS
{{Cradle of Humankind History of South Africa Australopithecus fossils Quaternary fossil record 1947 archaeological discoveries Archaeological discoveries in South Africa Archaeology of Southern Africa