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The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
in the
deer family Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
. Most adult male moose have distinctive broad, palmate ("open-hand shaped") antlers; most other members of the deer family have antlers with a dendritic ("twig-like") configuration. Moose typically inhabit boreal forests and temperate broadleaf and mixed forests of the Northern Hemisphere in temperate to
subarctic climate The subarctic climate (also called subpolar climate, or boreal climate) is a climate with long, cold (often very cold) winters, and short, warm to cool summers. It is found on large landmasses, often away from the moderating effects of an ocean, g ...
s. Hunting and other human activities have caused a reduction in the size of the moose's range over time. It has been reintroduced to some of its former habitats. Currently, most moose occur in
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
,
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
,
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
(with
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...
having the most of the lower 48 states),
New York State New York, officially the State of New York, is a state in the Northeastern United States. It is often called New York State to distinguish it from its largest city, New York City. With a total area of , New York is the 27th-largest U.S. sta ...
, Fennoscandia, the Baltic states,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
, Kazakhstan, and
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
. Its diet consists of both terrestrial and aquatic vegetation. Predators of moose include wolves, bears, humans and (rarely)
wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for " glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is a musc ...
s. Unlike most other deer species, moose do not form herds and are solitary animals, aside from calves who remain with their mother until the cow begins estrus (typically at 18 months after birth of the calf), at which point the cow chases them away. Although generally slow-moving and sedentary, moose can become aggressive, and move quickly if angered or startled. Their mating season in the autumn features energetic fights between males competing for a female.


Etymology and naming

''Alces alces'' is called a "moose" in North American English, but an "elk" in British English. The word "elk" in North American English refers to a completely different species of deer, ''Cervus canadensis'', also called the
wapiti The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The commo ...
. A mature male moose is called a bull, a mature female a cow, and an immature moose of either sex a calf. According to the ''
Oxford English Dictionary The ''Oxford English Dictionary'' (''OED'') is the first and foundational historical dictionary of the English language, published by Oxford University Press (OUP). It traces the historical development of the English language, providing a c ...
'', the etymology of the species is "of obscure history". In
Classical Antiquity Classical antiquity (also the classical era, classical period or classical age) is the period of cultural history between the 8th century BC and the 5th century AD centred on the Mediterranean Sea, comprising the interlocking civilizations of ...
, the animal was known as ''álkē'' in Greek and in
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
, words probably borrowed from a Germanic language or another language of northern Europe. By the 8th century, during the
Early Middle Ages The Early Middle Ages (or early medieval period), sometimes controversially referred to as the Dark Ages, is typically regarded by historians as lasting from the late 5th or early 6th century to the 10th century. They marked the start of the Mi ...
, the species was known as derived from the
Proto-Germanic Proto-Germanic (abbreviated PGmc; also called Common Germanic) is the reconstructed proto-language of the Germanic branch of the Indo-European languages. Proto-Germanic eventually developed from pre-Proto-Germanic into three Germanic br ...
: ''*elho-'', ''*elhon-'' and possibly connected with the . Later, the species became known in
Middle English Middle English (abbreviated to ME) is a form of the English language that was spoken after the Norman conquest of 1066, until the late 15th century. The English language underwent distinct variations and developments following the Old Englis ...
as ''elk'', ''elcke'', or ''elke'', appearing in the Latinized form ''alke'', with the spelling ''alce'' borrowed directly from . Noting that ''elk'' "is not the normal phonetic representative" of the Old English ''elch'', the ''Oxford English Dictionary'' derives ''elk'' from , itself from . The word "elk" has cognates in other
Indo-European languages The Indo-European languages are a language family native to the overwhelming majority of Europe, the Iranian plateau, and the northern Indian subcontinent. Some European languages of this family, English, French, Portuguese, Russian, D ...
, e.g. ''elg'' in Danish/ Norwegian; ''älg'' in Swedish; ''alnis'' in Latvian; ''Elch'' in
German German(s) may refer to: * Germany (of or related to) **Germania (historical use) * Germans, citizens of Germany, people of German ancestry, or native speakers of the German language ** For citizens of Germany, see also German nationality law **Ge ...
; and ''łoś'' in Polish. In the continental European languages, these forms of the word "elk" always refer to ''Alces alces''. The youngest elk bones in
Great Britain Great Britain is an island in the North Atlantic Ocean off the northwest coast of continental Europe. With an area of , it is the largest of the British Isles, the largest European island and the ninth-largest island in the world. It ...
were found in Scotland and are roughly 3,900 years old. The elk was probably extinct on the island before 900 AD. The word "elk" remained in usage because of English-speakers' familiarity with the species in Continental Europe; however, without any living animals around to serve as a reference, the meaning became rather vague, and by the 17th century "elk" had a meaning similar to "large deer". Dictionaries of the 18th century simply described "elk" as a deer that was "as large as a horse". Confusingly, the word "elk" is used in North America to refer to a different animal, ''
Cervus canadensis The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The com ...
'', which is also called by the Algonquian indigenous name, "wapiti". The British began colonizing America in the 17th century, and found two common species of deer for which they had no names. The wapiti appeared very similar to the red deer of Europe (which itself was then almost extinct in Southern Britain) although it was much larger and was not red; the two species are indeed closely related, though distinct behaviorally and genetically. The moose was a rather strange-looking deer to the colonists, and they often adopted local names for both. In the early days of American colonization, the wapiti was often called a gray moose and the moose was often called a black moose, but early accounts of the animals varied wildly, adding to the confusion. The word "moose" had first entered English by 1606 and is borrowed from the Algonquian languages (compare the Narragansett ''moos'' and Eastern Abenaki ''mos''; according to early sources, these were likely derived from ''moosu'', meaning "he strips off"), and possibly involved forms from multiple languages mutually reinforcing one another. The Proto-Algonquian form was ''*mo·swa''. Early European explorers in North America, particularly in Virginia where there were no moose, called the wapiti "elk" because of its size and resemblance to familiar-looking deer like the red deer.''The Book of Animal Ignorance: Everything You Think You Know Is Wrong'' By John Mitchinson, John Lloyd -- Harmony Books 2007 Page 141 The moose resembled the "German elk" (the moose of continental Europe), which was less familiar to the British colonists. For a long time neither species had an official name, but were called a variety of things. Eventually, in North America the wapiti became known as an elk while the moose retained its indigenous name. In 1736, Samuel Dale wrote to the Royal Society of Great Britain:
The common light-grey moose, called by the Indians, Wampoose, and the large or black-moose, which is the beast whose horns I herewith present. As to the grey moose, I take it to be no larger than what Mr. John Clayton, in his account of the Virginia Quadrupeds, calls the Elke ... was in all respects like those of our red-deer or stags, only larger ... The black moose is (by all that have hitherto writ of it) accounted a very large creature. ... The stag, buck, or male of this kind has a palmed horn, not like that of our common or fallow-deer, but the palm is much longer, and more like that of the ''German elke.''


Description and anatomy


Antlers

Bull moose have antlers like other members of the deer family. Cows select mates based on antler size. Bull moose use dominant displays of antlers to discourage competition and will spar or fight rivals. The size and growth rate of antlers is determined by diet and age; symmetry reflects health. The male's antlers grow as cylindrical beams projecting on each side of the head at right angles to the midline of the skull, and then fork. The lower prong of this fork may be either simple, or divided into two or three tines, with some flattening. Most moose have antlers that are broad and palmate (flat) with tines (points) along the outer edge. Within the ecologic range of the moose in Europe, those in northerly locales display the palmate pattern of antlers, while the antlers of European moose over the southerly portion of its range are typically of the cervina dendritic pattern and comparatively small, perhaps due to evolutionary pressures of hunting by humans, who prize the large palmate antlers. European moose with antlers intermediate between the palmate and the dendritic form are found in the middle of the north–south range. Moose with antlers have more acute hearing than those without antlers; a study of trophy antlers using a microphone found that the palmate antler acts as a parabolic reflector, amplifying sound at the moose's ear. The antlers of mature Alaskan adult bull moose (5 to 12 years old) have a normal maximum spread greater than . By the age of 13, moose antlers decline in size and symmetry. The widest spread recorded was across. An Alaskan moose also holds the record for the heaviest weight at . Antler beam diameter, not the number of tines, indicates age. In North America, moose (''A. a. americanus'') antlers are usually larger than those of Eurasian moose and have two lobes on each side, like a butterfly. Eurasian moose antlers resemble a seashell, with a single lobe on each side. In the North Siberian moose (''A. a. bedfordiae''), the posterior division of the main fork divides into three tines, with no distinct flattening. In the common moose (''A. a. alces'') this branch usually expands into a broad palmation, with one large tine at the base and a number of smaller snags on the free border. There is, however, a
Scandinavia Scandinavia; Sámi languages: /. ( ) is a subregion in Northern Europe, with strong historical, cultural, and linguistic ties between its constituent peoples. In English usage, ''Scandinavia'' most commonly refers to Denmark, Norway, and S ...
n breed of the common moose in which the antlers are simpler and recall those of the East Siberian animals. The palmation appears to be more marked in North American moose than in the typical Scandinavian moose. After the mating season males drop their antlers to conserve energy for the winter. A new set of antlers will then regrow in the spring. Antlers take three to five months to fully develop, making them one of the fastest growing animal organs. Antler growth is "nourished by an extensive system of blood vessels in the skin covering, which contains numerous hair follicles that give it a 'velvet' texture." This requires intense grazing on a highly-nutritious diet. By September the velvet is removed by rubbing and thrashing which changes the colour of the antlers. Immature bulls may not shed their antlers for the winter, but retain them until the following spring. Birds, carnivores and rodents eat dropped antlers as they are full of protein and moose themselves will eat antler velvet for the nutrients. If a bull moose is castrated, either by accidental or chemical means, he will shed his current set of antlers within two weeks and then immediately begin to grow a new set of misshapen and deformed antlers that he will wear the rest of his life without ever shedding again; similarly deformed antlers can result from a deficiency of testosterone caused by
cryptorchidism Cryptorchidism, also known as undescended testis, is the failure of one or both testes to descend into the scrotum. The word is from Greek () 'hidden' and () 'testicle'. It is the most common birth defect of the male genital tract. About 3% of ...
or old age. These deformed antlers are composed of living bone which is still growing or able to grow, since testosterone is needed to stop antler growth; they may take one of two forms. "Cactus antlers" or velericorn antlers usually retain the approximate shape of a normal moose's antlers but have numerous pearl-shaped exostoses on their surface; being made of living bone, they are easily broken but can grow back. Perukes () are constantly growing, tumor-like antlers with a distinctive appearance similar to coral. Like
roe deer The roe deer (''Capreolus capreolus''), also known as the roe, western roe deer, or European roe, is a species of deer. The male of the species is sometimes referred to as a roebuck. The roe is a small deer, reddish and grey-brown, and well-adapt ...
, moose are more likely to develop perukes, rather than cactus antlers, than the more developed cervine deer, but unlike roe deer, moose do not suffer fatal decalcification of the skull as a result of peruke growth, but rather can support their continued growth until they become too large to be fully supplied with blood. The distinctive-looking perukes (often referred to as "devil's antlers") are the source of several myths and legends among many groups of
Inuit Inuit (; iu, ᐃᓄᐃᑦ 'the people', singular: Inuk, , dual: Inuuk, ) are a group of culturally similar indigenous peoples inhabiting the Arctic and subarctic regions of Greenland, Labrador, Quebec, Nunavut, the Northwest Territorie ...
as well as several other tribes of indigenous peoples of North America. In extremely rare circumstances, a cow moose may grow antlers. This is usually attributed to a hormone imbalance.


Proboscis and olfaction

The moose proboscis is distinctive among the living cervids due to its large size; it also features nares that can be sealed shut when the moose is browsing aquatic vegetation. The moose proboscis likely evolved as an adaptation to aquatic browsing, with loss of the rhinarium, and development of a superior olfactory column separate from an inferior respiratory column.Pagano, Anthony Santino, Jeffrey T. Laitman, Kurt Albertine, and Samuel Marquez. "Evolution of the proboscis in the moose, Alces alces: Evidence from Morphology and Ecology." The FASEB Journal 33, no. 1_supplement (2019): 767-19. This separation contributes to the moose's keen sense of smell, which they employ to detect water sources, to find food under snow, and to detect mates or predators.


Hooves

As with all members of the order Artiodactyla (
even-toed ungulates The even-toed ungulates (Artiodactyla , ) are ungulates—hoofed animals—which bear weight equally on two (an even number) of their five toes: the third and fourth. The other three toes are either present, absent, vestigial, or pointing poste ...
), moose feet have two large keratinized hooves corresponding to the third and fourth toe, with two small posterolateral dewclaws (vestigial digits), corresponding to the second and fifth toe. The hoof of the fourth digit is broader than that of the third digit, while the inner hoof of the third digit is longer than that of the fourth digit. This foot configuration may favor striding on soft ground. The moose hoof splays under load, increasing surface area, which limits sinking of the moose foot into soft ground or snow, and which increases efficiency when swimming. The body weight per footprint surface area of the moose foot is intermediate between that of the pronghorn foot, (which have stiff feet lacking dewclaws—optimized for high-speed running) and the caribou foot (which are more rounded with large dewclaws, optimized for walking in deep snow). The moose's body weight per surface area of footprint is about twice that of the caribou. On firm ground, a bull moose leaves a visible impression of the dewclaws in its footprint, while a cow moose or calf does not leave a dewclaw impression. On soft ground or mud, bull, cow, and calf footprints may all show dewclaw impressions.


Skin and Fur

Moose skin is typical of the
deer family Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindeer ...
. Moose fur consists of four types of hair: eyelashes, vibrissae, guard hairs and wool hairs. Hair length and hair density varies according to season, age, and body region. The coat has two layers—a top layer of long guard hairs and a soft wooly undercoat. The guard hairs are hollow and filled with air for better insulation, which also helps them stay afloat when swimming.


Dewlap

Both male and female moose have a dewlap or bell, which is a fold of skin under the chin. Its exact function is unknown, but some morphologic analyses suggest a cooling (thermoregulatory) function. Other theories include a fitness signal in mating, as a visual and olfactory signal, or as a dominance signal by males, as are the antlers.


Tail

The tail is short (6 cm to 8 cm in length) and vestigial in appearance; unlike other ungulates the moose tail is too short to swish away insects.


Size

On average, an adult moose stands high at the shoulder, which is more than higher than the next-largest deer on average, the
wapiti The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The commo ...
. Males (or "bulls") normally weigh from and females (or "cows") typically weigh , depending on racial or clinal as well as individual age or nutritional variations. The head-and-body length is , with the vestigial tail adding only a further . The largest of all the races is the Alaskan subspecies (''A. a. gigas''), which can stand over at the shoulder, has a span across the antlers of and averages in males and in females. Typically, however, the antlers of a mature bull are between . The largest confirmed size for this species was a bull shot at the Yukon River in September 1897 that weighed and measured high at the shoulder.Wood, ''The Guinness Book of Animal Facts and Feats''. Sterling Pub Co Inc. (1983), There have been reported cases of even larger moose, including a bull killed in 2004 that weighed , and a bull that reportedly scaled , but none are authenticated and some may not be considered reliable. Among extant terrestrial animal species in
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
,
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, and Siberia, the moose is dwarfed only by two species of bison.


Ecology and biology


Diet

The moose is a browsing herbivore and is capable of consuming many types of plant or fruit. The average adult moose needs to consume per day to maintain its body weight. Much of a moose's energy is derived from terrestrial vegetation, mainly consisting of forbs and other non-grasses, and fresh shoots from trees such as willow and
birch A birch is a thin-leaved deciduous hardwood tree of the genus ''Betula'' (), in the family Betulaceae, which also includes alders, hazels, and hornbeams. It is closely related to the beech- oak family Fagaceae. The genus ''Betula'' cont ...
. As these terrestrial plants are rather low in
sodium Sodium is a chemical element with the symbol Na (from Latin ''natrium'') and atomic number 11. It is a soft, silvery-white, highly reactive metal. Sodium is an alkali metal, being in group 1 of the periodic table. Its only stable ...
, as much as half of their diet usually consists of aquatic plants, including lilies and pondweed, which while lower in energy content, provides the moose with its sodium requirements.''Biology by numbers: an encouragement to quantitative thinking'' By Richard F. Burton – Cambridge University Press 1998 Page 84-85 In winter, moose are often drawn to roadways, to lick salt that is used as a snow and ice melter. A typical moose, weighing , can eat up to of food per day. Moose lack upper front
teeth A tooth ( : teeth) is a hard, calcified structure found in the jaws (or mouths) of many vertebrates and used to break down food. Some animals, particularly carnivores and omnivores, also use teeth to help with capturing or wounding prey, ...
, but have eight sharp incisors on the lower jaw. They also have a tough tongue, lips and gums, which aid in the eating of woody vegetation. Moose have six pairs of large, flat molars and, ahead of those, six pairs of premolars, to grind up their food. A moose's upper lip is very sensitive, to help distinguish between fresh shoots and harder twigs, and is
prehensile Prehensility is the quality of an appendage or organ that has adapted for grasping or holding. The word is derived from the Latin term ''prehendere'', meaning "to grasp". The ability to grasp is likely derived from a number of different orig ...
, for grasping their food. In the summer, moose may use this prehensile lip for grabbing branches and pulling, stripping the entire branch of leaves in a single mouthful, or for pulling forbs, like dandelions, or aquatic plants up by the base, roots and all. A moose's diet often depends on its location, but they seem to prefer the new growths from deciduous trees with a high sugar content, such as white birch, trembling aspen and
striped maple ''Acer pensylvanicum'', known as the striped maple, moosewood, moose maple or goosefoot maple, is a small North American species of maple. The striped maple is a sequential hermaphrodite, meaning that it can change its sex throughout its lifetim ...
, among many others. To reach high branches, a moose may bend small saplings down, using its prehensile lip, mouth or body. For larger trees a moose may stand erect and walk upright on its hind legs, allowing it to reach branches up to or higher above the ground. Moose are excellent swimmers and are known to wade into water to eat aquatic plants. This trait serves a second purpose in cooling down the moose on summer days and ridding itself of
black flies A black fly or blackfly (sometimes called a buffalo gnat, turkey gnat, or white socks) is any member of the family Simuliidae of the Culicomorpha infraorder. It is related to the Ceratopogonidae, Chironomidae, and Thaumaleidae. Over 2,200 spec ...
. Moose are thus attracted to marshes and river banks during warmer months as both provide suitable vegetation to eat and water to wet themselves in. Moose have been known to dive over to reach plants on lake bottoms, and the complex snout may assist the moose in this type of feeding. Moose are the only deer that are capable of feeding underwater.''Deer of the World: Their Evolution, Behaviour, and Ecology'' By Valerius Geist -- Stackpole Books 1998 Page 237 As an adaptation for feeding on plants underwater, the nose is equipped with fatty pads and muscles that close the nostrils when exposed to water pressure, preventing water from entering the nose. Other species can pluck plants from the water too, but these need to raise their heads in order to swallow. Moose are not
grazing In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and ot ...
animals but browsers (concentrate selectors). Like giraffes, moose carefully select foods with less fiber and more concentrations of nutrients. Thus, the moose's digestive system has evolved to accommodate this relatively low-fiber diet. Unlike most hooved, domesticated animals ( ruminants), moose cannot digest hay, and feeding it to a moose can be fatal. The moose's varied and complex diet is typically expensive for humans to provide, and free-range moose require a lot of forested hectarage for sustainable survival, which is one of the main reasons moose have never been widely domesticated. File:Alces alces bark stripping.jpg, left,
Bark Bark may refer to: * Bark (botany), an outer layer of a woody plant such as a tree or stick * Bark (sound), a vocalization of some animals (which is commonly the dog) Places * Bark, Germany * Bark, Warmian-Masurian Voivodeship, Poland Arts, e ...
stripping File:Bull moose close up feeding on fireweed.JPG, Bull moose eating a fireweed plant File:Moose 983 LAB.jpg, Bull moose browses a beaver pond


Natural predators

A full-grown moose has few enemies except Siberian tigers (''Panthera tigris tigris'') which regularly prey on adult moose, but a pack of
gray wolves The wolf (''Canis lupus''; : wolves), also known as the gray wolf or grey wolf, is a large canine native to Eurasia and North America. More than thirty subspecies of ''Canis lupus'' have been recognized, and gray wolves, as popularly u ...
(''Canis lupus'') can still pose a threat, especially to females with calves.
Brown bear The brown bear (''Ursus arctos'') is a large bear species found across Eurasia and North America. In North America, the populations of brown bears are called grizzly bears, while the subspecies that inhabits the Kodiak Islands of Alaska is k ...
s (''Ursus arctos'') are also known to prey on moose of various sizes and are the only predator besides the wolf to attack moose both in Eurasia and North America. However, brown bears are more likely to take over a wolf kill or to take young moose than to hunt adult moose on their own. Black bears (''Ursus americanus'') and cougars (''Puma concolor'') can be significant predators of moose calves in May and June and can, in rare instances, prey on adults (mainly cows rather than the larger bulls).
Wolverine The wolverine (), (''Gulo gulo''; ''Gulo'' is Latin for " glutton"), also referred to as the glutton, carcajou, or quickhatch (from East Cree, ''kwiihkwahaacheew''), is the largest land-dwelling species of the family Mustelidae. It is a musc ...
s (''Gulo gulo'') are most likely to eat moose as carrion but have killed moose, including adults, when the large ungulates are weakened by harsh winter conditions. Orcas (''Orcinus orca'') are the moose's only confirmed marine predator as they have been known to prey on moose swimming between islands out of North America's Northwest Coast, however, there is at least one recorded instance of a moose preyed upon by a
Greenland shark The Greenland shark (''Somniosus microcephalus''), also known as the gurry shark, grey shark, or by the Kalaallisut name ''eqalussuaq'', is a large shark of the family Somniosidae ("sleeper sharks"), closely related to the Pacific and souther ...
(''Somniosus microcephalus''). In some areas, moose are the primary source of food for wolves. Moose usually flee upon detecting wolves. Wolves usually follow moose at a distance of , occasionally at a distance of . Attacks from wolves against young moose may last seconds, though sometimes they can be drawn out for days with adults. Sometimes, wolves will chase moose into shallow streams or onto frozen rivers, where their mobility is greatly impeded. Moose will sometimes stand their ground and defend themselves by charging at the wolves or lashing out at them with their powerful hooves. Wolves typically kill moose by tearing at their haunches and perineum, causing massive
blood loss Bleeding, hemorrhage, haemorrhage or blood loss, is blood escaping from the circulatory system from damaged blood vessels. Bleeding can occur internally, or externally either through a natural opening such as the mouth, nose, ear, urethra, ...
. Occasionally, a wolf may immobilise a moose by biting its sensitive nose, the pain of which can paralyze a moose. Wolf packs primarily target calves and elderly animals, but can and will take healthy, adult moose. Moose between the ages of two and eight are seldom killed by wolves. Though moose are usually hunted by packs, there are cases in which single wolves have successfully killed healthy, fully-grown moose. Research into moose predation suggests that their response to perceived threats is learned rather than instinctual. In practical terms this means moose are more vulnerable in areas where wolf or bear populations were decimated in the past but are now rebounding. These same studies suggest, however, that moose learn quickly and adapt, fleeing an area if they hear or smell wolves, bears, or scavenger birds such as ravens. Moose are also subject to various diseases and forms of parasitism. In northern Europe, the moose botfly is a parasite whose range seems to be spreading.


Parasites

Moose typically carry a heavy burden of parasites, both externally and internally. Parasitosis is an important cause of moose morbidity and mortality and also contributes to vulnerability to predators.


Ectoparasites

Ectoparasites of moose include the moose nose bot fly, and winter ticks.


Endoparasites

Endoparasites Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson has c ...
of moose include dog tapeworm, meningeal worm, lungworm, and roundworm.


Social structure and reproduction

Moose are mostly diurnal. They are generally solitary with the strongest bonds between mother and calf. Although moose rarely gather in groups, there may be several in close proximity during the mating season. Rutting and mating occurs in September and October. During the rut, mature bulls will cease feeding completely for a period of approximately two weeks; this fasting behavior has been attributed to neurophysiological changes related to redeployment of olfaction for detection of moose urine and moose cows. The males are polygamous and will seek several females to breed with. During this time both sexes will call to each other. Males produce heavy grunting sounds that can be heard from up to 500 meters away, while females produce wail-like sounds. Males will fight for access to females. Initially, the males assess which of them is dominant and one bull may retreat, however, the interaction can escalate to a fight using their antlers. Female moose have an eight-month gestation period, usually bearing one calf, or twins if food is plentiful, in May or June. Twinning can run as high as 30% to 40% with good nutrition Newborn moose have fur with a reddish hue in contrast to the brown appearance of an adult. The young will stay with the mother until just before the next young are born. The life span of an average moose is about 15–25 years. Moose populations are stable at 25 calves for every 100 cows at 1 year of age. With availability of adequate nutrition, mild weather, and low predation, moose have a huge potential for population expansion. File:Moose calves nursing.jpg, (newborn)
Calves nursing in spring. File:Cowcalflyingdown.JPG, (3 months)
Calves stay near their mothers at all times. File:Ninemomoose.JPG, (9 months)
This calf is almost ready to leave its mother. File:Mainstmoose.JPG, (10–11 months)
This yearling was probably recently chased away by its pregnant mother.


Aggression

Moose are not usually aggressive towards humans, but can be provoked or frightened to behave with aggression. In terms of raw numbers, they attack more people than bears and wolves combined, but usually with only minor consequences. In the Americas, moose injure more people than any other wild mammal, and worldwide, only hippopotamuses injure more. When harassed or startled by people or in the presence of a dog, moose may charge. Also, as with bears or any wild animal, moose that have become used to being fed by people may act aggressively when denied food. During the fall mating season, bulls may be aggressive toward humans because of the high hormone levels they experience. Cows with young calves are very protective and will attack humans who come too close, especially if they come between mother and calf. Unlike other dangerous animals, moose are not territorial, and do not view humans as food, and will therefore usually not pursue humans if they simply run away. Like any wild animal, moose are unpredictable. They are most likely to attack if annoyed or harassed, or if approached too closely. A moose that has been harassed may vent its anger on anyone in the vicinity, and they often do not make distinctions between their tormentors and innocent passers-by. Moose are very limber animals with highly flexible joints and sharp, pointed hooves, and are capable of kicking with both front and back legs. Unlike other large, hooved mammals, such as horses, moose can kick in all directions including sideways. Therefore, there is no safe side from which to approach. However, moose often give warning signs prior to attacking, displaying their aggression by means of body language. Maintained eye contact is usually the first sign of aggression, while laid-back ears or a lowered head is a definite sign of agitation. If the hairs on the back of the moose's neck and shoulders ( hackles) stand up, a charge is usually imminent. The Anchorage Visitor Centers warn tourists that "...a moose with its hackles raised is a thing to fear." Studies suggest that the calls made by female moose during the rut not only call the males but can actually induce a bull to invade another bull's harem and fight for control of it. This in turn means that the cow moose has at least a small degree of control over which bulls she mates with. Moose often show aggression to other animals as well; especially predators. Bears are common predators of moose calves and, rarely, adults. Alaskan moose have been reported to successfully fend off attacks from both black and brown bears. Moose have been known to stomp attacking wolves, which makes them less preferred as prey to the wolves. Moose are fully capable of killing bears and wolves. In one rare event, a female moose killed two adult male wolves. A moose of either sex that is confronted by danger may let out a loud roar, more resembling that of a predator than a prey animal. European moose are often more aggressive than North American moose, such as the moose in Sweden, which often become very agitated at the sight of a predator. However, like all ungulates known to attack predators, the more aggressive individuals are always darker in color, with the darkest coloring usually in areas facing the opponent, thus serving as a natural warning to other animals.


Habitat, range, and distribution


Habitat

Moose require habitat with adequate edible plants (e.g., pond grasses, young trees and shrubs), cover from predators, and protection from extremely hot or cold weather. Moose travel among different habitats with the seasons to address these requirements. Moose are cold-adapted mammals with thickened skin, dense, heat-retaining coat, and a low surface:volume ratio, which provides excellent cold tolerance but poor heat tolerance. Moose survive hot weather by accessing shade or cooling wind, or by immersion in cool water. In hot weather, moose are often found wading or swimming in lakes or ponds. When heat-stressed, moose may fail to adequately forage in summer and may not gain adequate body fat to survive the winter. Also, moose cows may not calve without adequate summer weight gain. Moose require access to both young forest for browsing and mature forest for shelter and cover. Forest disturbed by fire and logging promotes the growth of fodder for moose. Moose also require access to mineral licks, safe places for calving and aquatic feeding sites. Moose avoid areas with little or no snow as this increases the risk of predation by wolves and avoid areas with deep snow, as this impairs mobility. Thus, moose select habitat on the basis of trade-offs between risk of predation, food availability, and snow depth. With reintroduction of bison into boreal forest, there was some concern that bison would compete with moose for winter habitat, and thereby worsen the population decline of moose. However, this does not appear to be a problem. Moose prefer sub-alpine shrublands in early winter, while bison prefer wet sedge valley meadowlands in early winter. In late winter, moose prefer river valleys with deciduous forest cover or alpine terrain above the tree line, while bison preferred wet sedge meadowlands or sunny southern grassy slopes.


North America

After expanding for most of the 20th century, the moose population of North America has been in steep decline since the 1990s. Populations expanded greatly with improved habitat and protection, but now the moose population is declining rapidly. This decline has been attributed to opening of roads and landscapes into the northern range of moose, allowing deer to become populous in areas where they were not previously common. This encroachment by deer on moose habitat brought moose into contact with previously unfamiliar pathogens, including brainworm and liver fluke, and these parasites are believed to have contributed to the population decline of moose. In
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, the moose range includes almost all of
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
(excluding the
arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
and Vancouver Island), most of
Alaska Alaska ( ; russian: Аляска, Alyaska; ale, Alax̂sxax̂; ; ems, Alas'kaaq; Yup'ik: ''Alaskaq''; tli, Anáaski) is a state located in the Western United States on the northwest extremity of North America. A semi-exclave of the U ...
, northern
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
and upstate New York, the upper
Rocky Mountains The Rocky Mountains, also known as the Rockies, are a major mountain range and the largest mountain system in North America. The Rocky Mountains stretch in straight-line distance from the northernmost part of western Canada, to New Mexico ...
, northern
Minnesota Minnesota () is a state in the upper midwestern region of the United States. It is the 12th largest U.S. state in area and the 22nd most populous, with over 5.75 million residents. Minnesota is home to western prairies, now given over t ...
, northern
Wisconsin Wisconsin () is a state in the upper Midwestern United States. Wisconsin is the 25th-largest state by total area and the 20th-most populous. It is bordered by Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake M ...
,
Michigan Michigan () is a state in the Great Lakes region of the upper Midwestern United States. With a population of nearly 10.12 million and an area of nearly , Michigan is the 10th-largest state by population, the 11th-largest by area, and t ...
's Upper Peninsula, and Isle Royale in Lake Superior. This massive range, containing diverse habitats, contains four of the six North American subspecies. In the West, moose populations extend across Canada (
British Columbia British Columbia (commonly abbreviated as BC) is the westernmost province of Canada, situated between the Pacific Ocean and the Rocky Mountains. It has a diverse geography, with rugged landscapes that include rocky coastlines, sandy beaches, for ...
and
Alberta Alberta ( ) is one of the thirteen provinces and territories of Canada. It is part of Western Canada and is one of the three prairie provinces. Alberta is bordered by British Columbia to the west, Saskatchewan to the east, the Northwest T ...
). Isolated groups have been verified as far south as the mountains of Utah and Colorado and as far west as the Lake Wenatchee area of the Washington Cascades. In the northwestern US, the range includes
Wyoming Wyoming () is a state in the Mountain West subregion of the Western United States. It is bordered by Montana to the north and northwest, South Dakota and Nebraska to the east, Idaho to the west, Utah to the southwest, and Colorado to t ...
, Montana,
Idaho Idaho ( ) is a U.S. state, state in the Pacific Northwest region of the Western United States. To the north, it shares a small portion of the Canada–United States border with the province of British Columbia. It borders the states of Monta ...
, and smaller areas of
Washington Washington commonly refers to: * Washington (state), United States * Washington, D.C., the capital of the United States ** A metonym for the federal government of the United States ** Washington metropolitan area, the metropolitan area centered o ...
, and Oregon. Moose have extended their range southwards in the western Rocky Mountains, with initial sightings in Yellowstone National Park in 1868, and then to the northern slope of the
Uinta Mountains The Uinta Mountains ( ) are an east-west trending chain of mountains in northeastern Utah extending slightly into southern Wyoming in the United States. As a subrange of the Rocky Mountains, they are unusual for being the highest range in the ...
in Utah in the first half of the twentieth century. This is the southernmost naturally established moose population in the United States. In 1978, a few breeding pairs were reintroduced in western Colorado, and the state's moose population is now more than 2,400. In northeastern North America, the Eastern moose's history is very well documented: moose meat was often a staple in the diet of indigenous peoples for centuries. The common name "moose" was brought into English from the word used by those who lived in present day coastal
Rhode Island Rhode Island (, like ''road'') is a U.S. state, state in the New England region of the Northeastern United States. It is the List of U.S. states by area, smallest U.S. state by area and the List of states and territories of the United States ...
. The indigenous people often used moose hides for leather and its meat as an ingredient in pemmican, a type of dried jerky used as a source of sustenance in winter or on long journeys. The historical range of the subspecies extended from well into Quebec, the Maritimes, and Eastern Ontario south to include all of New England finally ending in the very northeastern tip of
Pennsylvania Pennsylvania (; ( Pennsylvania Dutch: )), officially the Commonwealth of Pennsylvania, is a state spanning the Mid-Atlantic, Northeastern, Appalachian, and Great Lakes regions of the United States. It borders Delaware to its southeast, ...
in the west, cutting off somewhere near the mouth of the
Hudson River The Hudson River is a river that flows from north to south primarily through eastern New York. It originates in the Adirondack Mountains of Upstate New York and flows southward through the Hudson Valley to the New York Harbor between Ne ...
in the south. The moose has been extinct in much of the eastern U.S. for as long as 150 years, due to colonial era overhunting and destruction of its habitat: Dutch, French, and British colonial sources all attest to its presence in the mid 17th century from Maine south to areas within of present-day Manhattan. However, by the 1870s, only a handful of moose existed in this entire region in very remote pockets of forest; less than 20% of suitable habitat remained. Since the 1980s, however, moose populations have rebounded, thanks to regrowth of plentiful food sources, abandonment of farmland, better land management, clean-up of pollution, and natural dispersal from the Canadian Maritimes and
Quebec Quebec ( ; )According to the Canadian government, ''Québec'' (with the acute accent) is the official name in Canadian French and ''Quebec'' (without the accent) is the province's official name in Canadian English is one of the thirte ...
. South of the Canada–US border,
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...
has most of the population with a 2012 headcount of about 76,000 moose. Dispersals from Maine over the years have resulted in healthy, growing populations each in
Vermont Vermont () is a U.S. state, state in the northeast New England region of the United States. Vermont is bordered by the states of Massachusetts to the south, New Hampshire to the east, and New York (state), New York to the west, and the Provin ...
and
New Hampshire New Hampshire is a U.S. state, state in the New England region of the northeastern United States. It is bordered by Massachusetts to the south, Vermont to the west, Maine and the Gulf of Maine to the east, and the Canadian province of Quebec t ...
, notably near bodies of water and as high up as above sea level in the mountains. In
Massachusetts Massachusetts (Massachusett: ''Muhsachuweesut Massachusett_writing_systems.html" ;"title="nowiki/> məhswatʃəwiːsət.html" ;"title="Massachusett writing systems">məhswatʃəwiːsət">Massachusett writing systems">məhswatʃəwiːsət'' En ...
, moose had gone extinct by 1870, but re-colonized the state in the 1960s, with the population expanding from Vermont and New Hampshire; by 2010, the population was estimated at 850–950. Moose reestablished populations in eastern New York and
Connecticut Connecticut () is the southernmost state in the New England region of the Northeastern United States. It is bordered by Rhode Island to the east, Massachusetts to the north, New York (state), New York to the west, and Long Island Sound to the ...
and appeared headed south towards the Catskill Mountains, a former habitat. In the Midwest U.S., moose are primarily limited to the upper Great Lakes region, but strays, primarily immature males, have been found as far south as eastern Iowa. For unknown reasons, the moose population is declining rapidly in the Midwest. Moose were successfully introduced on Newfoundland in 1878 and 1904, where they are now the dominant ungulate, and somewhat less successfully on Anticosti Island in the Gulf of Saint Lawrence. File:Alaskan moose pair (6862339335).jpg, Cow and bull moose File:Cow moose.jpg, Cow moose File:Alces alces (juvenile).jpg, Moose calf


Decline in population

Since the 1990s, moose populations have declined dramatically in much of temperate North America, although they remain stable in
Arctic The Arctic ( or ) is a polar regions of Earth, polar region located at the northernmost part of Earth. The Arctic consists of the Arctic Ocean, adjacent seas, and parts of Canada (Yukon, Northwest Territories, Nunavut), Danish Realm (Greenla ...
and subarctic regions. The exact causes of specific die-offs are not determined, but most documented mortality events were due to wolf predation, bacterial infection due to injuries sustained from predators, and
parasite Parasitism is a close relationship between species, where one organism, the parasite, lives on or inside another organism, the host, causing it some harm, and is adapted structurally to this way of life. The entomologist E. O. Wilson h ...
s from white-tailed deer to which moose have not developed a natural defense, such as liver flukes, brain worms and winter tick infestations. Predation of moose calves by brown bear is also significant. Landscape change from salvage logging of forest damage caused by the mountain pine beetle has resulted in greater foraging in logged areas by female moose, and this is the lead hypothesis as to why the moose population is declining in eastern North American forests, as this likely leads to increased predation. An alternate hypotheses among biologists for generalized, nonhunting declines in moose populations at the southern extent of their range is increasing heat stress brought on by the rapid seasonal temperature upswings as a result of human-induced climate change. Biologists studying moose populations typically use warm-season, heat-stress thresholds of between . However, the minor average temperature increase of 0.83–1.11 °C (1.5–2 °F), over the last 100 years, has resulted in milder winters that induce favorable conditions for ticks, parasites and other invasive species to flourish within the southern range of moose habitat in North America. The moose population in New Hampshire fell from 7,500 in the early 2000s to a 2014 estimate of 4,000 and in Vermont the numbers were down to 2,200 from a high of 5,000 animals in 2005. Much of the decline has been attributed to the winter tick, which, between 2017 and 2019, accounted for 74% of all winter mortality and 91% of winter calf deaths in Vermont. Moose with heavy tick infections will rub their fur down to the skin raw trying to get the ticks off, making them look white when their outer coat rubs off. Locals call them ghost moose. Loss of the insulating winter coat through attempts to rid the moose of winter tick increases the risk of hypothermia in winter.


Europe and Asia

In
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, moose are currently found in large numbers throughout
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of ...
,
Sweden Sweden, formally the Kingdom of Sweden,The United Nations Group of Experts on Geographical Names states that the country's formal name is the Kingdom of SwedenUNGEGN World Geographical Names, Sweden./ref> is a Nordic countries, Nordic c ...
,
Finland Finland ( fi, Suomi ; sv, Finland ), officially the Republic of Finland (; ), is a Nordic country in Northern Europe. It shares land borders with Sweden to the northwest, Norway to the north, and Russia to the east, with the Gulf of Bot ...
, Latvia,
Estonia Estonia, formally the Republic of Estonia, is a country by the Baltic Sea in Northern Europe. It is bordered to the north by the Gulf of Finland across from Finland, to the west by the sea across from Sweden, to the south by Latvia, an ...
,
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
, with more modest numbers in the southern
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
,
Belarus Belarus,, , ; alternatively and formerly known as Byelorussia (from Russian ). officially the Republic of Belarus,; rus, Республика Беларусь, Respublika Belarus. is a landlocked country in Eastern Europe. It is bordered by ...
, and northern
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inva ...
. They are also widespread through
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
on up through the borders with Finland south towards the border with Estonia, Belarus and Ukraine and stretching far away eastwards to the Yenisei River in Siberia. The European moose was native to most temperate areas with suitable habitat on the continent and even
Scotland Scotland (, ) is a country that is part of the United Kingdom. Covering the northern third of the island of Great Britain, mainland Scotland has a border with England to the southeast and is otherwise surrounded by the Atlantic Ocean to ...
from the end of the last Ice Age, as Europe had a mix of temperate boreal and deciduous forest. Up through Classical times, the species was certainly thriving in both Gaul and Magna Germania, as it appears in military and hunting accounts of the age. However, as the Roman era faded into medieval times, the beast slowly disappeared: soon after the reign of
Charlemagne Charlemagne ( , ) or Charles the Great ( la, Carolus Magnus; german: Karl der Große; 2 April 747 – 28 January 814), a member of the Carolingian dynasty, was King of the Franks from 768, King of the Lombards from 774, and the first E ...
, the moose disappeared from
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, where its range extended from Normandy in the north to the Pyrenees in the south. Farther east, it survived in
Alsace Alsace (, ; ; Low Alemannic German/ gsw-FR, Elsàss ; german: Elsass ; la, Alsatia) is a cultural region and a territorial collectivity in eastern France, on the west bank of the upper Rhine next to Germany and Switzerland. In 2020, it had ...
and the
Netherlands ) , anthem = ( en, "William of Nassau") , image_map = , map_caption = , subdivision_type = Sovereign state , subdivision_name = Kingdom of the Netherlands , established_title = Before independence , established_date = Spanish Netherl ...
until the 9th century as the marshlands in the latter were drained and the forests were cleared away for feudal lands in the former. It was gone from
Switzerland ). Swiss law does not designate a ''capital'' as such, but the federal parliament and government are installed in Bern, while other federal institutions, such as the federal courts, are in other cities (Bellinzona, Lausanne, Luzern, Neuchâtel ...
by the year 1000, from the western Czech Republic by 1300, from Mecklenburg in
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwee ...
by c. 1600, and from Hungary and the Caucasus since the 18th and 19th century, respectively. By the early 20th century, the last strongholds of the European moose appeared to be in Fennoscandian areas and patchy tracts of Russia, with a few migrants found in what is now Estonia and Lithuania. The USSR and Poland managed to restore portions of the range within its borders (such as the 1951 reintroduction into
Kampinos National Park Kampinos National Park ( pl, Kampinoski Park Narodowy) is a National Park in east-central Poland, in Masovian Voivodeship, on the north-west outskirts of Warsaw. It has a sister park agreement with Indiana Dunes National Park, Indiana, United Stat ...
and the later 1958 reintroduction in Belarus), but political complications limited the ability to reintroduce it to other portions of its range. Attempts in 1930 and again in 1967 in marshland north of Berlin were unsuccessful. At present in Poland, populations are recorded in the
Biebrza Biebrza ( lt, Bebras, '' be, Bobra'', ''german: Bober'') is a river in northeastern Poland, a tributary of the Narew river (near Wizna), with a length of and a basin area of 7,092 km2 (7,067 in Poland).Białowieża Forest Białowieża Forest; lt, Baltvyžių giria; pl, Puszcza Białowieska  ; russian: Беловежская пуща, Belovezhskaya Pushcha is a forest on the border between Belarus and Poland. It is one of the last and largest remaining pa ...
. It has migrated into other parts of Eastern Europe and has been spotted in eastern and southern Germany. Unsuccessful thus far in recolonizing these areas via natural dispersal from source populations in
Poland Poland, officially the Republic of Poland, is a country in Central Europe. It is divided into 16 administrative provinces called voivodeships, covering an area of . Poland has a population of over 38 million and is the fifth-most populou ...
,
Belarus Belarus,, , ; alternatively and formerly known as Byelorussia (from Russian ). officially the Republic of Belarus,; rus, Республика Беларусь, Respublika Belarus. is a landlocked country in Eastern Europe. It is bordered by ...
,
Ukraine Ukraine ( uk, Україна, Ukraïna, ) is a country in Eastern Europe. It is the second-largest European country after Russia, which it borders to the east and northeast. Ukraine covers approximately . Prior to the ongoing Russian inva ...
,
Czech Republic The Czech Republic, or simply Czechia, is a landlocked country in Central Europe. Historically known as Bohemia, it is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the southeast. The ...
, and
Slovakia Slovakia (; sk, Slovensko ), officially the Slovak Republic ( sk, Slovenská republika, links=no ), is a landlocked country in Central Europe. It is bordered by Poland to the north, Ukraine to the east, Hungary to the south, Austria to the ...
, it appears to be having more success migrating south into the
Caucasus The Caucasus () or Caucasia (), is a region between the Black Sea and the Caspian Sea, mainly comprising Armenia, Azerbaijan, Georgia, and parts of Southern Russia. The Caucasus Mountains, including the Greater Caucasus range, have historica ...
. It is listed under Appendix III of the Bern Convention.Alces alces (Eurasian Elk, Moose, Elk, Eurasian Moose, European Elk, Siberian Elk)
. Iucnredlist.org. Retrieved on January 9, 2011.
In 2008, two moose were reintroduced into the Scottish Highlands in
Alladale Wilderness Reserve Alladale Wilderness Reserve is a highland estate in the Scottish Highlands. The estate was purchased in 2003 by conservationist and philanthropist Paul Lister, who hopes to recreate a wooded landscape and reintroduce native animals including Pr ...
. The East Asian moose populations confine themselves mostly to the territory of
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
, with much smaller populations in
Mongolia Mongolia; Mongolian script: , , ; lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia to the north and China to the south. It covers an area of , with a population of just 3.3 million ...
and Northeastern
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
. Moose populations are relatively stable in Siberia and increasing on the Kamchatka Peninsula. In Mongolia and China, where poaching took a great toll on moose, forcing them to near extinction, they are protected, but enforcement of the policy is weak and demand for traditional medicines derived from deer parts is high. In 1978, the Regional Hunting Department transported 45 young moose to the center of Kamchatka. These moose were brought from Chukotka, home to the largest moose on the planet. Kamchatka now regularly is responsible for the largest trophy moose shot around the world each season. As it is a fertile environment for moose, with a milder climate, less snow, and an abundance of food, moose quickly bred and settled along the valley of the Kamchatka River and many surrounding regions. The population in the past 20 years has risen to over 2,900 animals. The size of the moose varies. Following Bergmann's rule, population in the south (''A. a. cameloides'') usually grow smaller, while moose in the north and northeast (''A. a. buturlini'') can match the imposing sizes of the Alaskan moose (''A. a. gigas'') and are prized by trophy hunters.


New Zealand

In 1900, an attempt to introduce moose into the Hokitika area failed; then in 1910 ten moose (four bulls and six cows) were introduced into Fiordland. This area is considered a less than suitable habitat, and subsequent low numbers of sightings and kills have led to some presumption of this population's failure. The last proven sighting of a moose in New Zealand was in 1952. However, a moose antler was found in 1972, and DNA tests showed that hair collected in 2002 was from a moose. There has been extensive searching, and while automated cameras failed to capture photographs, evidence was seen of bedding spots, browsing, and antler marks.


Paleontology

Moose are an old genus. Like its relatives, '' Odocoileus'' and ''
Capreolus ''Capreolus'' is a genus of deer, the roe deer. Etymology English ''roe'' is from Old English ''ra'' or ''rá'', from ''raha'', from Proto-Germanic ''*raikhaz'', cognate to Old Norse ''ra'', Old Saxon ''reho'', Middle Dutch and Dutch '' ...
'', the genus ''Alces'' gave rise to very few species that endured for long periods of time. This differs from the megacerines, such as the Irish elk, which evolved many species before going extinct. Some scientists, such as Adrian Lister, group the moose and all its extinct relatives into one genus, ''Alces'', while others, such as Augusto Azzaroli, restrict ''Alces'' to the living species, placing the fossil species into the genera '' Cervalces'' (stag moose) and ''Libralces'' (source moose). The earliest known species in the moose lineage is '' Libralces gallicus'' (French moose), which lived in the
Pliocene epoch The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58wapiti The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The commo ...
. However, the rest of its skull structure, skeletal structure and teeth bore strong resemblance to those features that are unmistakable in modern moose, indicating a similar diet. Its antlers consisted of a horizontal bar long, with no tines, ending in small palmations. Its skull and neck structure suggest an animal that fought using high-speed impacts, much like the Dall sheep, rather than locking and twisting antlers the way modern moose combat. Their long legs and bone structure suggest an animal that was adapted to running at high speeds over rough terrain.''Deer of the world: their evolution, behaviour, and ecology'' By Valerius Geist - Page 244-250''Morphological Change in Quaternary Mammals of North America'' by Robert Allen Martin, Anthony D. Barnosky - Cambridge University Press 1993 Page 178-181 ''Libralces'' existed until the middle
Pleistocene epoch The Pleistocene ( , often referred to as the '' Ice age'') is the geological epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was finally confirmed i ...
and were followed briefly by a species called ''Cervalces carnutorum'' (moose of the Carnutes). The main differences between the two consisted of shortening of the horizontal bar in the antlers and broadening of the palmations, indicating a likely change from open plains to more forested environments, and skeletal changes in the joints and toes that suggest an adaptation to marshy,
taiga Taiga (; rus, тайга́, p=tɐjˈɡa; relates to Mongolic and Turkic languages), generally referred to in North America as a boreal forest or snow forest, is a biome characterized by coniferous forests consisting mostly of pines, sp ...
environments. ''Cervalces carnutorum'' was soon followed by a much larger species called '' Cervalces latifrons'' (broad-fronted stag-moose). The Pleistocene epoch was a time of gigantism, in which most species were much larger than their descendants of today, including exceptionally large lions, hippopotamuses, mammoths, and deer. Many fossils of ''Cervalces latifrons'' have been found in Siberia, dating from about 1.2 to 0.5 million years ago. This is most likely the time at which the species migrated from the Eurasian continent to North America. Like its descendants, it inhabited mostly northern latitudes, and was probably well-adapted to the cold. ''C. latifrons'' was the largest deer known to have ever existed, standing more than tall at the shoulders. This is bigger than even the Irish elk (megacerine), which was tall at the shoulders. Its antlers were smaller than the Irish elk's, but comparable in size to those of ''L. gallicus''. However, the antlers had a shorter horizontal bar and larger palmations, more resembling those of a modern moose. ''Alces alces'' (the modern moose) appeared during the late Pleistocene epoch. The species arrived in North America at the end of the Pleistocene and coexisted with a late-surviving variety or relative of ''C. latifrons'', which Azzaroli classified as a separate species called '' Cervalces scotti'', or the American stag-moose.


Populations

North America: * ''In Canada'': There are an estimated 500,000 to 1,000,000 moose, with 150,000 in Newfoundland in 2007 descended from just four that were introduced in the 1900s. * ''In United States'': There are estimated to be around 300,000: ** Alaska: The state's Department of Fish and Game estimated 200,000 in 2011. ** Northeast: A wildlife ecologist estimated 50,000 in New York and New England in 2007, with expansion expected. ** Rocky Mountain states: Wyoming is said to have the largest share in its six-state region, and its Fish and Game Commission estimated 7,692 in 2009. ** Upper Midwest: Michigan 2000 on Isle Royale (2019) and an estimated 433 (in its Upper Peninsula) in 2011, Wisconsin, 20–40 (close to its border with Michigan) in 2003, Minnesota 5600 in its northeast in 2010, and under 100 in its northwest in 2009; North Dakota closed, due to low moose population, one of its moose-hunting geographic units in 2011, and issued 162 single-kill licenses to hunters, each restricted to one of the remaining nine units. Europe and Asia: * ''Finland'': In 2009, there was a summer population of 115,000. * ''Norway'': In 2009, there were a winter population of around 120,000. In 2015 31,131 moose were shot. In 1999, a record number of 39,422 moose were shot. * ''Latvia'': in 2015, there were 21,000. * ''Estonia'': 13,260 * ''Lithuania'': around 14,000 in 2016 * ''Poland'': 28,000 * ''Czech Republic'': maximum of 50 * ''Russia'': In 2007, there were approximately 600,000. * ''Sweden'': Summer population is estimated to be 300,000–400,000. Around 100,000 are shot each fall. About 10,000 are killed in traffic accidents yearly.


Subspecies


Relationship with humans


History

European
rock drawings A petroglyph is an image created by removing part of a rock surface by incising, picking, carving, or abrading, as a form of rock art. Outside North America, scholars often use terms such as "carving", "engraving", or other descriptions ...
and cave paintings reveal that moose have been hunted since the Stone Age. Excavations in Alby, Sweden, adjacent to the Stora Alvaret have yielded moose antlers in wooden hut remains from 6000 BCE, indicating some of the earliest moose hunting in northern Europe. In northern Scandinavia one can still find remains of trapping pits used for hunting moose. These pits, which can be up to in area and deep, would have been camouflaged with branches and leaves. They would have had steep sides lined with planks, making it impossible for the moose to escape once it fell in. The pits are normally found in large groups, crossing the moose's regular paths and stretching over several km. Remains of wooden fences designed to guide the animals toward the pits have been found in bogs and peat. In Norway, an early example of these trapping devices has been dated to around 3700 BC. Trapping elk in pits is an extremely effective hunting method. As early as the 16th century the Norwegian government tried to restrict their use; nevertheless, the method was in use until the 19th century. The earliest recorded description of the moose is in
Julius Caesar Gaius Julius Caesar (; ; 12 July 100 BC – 15 March 44 BC), was a Roman general and statesman. A member of the First Triumvirate, Caesar led the Roman armies in the Gallic Wars before defeating his political rival Pompey in a civil war, an ...
's '' Commentarii de Bello Gallico'', where it is described thus:
There are also nimals which are called ''
alces The moose (in North America) or elk (in Eurasia) (''Alces alces'') is a member of the New World deer subfamily and is the only species in the genus ''Alces''. It is the largest and heaviest extant species in the deer family. Most adult mal ...
'' (moose). The shape of these, and the varied color of their skins, is much like roes, but in size they surpass them a little and are destitute of horns, and have legs without joints and ligatures; nor do they lie down for the purpose of rest, nor, if they have been thrown down by any accident, can they raise or lift themselves up. Trees serve as beds to them; they lean themselves against them, and thus reclining only slightly, they take their rest; when the huntsmen have discovered from the footsteps of these animals whither they are accustomed to betake themselves, they either undermine all the trees at the roots, or cut into them so far that the upper part of the trees may appear to be left standing. When they have leant upon them, according to their habit, they knock down by their weight the unsupported trees, and fall down themselves along with them.
In book 8, chapter 16 of Pliny the Elder's '' Natural History'' from 77 CE, the elk and an animal called achlis, which is presumably the same animal, are described thus:
... there is, also, the moose, which strongly resembles our steers, except that it is distinguished by the length of the ears and of the neck. There is also the achlis, which is produced in the land of Scandinavia; it has never been seen in this city, although we have had descriptions of it from many persons; it is not unlike the moose, but has no joints in the hind leg. Hence, it never lies down, but reclines against a tree while it sleeps; it can only be taken by previously cutting into the tree, and thus laying a trap for it, as otherwise, it would escape through its swiftness. Its upper lip is so extremely large, for which reason it is obliged to go backwards when grazing; otherwise, by moving onwards, the lip would get doubled up.


As food

Moose are hunted as a game species in many of the countries where they are found. Moose meat tastes, wrote Henry David Thoreau in "The Maine Woods", "like tender beef, with perhaps more flavour; sometimes like veal". While the flesh has
protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residues. Proteins perform a vast array of functions within organisms, including catalysing metabolic reactions, DNA replication, res ...
levels similar to those of other comparable
red meat In gastronomy, red meat is commonly red when raw and a dark color after it is cooked, in contrast to white meat, which is pale in color before and after cooking. In culinary terms, only flesh from mammals or fowl (not fish) is classified a ...
s (e.g.
beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantit ...
, deer and
wapiti The elk (''Cervus canadensis''), also known as the wapiti, is one of the largest species within the deer family, Cervidae, and one of the largest terrestrial mammals in its native range of North America and Central and East Asia. The commo ...
), it has a low fat content, and the fat that is present consists of a higher proportion of polyunsaturated fats than saturated fats. Dr.
Valerius Geist Valerius Geist (2 February 1938 – 6 July 2021) was a Canadian biologist and a professor emeritus in the Faculty of Environmental Design at the University of Calgary. He was a specialist on the biology, behavior, and social dynamics of North Am ...
, who emigrated to Canada from the Soviet Union, wrote in his 1999 book ''Moose: Behaviour, Ecology, Conservation'': Boosting moose populations in Alaska for hunting purposes is one of the reasons given for allowing aerial or airborne methods to remove wolves in designated areas, e.g., Craig Medred: "A kill of 124 wolves would thus translate to he survival of1488 moose or 2976 caribou or some combination thereof". Some scientists believe that this artificial inflation of game populations is actually detrimental to both caribou and moose populations as well as the ecosystem as a whole. This is because studies have shown that when these game populations are artificially boosted, it leads to both habitat destruction and a crash in these populations.


Consumption of offal

Cadmium Cadmium is a chemical element with the Symbol (chemistry), symbol Cd and atomic number 48. This soft, silvery-white metal is chemically similar to the two other stable metals in group 12 element, group 12, zinc and mercury (element), mercury. Li ...
levels are high in Finnish moose
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
and kidneys, with the result that consumption of these organs from moose more than one year old is prohibited in Finland. As a result of a study reported in 1988, the
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
Ministry of Natural Resources An environmental ministry is a national or subnational government agency politically responsible for the environment and/or natural resources. Various other names are commonly used to identify such agencies, such as Ministry of the Environment, ...
recommended against the consumption of moose and deer kidneys and livers. Levels of cadmium were found to be considerably higher than in Scandinavia. The
New Brunswick New Brunswick (french: Nouveau-Brunswick, , locally ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. It is the only province with both English and ...
Department of Natural Resources advises hunters not to consume cervid offal. Cadmium intake has been found to be elevated amongst all consumers of moose meat, though the meat was found to contribute only slightly to the daily cadmium intake. However the consumption of moose liver or kidneys significantly increased cadmium intake, with the study revealing that heavy consumers of moose organs have a relatively narrow safety margin below the levels which would probably cause adverse
health effects Health effects (or health impacts) are changes in health resulting from exposure to a source. Health effects are an important consideration in many areas, such as hygiene, pollution studies, occupational safety and health, ( utrition and health sc ...
.


Vehicle collisions

The center of mass of a moose is above the hood of most
passenger cars A car or automobile is a motor vehicle with wheels. Most definitions of ''cars'' say that they run primarily on roads, seat one to eight people, have four wheels, and mainly transport people instead of goods. The year 1886 is regarded as t ...
. In a collision, the impact crushes the front roof beams and individuals in the front seats. Collisions of this type are frequently fatal; seat belts and airbags offer little protection. In collisions with higher vehicles (such as trucks), most of the deformation is to the front of the vehicle and the passenger compartment is largely spared. Moose collisions have prompted the development of a vehicle test referred to as the " moose test" ( sv, Älgtest, german: Elchtest). A Massachusetts study found that moose–vehicular collisions had a very high human fatality rate and that such collisions caused the death of 3% of the Massachusetts moose population annually. Moose warning signs are used on roads in regions where there is a danger of collision with the animal. The triangular warning signs common in Sweden, Norway, and Finland have become coveted souvenirs among tourists traveling in these countries, causing road authorities so much expense that the moose signs have been replaced with imageless generic warning signs in some regions. In Ontario, Canada, an estimated 265 moose die each year as a result of collision with trains. Moose–train collisions were more frequent in winters with above-average snowfall. In January 2008, the Norwegian newspaper '' Aftenposten'' estimated that some 13,000 moose had died in collisions with Norwegian trains since 2000. The state agency in charge of railroad infrastructure (Jernbaneverket) plans to spend 80 million Norwegian kroner to reduce collision rate in the future by fencing the railways, clearing vegetation from near the tracks, and providing alternative snow-free feeding places for the animals elsewhere. In the Canadian province of
New Brunswick New Brunswick (french: Nouveau-Brunswick, , locally ) is one of the thirteen provinces and territories of Canada. It is one of the three Maritime provinces and one of the four Atlantic provinces. It is the only province with both English and ...
, collisions between automobiles and moose are frequent enough that all new highways have fences to prevent moose from accessing the road, as has long been done in Finland, Norway, and Sweden. A demonstration project, Highway 7 between Fredericton and Saint John, which has one of the highest frequencies of moose collisions in the province, did not have these fences until 2008, although it was and continues to be extremely well signed. Newfoundland and Labrador recommended that motorists use caution between dusk and dawn because that is when moose are most active and most difficult to see, increasing the risk of collisions. Local moose sightings are often reported on radio stations so that motorists can take care while driving in particular areas. An electronic "moose detection system" was installed on two sections of the Trans-Canada Highway in Newfoundland in 2011, but the system proved unreliable and was removed in 2015. In Sweden, a road will not be fenced unless it experiences at least one moose accident per km per year. In eastern Germany, where the scarce population is slowly increasing, there were two road accidents involving moose since 2000. File:NO road sign 146.1.svg, Norwegian road sign. File:Moosecrossingkenaiak.JPG, Warning sign in Alaska where trees and brush are trimmed along high moose crossing areas so that moose can be seen as they approach the road. File:Moose crossing a road.jpg, Moose (''A. a. gigas'') crossing a road in Alaska. File:Moose crossing warning sign.jpg, Canadian road sign.


Domestication

Domestication of moose was investigated in the
Soviet Union The Soviet Union,. officially the Union of Soviet Socialist Republics. (USSR),. was a transcontinental country that spanned much of Eurasia from 1922 to 1991. A flagship communist state, it was nominally a federal union of fifteen nationa ...
before
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the World War II by country, vast majority of the world's countries—including all of the great power ...
. Early experiments were inconclusive, but with the creation of a moose farm at
Pechora-Ilych Nature Reserve Pechora-Ilych Nature Reserve (russian: Печоро-Илычский заповедник, ''Pechoro-Ilychsky zapovednik'') is a Russian 'zapovednik' (strict nature reserve) in the Komi Republic, Russia. It currently occupies 7,213 square kilometer ...
in 1949, a small-scale moose domestication program was started, involving attempts at
selective breeding Selective breeding (also called artificial selection) is the process by which humans use animal breeding and plant breeding to selectively develop particular phenotypic traits (characteristics) by choosing which typically animal or plant ...
of animals on the basis of their behavioural characteristics. Since 1963, the program has continued at Kostroma Moose Farm, which had a herd of 33
tame Tame may refer to: *Taming, the act of training wild animals *River Tame, Greater Manchester *River Tame, West Midlands and the Tame Valley *Tame, Arauca, a Colombian town and municipality * "Tame" (song), a song by the Pixies from their 1989 alb ...
moose as of 2003. Although at this stage the farm is not expected to be a profit-making enterprise, it obtains some income from the sale of
moose milk Moose milk, also known as elk milk, refers to milk produced by moose (''Alces alces''). Though it is most commonly consumed by moose calves, its production has also been commercialised in Russia, Sweden and Canada. Nutritional content Moose ...
and from visiting tourist groups. Its main value, however, is seen in the opportunities it offers for the research in the
physiology Physiology (; ) is the scientific study of functions and mechanisms in a living system. As a sub-discipline of biology, physiology focuses on how organisms, organ systems, individual organs, cells, and biomolecules carry out the chemic ...
and behavior of the moose, as well as in the insights it provides into the general principles of animal domestication. In Sweden, there was a debate in the late 18th century about the national value of using the moose as a domestic animal. Among other things, the moose was proposed to be used in postal distribution, and there was a suggestion to develop a moose-mounted cavalry. Such proposals remained unimplemented, mainly because the extensive hunting for moose that was deregulated in the 1790s nearly drove it to extinction. While there have been documented cases of individual moose being used for riding and/or pulling carts and sleds, Björklöf concludes no wide-scale usage has occurred outside fairy tales.


Heraldry

As one of the Canadian national symbols, the moose occurs on several Canadian coats of arms, including Newfoundland and Labrador, and
Ontario Ontario ( ; ) is one of the thirteen provinces and territories of Canada.Ontario is located in the geographic eastern half of Canada, but it has historically and politically been considered to be part of Central Canada. Located in Central Ca ...
. Moose is also a common coat of arms in Europe as well, for example in Finland it appears on the coats of arms of Hirvensalmi and Mäntsälä municipalities. The Seal of Michigan features a moose.


See also

* Älgen Stolta, a rare example of a domesticated moose


References


Further reading

*
Alces: A journal devoted to the biology and management of moose (Alces alces)
' Centre for Northern Forest Ecosystem Research. * * * *


External links

*
A moose in the National Nature Park "Losinyj Ostrov" ("Moose" or "Elk" Island) in Russia
* from the
National Museum of Natural History The National Museum of Natural History is a natural history museum administered by the Smithsonian Institution, located on the National Mall in Washington, D.C., United States. It has free admission and is open 364 days a year. In 2021, with 7. ...
{{Authority control Holarctic fauna Mammals of Asia Mammals of Europe Mammals of North America Mammals described in 1758 Taxa named by Carl Linnaeus Symbols of Alaska Capreolinae