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biogeography Biogeography is the study of the distribution of species and ecosystems in geographic space and through geological time. Organisms and biological communities often vary in a regular fashion along geographic gradients of latitude, elevation, ...
, the Mediterranean Basin (; also known as the Mediterranean Region or sometimes Mediterranea) is the region of lands around the
Mediterranean Sea The Mediterranean Sea is a sea connected to the Atlantic Ocean, surrounded by the Mediterranean Basin and almost completely enclosed by land: on the north by Western and Southern Europe and Anatolia, on the south by North Africa, and on ...
that have mostly a
Mediterranean climate A Mediterranean climate (also called a dry summer temperate climate ''Cs'') is a temperate climate sub-type, generally characterized by warm, dry summers and mild, fairly wet winters; these weather conditions are typically experienced in the ...
, with mild to cool, rainy
winter Winter is the coldest season of the year in polar and temperate climates. It occurs after autumn and before spring. The tilt of Earth's axis causes seasons; winter occurs when a hemisphere is oriented away from the Sun. Different cultur ...
s and warm to hot, dry
summer Summer is the hottest of the four temperate seasons, occurring after spring and before autumn. At or centred on the summer solstice, the earliest sunrise and latest sunset occurs, daylight hours are longest and dark hours are shortest, wit ...
s, which supports characteristic
Mediterranean forests, woodlands, and scrub Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
vegetation.


Geography

The Mediterranean Basin covers portions of three continents:
Europe Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
,
Africa Africa is the world's second-largest and second-most populous continent, after Asia in both cases. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 6% of Earth's total surface area ...
, and
Asia Asia (, ) is one of the world's most notable geographical regions, which is either considered a continent in its own right or a subcontinent of Eurasia, which shares the continental landmass of Afro-Eurasia with Africa. Asia covers an are ...
. It is distinct from the
drainage basin A drainage basin is an area of land where all flowing surface water converges to a single point, such as a river mouth, or flows into another body of water, such as a lake or ocean. A basin is separated from adjacent basins by a perimeter, ...
, which extends much further south and north due to major rivers ending in the Mediterranean Sea, such as the
Nile The Nile, , Bohairic , lg, Kiira , Nobiin: Áman Dawū is a major north-flowing river in northeastern Africa. It flows into the Mediterranean Sea. The Nile is the longest river in Africa and has historically been considered the longest riv ...
and
Rhône The Rhône ( , ; wae, Rotten ; frp, Rôno ; oc, Ròse ) is a major river in France and Switzerland, rising in the Alps and flowing west and south through Lake Geneva and southeastern France before discharging into the Mediterranean Sea. At Ar ...
. Conversely, the Mediterranean Basin includes regions not in the drainage basin. It has a varied and contrasting topography. The Mediterranean Region offers an ever-changing landscape of high mountains, rocky shores, impenetrable scrub, semi-arid steppes, coastal wetlands, sandy beaches and a myriad islands of various shapes and sizes dotted amidst the clear blue sea. Contrary to the classic sandy beach images portrayed in most tourist brochures, the Mediterranean is surprisingly hilly. Mountains can be seen from almost anywhere. By definition, the Mediterranean Basin extends from
Macaronesia Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of Atlantic oceanic islands ...
in the west, to the
Levant The Levant () is an approximate historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology and other cultural contexts, it is ...
in the east, although some places may or may not be included depending on the view, as is the case with Macaronesia: some definitions only include
Madeira ) , anthem = ( en, "Anthem of the Autonomous Region of Madeira") , song_type = Regional anthem , image_map=EU-Portugal_with_Madeira_circled.svg , map_alt=Location of Madeira , map_caption=Location of Madeira , subdivision_type=Sovereign st ...
and the
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
while others include the whole Macaronesia (with the
Azores ) , motto =( en, "Rather die free than subjected in peace") , anthem= ( en, "Anthem of the Azores") , image_map=Locator_map_of_Azores_in_EU.svg , map_alt=Location of the Azores within the European Union , map_caption=Location of the Azores wi ...
and
Cape Verde , national_anthem = () , official_languages = Portuguese , national_languages = Cape Verdean Creole , capital = Praia , coordinates = , largest_city = capital , demonym ...
). In Western Asia, it covers the western and southern portions of the peninsula of Anatolia, as far as
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
, but excluding the temperate-climate mountains of central Turkey. It includes the Mediterranean
Levant The Levant () is an approximate historical geographical term referring to a large area in the Eastern Mediterranean region of Western Asia. In its narrowest sense, which is in use today in archaeology and other cultural contexts, it is ...
at the eastern end of the Mediterranean, bounded on the east and south by the
Syrian Syrians ( ar, سُورِيُّون, ''Sūriyyīn'') are an Eastern Mediterranean ethnic group indigenous to the Levant. They share common Levantine Semitic roots. The cultural and linguistic heritage of the Syrian people is a blend of both indi ...
and
Negev The Negev or Negeb (; he, הַנֶּגֶב, hanNegév; ar, ٱلنَّقَب, an-Naqab) is a desert and semidesert region of southern Israel. The region's largest city and administrative capital is Beersheba (pop. ), in the north. At its sout ...
deserts. The northern portion of the
Maghreb The Maghreb (; ar, الْمَغْرِب, al-Maghrib, lit=the west), also known as the Arab Maghreb ( ar, المغرب العربي) and Northwest Africa, is the western part of North Africa and the Arab world. The region includes Algeria, ...
region of northwestern Africa has a Mediterranean climate, separated from the Sahara Desert, which extends across
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
, by the
Atlas Mountains The Atlas Mountains are a mountain range in the Maghreb in North Africa. It separates the Sahara Desert from the Mediterranean Sea and the Atlantic Ocean; the name "Atlantic" is derived from the mountain range. It stretches around through ...
. In the eastern Mediterranean the Sahara extends to the southern shore of the Mediterranean, with the exception of the northern fringe of the peninsula of
Cyrenaica Cyrenaica ( ) or Kyrenaika ( ar, برقة, Barqah, grc-koi, Κυρηναϊκή ��παρχίαKurēnaïkḗ parkhíā}, after the city of Cyrene), is the eastern region of Libya. Cyrenaica includes all of the eastern part of Libya between ...
in
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Suda ...
, which has a dry Mediterranean climate. Europe lies to the north of the Mediterranean. The European portion of the Mediterranean Basin loosely corresponds to
Southern Europe Southern Europe is the southern region of Europe. It is also known as Mediterranean Europe, as its geography is essentially marked by the Mediterranean Sea. Definitions of Southern Europe include some or all of these countries and regions: Alb ...
. The three large Southern European peninsulas, the
Iberian Peninsula The Iberian Peninsula (), ** * Aragonese and Occitan: ''Peninsula Iberica'' ** ** * french: Péninsule Ibérique * mwl, Península Eibérica * eu, Iberiar penintsula also known as Iberia, is a peninsula in southwestern Europe, def ...
, Italian Peninsula, and the
Balkan Peninsula The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
, extend into and comprise much of the Mediterranean-climate zone. A system of folded mountains, including the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
dividing
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
from
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
, the
Alps The Alps () ; german: Alpen ; it, Alpi ; rm, Alps ; sl, Alpe . are the highest and most extensive mountain range system that lies entirely in Europe, stretching approximately across seven Alpine countries (from west to east): France, Sw ...
dividing
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
from
Central Europe Central Europe is an area of Europe between Western Europe and Eastern Europe, based on a common historical, social and cultural identity. The Thirty Years' War (1618–1648) between Catholicism and Protestantism significantly shaped the a ...
, the
Dinaric Alps The Dinaric Alps (), also Dinarides, are a mountain range in Southern and Southcentral Europe, separating the continental Balkan Peninsula from the Adriatic Sea. They stretch from Italy in the northwest through Slovenia, Croatia, Bosnia and Herz ...
along the eastern
Adriatic The Adriatic Sea () is a body of water separating the Italian Peninsula from the Balkan Peninsula. The Adriatic is the northernmost arm of the Mediterranean Sea, extending from the Strait of Otranto (where it connects to the Ionian Sea) to the ...
, and the
Balkan The Balkans ( ), also known as the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throughout the who ...
and
Rila Rila ( bg, Рила, ) is the highest mountain range of Bulgaria, the Balkan Peninsula and Southeast Europe. It is situated in southwestern Bulgaria and forms part of the Rila– Rhodope Massif. The highest summit is Musala at an elevation of 2, ...
- Rhodope mountains of the Balkan Peninsula divide the Mediterranean from the temperate climate regions of Western, Northwestern or
Northern Europe The northern region of Europe has several definitions. A restrictive definition may describe Northern Europe as being roughly north of the southern coast of the Baltic Sea, which is about 54°N, or may be based on other geographical factors ...
,
Central Europe Central Europe is an area of Europe between Western Europe and Eastern Europe, based on a common historical, social and cultural identity. The Thirty Years' War (1618–1648) between Catholicism and Protestantism significantly shaped the a ...
, and
Eastern Europe Eastern Europe is a subregion of the European continent. As a largely ambiguous term, it has a wide range of geopolitical, geographical, ethnic, cultural, and socio-economic connotations. The vast majority of the region is covered by Russia, whi ...
.


Geology and paleoclimatology

The Mediterranean Basin was shaped by the ancient collision of the northward-moving African–Arabian continent with the stable Eurasian continent. As Africa–Arabia moved north, it closed the former
Tethys Sea The Tethys Ocean ( el, Τηθύς ''Tēthús''), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean that covered most of the Earth during much of the Mesozoic Era and early Cenozoic Era, located between the ancient continents ...
, which formerly separated Eurasia from the ancient super continent of
Gondwana Gondwana () was a large landmass, often referred to as a supercontinent, that formed during the late Neoproterozoic (about 550 million years ago) and began to break up during the Jurassic period (about 180 million years ago). The final sta ...
, of which Africa was part. At about the same time, 170 mya in the
Jurassic The Jurassic ( ) is a geologic period and stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately Mya. The Jurassic constitutes the middle period of ...
period, a small Neotethys ocean basin formed shortly before the Tethys Sea was closed at the eastern end. The collision pushed up a vast system of mountains, extending from the
Pyrenees The Pyrenees (; es, Pirineos ; french: Pyrénées ; ca, Pirineu ; eu, Pirinioak ; oc, Pirenèus ; an, Pirineus) is a mountain range straddling the border of France and Spain. It extends nearly from its union with the Cantabrian Mountains to ...
in Spain to the
Zagros Mountains The Zagros Mountains ( ar, جبال زاغروس, translit=Jibal Zaghrus; fa, کوه‌های زاگرس, Kuh hā-ye Zāgros; ku, چیاکانی زاگرۆس, translit=Çiyakani Zagros; Turkish: ''Zagros Dağları''; Luri: ''Kuh hā-ye Zāgr ...
in
Iran Iran, officially the Islamic Republic of Iran, and also called Persia, is a country located in Western Asia. It is bordered by Iraq and Turkey to the west, by Azerbaijan and Armenia to the northwest, by the Caspian Sea and Turkmeni ...
. This episode of mountain building, known as the Alpine orogeny, occurred mostly during the
Oligocene The Oligocene ( ) is a geologic epoch of the Paleogene Period and extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that define the epoch are well identified but t ...
(34 to 23 million years ago ( mya)) and
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
(23 to 5.3 mya) epochs. The Neotethys became larger during these collisions and associated folding and subduction. About 6 mya during the late Miocene, the Mediterranean was closed at its western end by drifting Africa, which caused the entire sea to evaporate. There followed several (debated) episodes of sea drawdown and re-flooding known as the
Messinian Salinity Crisis The Messinian salinity crisis (MSC), also referred to as the Messinian event, and in its latest stage as the Lago Mare event, was a geological event during which the Mediterranean Sea went into a cycle of partial or nearly complete desiccation (d ...
, which ended when the Atlantic last re-flooded the basin at the end of the Miocene. Recent research has suggested that a desiccation-flooding cycle may have repeated several times during the last 630,000 years of the Miocene epoch, which could explain several events of large amounts of salt deposition. Recent studies, however, show that repeated desiccation and re-flooding is unlikely from a geodynamic point of view. The end of the Miocene also marked a change in the Mediterranean Basin's climate. Fossil evidence shows that the Mediterranean Basin had a relatively humid subtropical climate with summer rainfall during the Miocene, which supported
laurel forest Laurel forest, also called laurisilva or laurissilva, is a type of subtropical forest found in areas with high humidity and relatively stable, mild temperatures. The forest is characterized by broadleaf tree species with evergreen, glossy and elo ...
s. The shift to a Mediterranean climate occurred within the last 3.2–2.8 million years, during the
Pliocene The Pliocene ( ; also Pleiocene) is the epoch in the geologic time scale that extends from 5.333 million to 2.58Macaronesia Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of Atlantic oceanic islands ...
off the Atlantic coast of Iberia and North Africa, and the present Mediterranean vegetation evolved, dominated by coniferous trees and sclerophyllous trees and shrubs, with small, hard, waxy leaves that prevent moisture loss in the dry summers. Much of these forests and shrublands have been altered beyond recognition by thousands of years of human habitation. There are now very few relatively intact natural areas in what was once a heavily wooded region.


Flora and fauna

Phytogeographically Phytogeography (from Greek φυτόν, ''phytón'' = "plant" and γεωγραφία, ''geographía'' = "geography" meaning also distribution) or botanical geography is the branch of biogeography that is concerned with the geographic distribution o ...
, the Mediterranean Basin together with the nearby Atlantic coast, the
Mediterranean woodlands and forests The Mediterranean woodlands and forests is an ecoregion in the coastal plains, hills, and mountains bordering the Mediterranean Sea and Atlantic Ocean in North Africa. It has a Mediterranean climate, and is in the Mediterranean forests, woodlands ...
and
Mediterranean dry woodlands and steppe The Mediterranean dry woodlands and steppe is a Mediterranean forests, woodlands, and scrub ecoregion of North Africa. It occupies interior plateaus and mountain ranges of the Maghreb region, lying generally between the coastal Mediterranean wo ...
of
North Africa North Africa, or Northern Africa is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region, and it is sometimes defined as stretching from the Atlantic shores of Mauritania in ...
, the
Black Sea The Black Sea is a marginal mediterranean sea of the Atlantic Ocean lying between Europe and Asia, east of the Balkans, south of the East European Plain, west of the Caucasus, and north of Anatolia. It is bounded by Bulgaria, Georgia, Rom ...
coast of northeastern
Anatolia Anatolia, tr, Anadolu Yarımadası), and the Anatolian plateau, also known as Asia Minor, is a large peninsula in Western Asia and the westernmost protrusion of the Asian continent. It constitutes the major part of modern-day Turkey. The re ...
, the southern coast of
Crimea Crimea, crh, Къырым, Qırım, grc, Κιμμερία / Ταυρική, translit=Kimmería / Taurikḗ ( ) is a peninsula in Ukraine, on the northern coast of the Black Sea, that has been occupied by Russia since 2014. It has a p ...
between
Sevastopol Sevastopol (; uk, Севасто́поль, Sevastópolʹ, ; gkm, Σεβαστούπολις, Sevastoúpolis, ; crh, Акъя́р, Aqyár, ), sometimes written Sebastopol, is the largest city in Crimea, and a major port on the Black Sea ...
and Feodosiya and the Black Sea coast between
Anapa Anapa (russian: Ана́па, ) is a town in Krasnodar Krai, Russia, located on the northern coast of the Black Sea near the Sea of Azov. Population: History The area around Anapa was settled in antiquity. It was originally a major seaport ...
and
Tuapse Tuapse (russian: Туапсе́; ady, Тӏуапсэ ) is a town in Krasnodar Krai, Russia, situated on the northeast shore of the Black Sea, south of Gelendzhik and north of Sochi. Population: Tuapse is a sea port and the northern center of ...
in
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-ei ...
forms the ''Mediterranean Floristic Region'', which belongs to the Tethyan Subkingdom of the
Boreal Kingdom The Boreal Kingdom or Holarctic Kingdom (Holarctis) is a floristic kingdom identified by botanist Ronald Good (and later by Armen Takhtajan), which includes the temperate to Arctic portions of North America and Eurasia. Its flora is inher ...
and is enclosed between the
Circumboreal The Circumboreal Region in phytogeography is a floristic region within the Holarctic Kingdom in Eurasia and North America, as delineated by such geobotanists as Josias Braun-Blanquet and Armen Takhtajan. It is the largest floristic region in ...
, Irano-Turanian, Saharo-Arabian and
Macaronesian Macaronesia (Portuguese: ''Macaronésia,'' Spanish: ''Macaronesia'') is a collection of four volcanic archipelagos in the North Atlantic, off the coasts of Africa and Europe. Each archipelago is made up of a number of Atlantic oceanic island ...
floristic regions A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both re ...
. The Mediterranean Region was first proposed by German botanist
August Grisebach August Heinrich Rudolf Grisebach () was a German botanist and phytogeographer. He was born in Hannover on 17 April 1814 and died in Göttingen on 9 May 1879. Biography Grisebach studied at the Lyceum in Hanover, the cloister-school at Ilfe ...
in the late 19th century. The
monotypic In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of genera, the term "unispe ...
Drosophyllaceae ''Drosophyllum'' ( , rarely ) is a genus of carnivorous plants containing the single species ''Drosophyllum lusitanicum'', commonly known as Portuguese sundew or dewy pine. In appearance, it is similar to the related genus '' Drosera'' (the sun ...
, recently segregated from
Droseraceae Droseraceae is a family of carnivorous flowering plants, also known as the sundew family. It consists of approximately 180 species in three extant genera. Representatives of the Droseraceae are found on all continents except Antarctica. Desc ...
, is the only plant family
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the region. Among the endemic plant genera are: *''
Anagyris ''Anagyris'' (Spanish: ''oro de risco'') is a genus of flowering plants in the family Fabaceae. It belongs to the subfamily Faboideae. Species ''Anagyris'' comprises the following species: * '' Anagyris foetida'' L. * '' Anagyris latifolia'' Br ...
'' *''
Andryala ''Andryala'' is a genus of flowering plants in the family Asteraceae. It is native to Europe, North Africa, and the Middle East The Middle East ( ar, الشرق الأوسط, ISO 233: ) is a geopolitical region commonly encompassing ...
'' *''
Aphyllanthes ''Aphyllanthes'' is a genus of flowering plants with only one species, ''Aphyllanthes monspeliensis'', endemic to the western Mediterranean region. It is the only genus in the Aphyllanthoideae, a subfamily of the family Family (from la, f ...
'' *''
Argania ''Argania'' (Tashelhit: ⴰⵔⴳⴰⵏ ''Argan'') is a genus of flowering plants containing the sole species ''Argania spinosa'', known as argan, a tree endemic to the calcareous semidesert Sous valley of southwestern Morocco and to the regio ...
'' *''
Argantoniella ''Satureja'' is a genus of aromatic plants of the family Lamiaceae, related to rosemary and thyme. It is native to North Africa, southern and southeastern Europe, the Middle East, and Central Asia. A few New World species were formerly included ...
'' *'' Bellardia'' *'' Biserrula'' *''
Bivonaea ''Bivonaea'' is a genus of flowering plants belonging to the family Brassicaceae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifer ...
'' *''
Bolanthus ''Bolanthus'' is a genus of flowering plants belonging to the family Caryophyllaceae. Its native range is Greece to Jordan. Species: *''Bolanthus aziz-sancarii'' *''Bolanthus cherlerioides'' *''Bolanthus confertifolius'' *''Bolanthus filica ...
'' *'' Boleum'' *'' Callicotome'' *'' Ceratocapnos'' *'' Ceratonia'' *'' Chamaerops'' *''
Chronanthus ''Cytisus'' is a genus of flowering plants in the family Fabaceae, native to open sites (typically scrub and heathland) in Europe, western Asia and North Africa. It belongs to the subfamily Faboideae, and is one of several genera in the tri ...
'' *''
Cladanthus ''Cladanthus'' is a genus of plants in the sunflower family, native to the Mediterranean region. ; Species * '' Cladanthus arabicus'' (L.) Cass. -SoutheasternSpain, Western Sahara, Morocco, Algeria, Libya, Tunisia, Malta, Sicily * '' Cladanthus ...
'' *''
Coridothymus ''Thymbra'', common name Mediterranean thyme, is a genus of plants in the family Lamiaceae. As currently categorized, the genus has seven species and one subspecies. It is native to the Mediterranean region of southern Europe, North Africa, and ...
'' *''
Didesmus ''Didesmus'' is a genus of flowering plants belonging to the family Brassicaceae Brassicaceae () or (the older) Cruciferae () is a medium-sized and economically important family of flowering plants commonly known as the mustards, the crucifers, ...
'' *'' Dorystoechas'' *''
Drosophyllum ''Drosophyllum'' ( , rarely ) is a genus of carnivorous plants containing the single species ''Drosophyllum lusitanicum'', commonly known as Portuguese sundew or dewy pine. In appearance, it is similar to the related genus '' Drosera'' (the sun ...
'' *'' Euzomodendron'' *''
Fedia ''Valeriana'' is a genus of flowering plants in the family Caprifoliaceae, members of which may by commonly known as valerians. It contains many species, including the garden valerian, ''Valeriana officinalis''. Species are native to all contine ...
'' *''
Guiraoa ''Guiraoa'' is a monotypic genus of flowering plants belonging to the family Brassicaceae. It only contains one species, ''Guiraoa arvensis'' Coss. It is native to Spain. The genus name of ''Guiraoa'' is in honour of Ángel Guirao y Navarro (18 ...
'' *'' Gyrocarion'' *''
Helicodiceros ''Helicodiceros muscivorus'', the dead horse arum lily, is an ornamental plant native to Corsica, Sardinia and the Balearic Islands. It is the only species in the genus ''Helicodiceros''. Within the family Araceae the plant is part of the subfa ...
'' *''
Hermodactylus ''Iris tuberosa'' (formerly ''Hermodactylus tuberosus'') is a species of tuberous flowering plant of the genus ''Iris'', with the common names snake's-head, snake's-head iris, widow iris, black iris, or velvet flower-de-luce. Distribution A na ...
'' *'' Hutera'' *'' Hymenocarpus'' *''
Ionopsidium ''Ionopsidium'' is a genus of plants in the family (biology), family Brassicaceae, endemism, endemic to the western Mediterranean region. Species References

* Brassicaceae Brassicaceae genera {{Brassicales-stub ...
'' *''
Lafuentea ''Lafuentea'' is a genus of flowering plants belonging to the family Plantaginaceae. Its native range is the Western Mediterranean, where it is found in Morocco and Spain. The genus name of ''Lafuentea'' is in honour of Tadeo Lafuente (b. c. 1780 ...
'' *'' Lagoecia'' *''
Leuzea ''Rhaponticum'' is a genus of flowering plants in the tribe Cardueae within the family Asteraceae. Description ''Rhaponticum'' species are perennial herbaceous plants with simple, rarely branched stems. The leaves are simple to pinnatifid. Th ...
'' *''
Lycocarpus ''Lycocarpus'' is a genus of flowering plants belonging to the family Brassicaceae. Its native range is Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto ...
'' *''
Malope ''Malope'' is a genus of three species in the mallow botanical family (Malvaceae). Of the three species, '' Malope trifida'' is often used as an ornamental plant Ornamental plants or garden plants are plants that are primarily grown for thei ...
'' *''
Morisia ''Morisia'' is a monotypic genus of ornamental plant in the family Brassicaceae. Its only species is ''Morisia monanthos'', also called ''M. hypogaea''. It is native to the Mediterranean islands of Corsica and Sardinia Sardinia ( ; it, Sa ...
'' *''
Ortegia ''Ortegia'' is a monotypic genus of flowering plants belonging to the family Caryophyllaceae. It only contains one known species, Ortegia hispanica L. It is part of the tribe Polycarpaeae, clustering with '' Cardionema'' and '' Illecebrum''. Its ...
'' *''
Petagnia ''Petagnia'' is a genus of flies in the family Tachinidae. Species *'' Petagnia occlusa'' Rondani, 1856 Distribution Czech Republic, Poland, Bulgaria, Italy, Slovenia, Turkey, Austria, France, Switzerland, Transcaucasia The South Cau ...
'' *''
Petromarula ''Petromarula'' is a genus of plants in the family Campanulaceae. There is only one known species, ''Petromarula pinnata'', endemic to the island of Crete in the Mediterranean. The name "Petromarula" means "rock lettuce" in Greek, a reference to ...
'' *'' Phillyrea'' *''
Preslia Preslia is a peer-reviewed scientific journal publishing original research papers on plant systematics, morphology, phytogeography, ecology and vegetation science, with a geographical focus on central Europe. It has been published by the Cz ...
'' *''
Putoria ''Plocama'' is a genus of flowering plants in the family Rubiaceae. It was described by William Aiton in 1789. It is distributed from the Canary Islands to northwestern India. The genus was expanded in 2007 when several other Rubiaceae gener ...
'' *'' Rothmaleria'' *''
Rosmarinus Rosmarinus ( ) is a small taxonomic clade of woody, perennial herbs with fragrant evergreen needle-like leaves in the family Lamiaceae, native to the Mediterranean Basin. In 2017 the species in the genus ''Rosmarinus'' were moved into the large ...
'' *''
Rupicapnos ''Rupicapnos'' is a genus of flowering plants belonging to the family Papaveraceae The Papaveraceae are an economically important family of about 42 genera and approximately 775 known species of flowering plants in the order Ranunculales, in ...
'' *'' Santolina'' *''
Staehelina ''Staehelina'' is a genus of flowering plants in the tribe Cardueae within the family Asteraceae. Species *'' Staehelina dubia'' L. - western Mediterranean *'' Staehelina petiolata'' ( L.) Hilliard & B.L.Burtt - Crete + Karpathos Island *'' S ...
'' *''
Soleirolia ''Soleirolia soleirolii'' (, syn. ''Helxine soleirolii'') is a plant in the nettle family. It has a number of common names, including baby's tears, angel's tears, peace in the home, bits and pieces, bread and cheese, Corsican creeper, Corsican ...
'' *'' Spartium'' *''
Tetraclinis ''Tetraclinis'' (also called arar, araar or Sictus tree) is a genus of evergreen coniferous trees in the cypress family Cupressaceae, containing only one species, ''Tetraclinis articulata'', also known as Thuja articulata,Trachelium'' *'' Tremastelma'' *''
Triplachne ''Triplachne'' is a genus of seaside plants in the grass family, native to shorelines in the Mediterranean region from the Canary Islands to Turkey. The only known species is ''Triplachne nitens'', called shining grass. It is native to Spain (in ...
'' *'' Vella'' The genera ''
Aubrieta ''Aubrieta'' (often misspelled as Aubretia) is a genus of about 20 species of flowering plants in the cabbage family Brassicaceae. The genus is named after Claude Aubriet, a French flower painter. It originates from southern Europe east to centr ...
'', ''
Sesamoides ''Sesamoides'' is a genus of flowering plants belonging to the family Resedaceae Resedaceae is a family of mostly herbaceous dicotyledonous plants comprising 107 known species in 8 to 12 genera: *'' Borthwickia'' - 1 species, sometimes placed i ...
'', ''
Cynara ''Cynara'' is a genus of thistle-like perennial plants in the family Asteraceae. They are native to the Mediterranean region, the Middle East, northwestern Africa, and the Canary Islands. The genus name comes from the Greek ''kynara'', which me ...
'', '' Dracunculus'', ''
Arisarum ''Arisarum'' is a genus of flowering plants in the family Araceae. It is native to the Mediterranean region, east to the Caucasus and west to Macaronesia. Species Accepted species: Natural Hybrids # '' Arisarum × aspergillum'' Dunal - Spai ...
'' and ''
Biarum ''Biarum'' is a genus of flowering plants in the family Araceae. It is composed of plants that are native to the Middle East, southern Europe (Spain, Portugal, Italy, Balkans), and North Africa. ''Biarum'' are often found growing in rock crev ...
'' are nearly endemic. Among the endemic species prominent in the Mediterranean vegetation are the
Aleppo pine ''Pinus halepensis'', commonly known as the Aleppo pine, also known as the Jerusalem pine, is a pine native to the Mediterranean region. Description ''Pinus halepensis'' is a small to medium-sized tree, tall, with a trunk diameter up to , ex ...
, stone pine,
Mediterranean cypress ''Cupressus sempervirens'', the Mediterranean cypress (also known as Italian cypress, Tuscan cypress, Persian cypress, or pencil pine), is a species of cypress native to the eastern Mediterranean region, in northeast Libya, southern Albania, sou ...
,
bay laurel ''Laurus nobilis'' is an aromatic evergreen tree or large shrub with green, glabrous (smooth) leaves. It is in the flowering plant family Lauraceae. It is native to the Mediterranean region and is used as bay leaf for seasoning in cookin ...
,
Oriental sweetgum ''Liquidambar Orientalis'', commonly known as oriental sweetgum or Turkish sweetgum, is a deciduous tree in the genus ''Liquidambar'', native to the eastern Mediterranean region, that occurs as pure stands mainly in the floodplains of southwest ...
, holm oak,
kermes oak ''Quercus coccifera'', the kermes oak, is an oak bush in the '' Ilex'' section of the genus. It is native to the Mediterranean region and Northern African Maghreb, south to north from Morocco to France and west to east from Portugal to Cyprus ...
, strawberry tree,
Greek strawberry tree Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: * Greeks, an ethnic group. * Greek language, a branch of the Indo-European language family. ** Proto-Greek language, the assumed last common ances ...
, mastic,
terebinth ''Pistacia terebinthus'' also called the terebinth and the turpentine tree, is a deciduous tree species of the genus '' Pistacia'', native to the Mediterranean region from the western regions of Morocco and Portugal to Greece and western an ...
, common myrtle,
oleander ''Nerium oleander'' ( ), most commonly known as oleander or nerium, is a shrub or small tree cultivated worldwide in temperate and subtropical areas as an ornamental and landscaping plant. It is the only species currently classified in the ge ...
, ''
Acanthus mollis ''Acanthus mollis'', commonly known as bear's breeches, sea dock, bear's foot plant, sea holly, gator plant or oyster plant, is a species of plant in the family Acanthaceae and is native to the Mediterranean region. It is a leafy, clump-forming ...
'' and '' Vitex agnus-castus''. Moreover, many plant
taxa In biology, a taxon (back-formation from ''taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular nam ...
are shared with one of the four neighboring floristic regions only. According to different versions of
Armen Takhtajan Armen Leonovich Takhtajan or Takhtajian ( hy, Արմեն Լևոնի Թախտաջյան; russian: Армен Леонович Тахтаджян; surname also transliterated Takhtadjan, Takhtadzhi︠a︡n or Takhtadzhian, pronounced takh-tuh-JA ...
's delineation, the Mediterranean Region is further subdivided into seven to nine floristic provinces: Southwestern Mediterranean (or Southern Moroccan and Southwestern Mediterranean), Ibero-Balearian (or Iberian and Balearian), Liguro-Tyrrhenian, Adriatic,
East Mediterranean Eastern Mediterranean is a loose definition of the eastern approximate half, or third, of the Mediterranean Sea, often defined as the countries around the Levantine Sea. It typically embraces all of that sea's coastal zones, referring to commun ...
, South Mediterranean and Crimeo-Novorossiysk. The Mediterranean Basin is the largest of the world's five
Mediterranean forests, woodlands, and scrub Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
regions. It is home to a number of plant communities, which vary with rainfall, elevation, latitude, and soil. *
Scrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It ...
s occur in the driest areas, especially areas near the seacoast where wind and salt spray are frequent. Low, soft-leaved scrublands around the Mediterranean are known as ''garrigar'' in
Catalan Catalan may refer to: Catalonia From, or related to Catalonia: * Catalan language, a Romance language * Catalans, an ethnic group formed by the people from, or with origins in, Northern or southern Catalonia Places * 13178 Catalan, asteroid #1 ...
, '' garrigue'' in
French French (french: français(e), link=no) may refer to: * Something of, from, or related to France ** French language, which originated in France, and its various dialects and accents ** French people, a nation and ethnic group identified with Franc ...
, ''phrygana'' in
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
, ''tomillares'' in Spanish, and ''batha'' in
Hebrew Hebrew (; ; ) is a Northwest Semitic language of the Afroasiatic language family. Historically, it is one of the spoken languages of the Israelites and their longest-surviving descendants, the Jews and Samaritans. It was largely preserved ...
. *
Shrubland Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It ...
s are dense thickets of evergreen sclerophyll
shrubs A shrub (often also called a bush) is a small-to-medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees ...
and small trees and are the most common plant community around the Mediterranean. Mediterranean shrublands are known as '' màquia'' in Catalan, '' macchia'' in Italian, ''
maquis Maquis may refer to: Resistance groups * Maquis (World War II), predominantly rural guerrilla bands of the French Resistance * Spanish Maquis, guerrillas who fought against Francoist Spain in the aftermath of the Spanish Civil War * The netwo ...
'' in French, and "
matorral 300px, Springtime in Chilean matorral a few kilometers north of Santiago along the Pan-American Highway Matorral is a Spanish language, Spanish word, along with ''tomillares'', for shrubland, thicket or bushes. It is used in naming and describin ...
" in Spanish. In some places, shrublands are the mature vegetation type, and in other places the result of the degradation of former forest or woodland by logging or
overgrazing Overgrazing occurs when plants are exposed to intensive grazing for extended periods of time, or without sufficient recovery periods. It can be caused by either livestock in poorly managed agricultural applications, game reserves, or nature res ...
, or disturbance by major fires. *
Savanna A savanna or savannah is a mixed woodland- grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground ...
s and
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s occur around the Mediterranean, usually dominated by annual
grass Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns a ...
es. *
Woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
s are usually dominated by oak and
pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family (biology), family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. The World Flora Online created by the Royal Botanic Gardens, Kew and Missouri Botanic ...
, mixed with other sclerophyll and coniferous trees. *
Forest A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s are distinct from woodlands in having a closed canopy, and occur in the areas of highest rainfall and in
riparian A riparian zone or riparian area is the interface between land and a river or stream. Riparian is also the proper nomenclature for one of the terrestrial biomes of the Earth. Plant habitats and communities along the river margins and banks a ...
zones along rivers and streams where they receive summer water. Mediterranean forests are generally composed of evergreen trees, predominantly oak and pine. At higher elevations Mediterranean forests transition to mixed broadleaf and tall conifer forests similar to temperate zone forests. The Mediterranean Basin is home to considerable
biodiversity Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity'') ...
, including 22,500
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
.
Conservation International Conservation International (CI) is an American nonprofit environmental organization headquartered in Crystal City, Arlington, Virginia. CI's work focuses on science, policy and partnership with businesses, governments and communities. The org ...
designates the region as a biodiversity hotspot, because of its rich biodiversity and its threatened status. The Mediterranean Basin has an area of 2,085,292 km2, of which only 98,009 km2 remains undisturbed. Endangered
mammal Mammals () are a group of vertebrate animals constituting the class Mammalia (), characterized by the presence of mammary glands which in females produce milk for feeding (nursing) their young, a neocortex (a region of the brain), fur ...
s of the Mediterranean Basin include the Mediterranean monk seal, the
Barbary macaque The Barbary macaque (''Macaca sylvanus''), also known as Barbary ape, is a macaque species native to the Atlas Mountains of Algeria, Libya, Tunisia and Morocco, along with a small introduced population in Gibraltar. It is the type species of the ...
, and the
Iberian lynx The Iberian lynx (''Lynx pardinus'') is a wild cat species endemic to the Iberian Peninsula in southwestern Europe. It is listed as Endangered on the IUCN Red List. In the 20th century, the Iberian lynx population had declined because of overhun ...
.


Ecoregions

The WWF identifies 22
Mediterranean forests, woodlands, and scrub Mediterranean forests, woodlands, and scrub is a biome defined by the World Wide Fund for Nature. The biome is generally characterized by dry summers and rainy winters, although in some areas rainfall may be uniform. Summers are typically hot in ...
ecoregions in the Mediterranean Basin, most of which featuring
sclerophyll Sclerophyll is a type of vegetation that is adapted to long periods of dryness and heat. The plants feature hard leaves, short internodes (the distance between leaves along the stem) and leaf orientation which is parallel or oblique to direct ...
plant species: * Aegean and Western Turkey sclerophyllous and mixed forests (
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
,
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...
,
Bulgaria Bulgaria (; bg, България, Bǎlgariya), officially the Republic of Bulgaria,, ) is a country in Southeast Europe. It is situated on the eastern flank of the Balkans, and is bordered by Romania to the north, Serbia and North Macedo ...
) * Anatolian conifer and deciduous mixed forests (
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
) * Canary Islands dry woodlands and forests (
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) *
Corsican montane broadleaf and mixed forests The Corsican montane broadleaf and mixed forests ecoregion, in the Mediterranean forests, woodlands, and scrub biome, are on the island of Corsica. The ecoregion includes the high-altitude regions of Corsica's mountain ranges. Setting The ecoregi ...
(
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
) *
Crete Mediterranean forests The Crete Mediterranean forests is a terrestrial ecoregion that encompasses Greek the island of Crete. The island has a Mediterranean climate, and is in the Mediterranean forests, woodlands, and scrub biome found in the lands in and around the Me ...
(
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
) *
Cyprus Mediterranean forests The Cyprus Mediterranean forests is a terrestrial ecoregion that encompasses the island of Cyprus. The island has a Mediterranean climate, and is in the Mediterranean forests, woodlands, and scrub biome found in the lands in and around the Medit ...
(
Cyprus Cyprus ; tr, Kıbrıs (), officially the Republic of Cyprus,, , lit: Republic of Cyprus is an island country located south of the Anatolian Peninsula in the eastern Mediterranean Sea. Its continental position is disputed; while it is ...
) * Eastern Mediterranean conifer–sclerophyllous–broadleaf forests (
Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus lie ...
,
Iraq Iraq,; ku, عێراق, translit=Êraq officially the Republic of Iraq, '; ku, کۆماری عێراق, translit=Komarî Êraq is a country in Western Asia. It is bordered by Turkey to Iraq–Turkey border, the north, Iran to Iran–Iraq ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
, Palestine,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
) *
Iberian conifer forests The Iberian conifer forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It includes the mountain forests of southern and central Spain. The ecoregion has montane Mediterranean climate. Rainfall averages 1,10 ...
(
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) *
Iberian sclerophyllous and semi-deciduous forests The Iberian sclerophyllous and semi-deciduous forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It occupies the interior valleys and plateaus of the Iberian Peninsula. The ecoregion lies mostly in Spain, an ...
(
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of th ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) * Illyrian deciduous forests (
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and share ...
,
Bosnia and Herzegovina Bosnia and Herzegovina ( sh, / , ), abbreviated BiH () or B&H, sometimes called Bosnia–Herzegovina and often known informally as Bosnia, is a country at the crossroads of south and southeast Europe, located in the Balkans. Bosnia and ...
,
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = " Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capi ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Montenegro ) , image_map = Europe-Montenegro.svg , map_caption = , image_map2 = , capital = Podgorica , coordinates = , largest_city = capital , official_languages = ...
,
Slovenia Slovenia ( ; sl, Slovenija ), officially the Republic of Slovenia (Slovene: , abbr.: ''RS''), is a country in Central Europe. It is bordered by Italy to the west, Austria to the north, Hungary to the northeast, Croatia to the southeast, and ...
) * Italian sclerophyllous and semi-deciduous forests (
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
San Marino San Marino (, ), officially the Republic of San Marino ( it, Repubblica di San Marino; ), also known as the Most Serene Republic of San Marino ( it, Serenissima Repubblica di San Marino, links=no), is the fifth-smallest country in the world an ...
) * Mediterranean acacia–argania dry woodlands (
Western Sahara Western Sahara ( '; ; ) is a disputed territory on the northwest coast and in the Maghreb region of North and West Africa. About 20% of the territory is controlled by the self-proclaimed Sahrawi Arab Democratic Republic (SADR), while the ...
,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
,
Canary Islands The Canary Islands (; es, :es:Canarias, Canarias, ), also known informally as the Canaries, are a Spanish Autonomous communities of Spain, autonomous community and archipelago in the Atlantic Ocean, in Macaronesia. At their closest point to ...
(
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
)) *
Mediterranean dry woodlands and steppe The Mediterranean dry woodlands and steppe is a Mediterranean forests, woodlands, and scrub ecoregion of North Africa. It occupies interior plateaus and mountain ranges of the Maghreb region, lying generally between the coastal Mediterranean wo ...
(
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
,
Egypt Egypt ( ar, مصر , ), officially the Arab Republic of Egypt, is a List of transcontinental countries, transcontinental country spanning the North Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via a land bridg ...
,
Libya Libya (; ar, ليبيا, Lībiyā), officially the State of Libya ( ar, دولة ليبيا, Dawlat Lībiyā), is a country in the Maghreb region in North Africa. It is bordered by the Mediterranean Sea to the north, Egypt to the east, Suda ...
,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
,
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
) *
Mediterranean woodlands and forests The Mediterranean woodlands and forests is an ecoregion in the coastal plains, hills, and mountains bordering the Mediterranean Sea and Atlantic Ocean in North Africa. It has a Mediterranean climate, and is in the Mediterranean forests, woodlands ...
(
Algeria ) , image_map = Algeria (centered orthographic projection).svg , map_caption = , image_map2 = , capital = Algiers , coordinates = , largest_city = capital , relig ...
,
Morocco Morocco (),, ) officially the Kingdom of Morocco, is the westernmost country in the Maghreb region of North Africa. It overlooks the Mediterranean Sea to the north and the Atlantic Ocean to the west, and has land borders with Algeria to A ...
,
Tunisia ) , image_map = Tunisia location (orthographic projection).svg , map_caption = Location of Tunisia in northern Africa , image_map2 = , capital = Tunis , largest_city = capital , ...
) *
Northeastern Spain and Southern France Mediterranean forests The Northeastern Spain and Southern France Mediterranean forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It occupies the Mediterranean coastal region of northeastern Spain and Southern France, and the B ...
(
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Monaco Monaco (; ), officially the Principality of Monaco (french: Principauté de Monaco; Ligurian: ; oc, Principat de Mónegue), is a sovereign city-state and microstate on the French Riviera a few kilometres west of the Italian region of Lig ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) * Northwest Iberian montane forests (
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of th ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) * Pindus Mountains mixed forests (
Albania Albania ( ; sq, Shqipëri or ), or , also or . officially the Republic of Albania ( sq, Republika e Shqipërisë), is a country in Southeastern Europe. It is located on the Adriatic and Ionian Seas within the Mediterranean Sea and share ...
,
Greece Greece,, or , romanized: ', officially the Hellenic Republic, is a country in Southeast Europe. It is situated on the southern tip of the Balkans, and is located at the crossroads of Europe, Asia, and Africa. Greece shares land borders wi ...
,
North Macedonia North Macedonia, ; sq, Maqedonia e Veriut, (Macedonia before February 2019), officially the Republic of North Macedonia,, is a country in Southeast Europe. It gained independence in 1991 as one of the successor states of Socialist Feder ...
) * South Apennine mixed montane forests (
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
) *
Southeastern Iberian shrubs and woodlands The Southeastern Iberian shrubs and woodlands is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It lies southwestern coastal Spain, along the Mediterranean Sea. Geography The ecoregion covers coastal low plains, ...
(
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) * Southern Anatolian montane conifer and deciduous forests (
Lebanon Lebanon ( , ar, لُبْنَان, translit=lubnān, ), officially the Republic of Lebanon () or the Lebanese Republic, is a country in Western Asia. It is located between Syria to the north and east and Israel to the south, while Cyprus lie ...
,
Israel Israel (; he, יִשְׂרָאֵל, ; ar, إِسْرَائِيل, ), officially the State of Israel ( he, מְדִינַת יִשְׂרָאֵל, label=none, translit=Medīnat Yīsrāʾēl; ), is a country in Western Asia. It is situated ...
,
Jordan Jordan ( ar, الأردن; tr. ' ), officially the Hashemite Kingdom of Jordan,; tr. ' is a country in Western Asia. It is situated at the crossroads of Asia, Africa, and Europe, within the Levant region, on the East Bank of the Jordan Rive ...
,
Syria Syria ( ar, سُورِيَا or سُورِيَة, translit=Sūriyā), officially the Syrian Arab Republic ( ar, الجمهورية العربية السورية, al-Jumhūrīyah al-ʻArabīyah as-Sūrīyah), is a Western Asian country loc ...
,
Turkey Turkey ( tr, Türkiye ), officially the Republic of Türkiye ( tr, Türkiye Cumhuriyeti, links=no ), is a transcontinental country located mainly on the Anatolian Peninsula in Western Asia, with a small portion on the Balkan Peninsula ...
) *
Southwest Iberian Mediterranean sclerophyllous and mixed forests The Southwest Iberian Mediterranean sclerophyllous and mixed forests is a Mediterranean forests, woodlands, and scrub ecoregion in southwestern Europe. It occupies the southwestern Iberian Peninsula, encompassing coastal lowlands and mountains in ...
(
Portugal Portugal, officially the Portuguese Republic ( pt, República Portuguesa, links=yes ), is a country whose mainland is located on the Iberian Peninsula of Southwestern Europe, and whose territory also includes the Atlantic archipelagos of th ...
,
Spain , image_flag = Bandera de España.svg , image_coat = Escudo de España (mazonado).svg , national_motto = '' Plus ultra'' (Latin)(English: "Further Beyond") , national_anthem = (English: "Royal March") , ...
) * Tyrrhenian–Adriatic sclerophyllous and mixed forests (
Croatia , image_flag = Flag of Croatia.svg , image_coat = Coat of arms of Croatia.svg , anthem = " Lijepa naša domovino"("Our Beautiful Homeland") , image_map = , map_caption = , capi ...
,
France France (), officially the French Republic ( ), is a country primarily located in Western Europe. It also comprises of Overseas France, overseas regions and territories in the Americas and the Atlantic Ocean, Atlantic, Pacific Ocean, Pac ...
,
Italy Italy ( it, Italia ), officially the Italian Republic, ) or the Republic of Italy, is a country in Southern Europe. It is located in the middle of the Mediterranean Sea, and its territory largely coincides with the homonymous geographical ...
,
Malta Malta ( , , ), officially the Republic of Malta ( mt, Repubblika ta' Malta ), is an island country in the Mediterranean Sea. It consists of an archipelago, between Italy and Libya, and is often considered a part of Southern Europe. It lies ...
) .


History

Neanderthal Neanderthals (, also ''Homo neanderthalensis'' and erroneously ''Homo sapiens neanderthalensis''), also written as Neandertals, are an Extinction, extinct species or subspecies of archaic humans who lived in Eurasia until about 40,000 years ag ...
s inhabited western Asia and the non-glaciated portions of Europe starting about 230,000 years ago. Modern humans moved into western Asia from Africa less than 100,000 years ago. Modern humans, known as
Cro-Magnon Early European modern humans (EEMH), or Cro-Magnons, were the first early modern humans (''Homo sapiens'') to settle in Europe, migrating from Western Asia, continuously occupying the continent possibly from as early as 56,800 years ago. They i ...
s, moved into Europe approximately 50–40,000 years ago. The most recent glacial period, the
Wisconsin glaciation The Wisconsin Glacial Episode, also called the Wisconsin glaciation, was the most recent glacial period of the North American ice sheet complex. This advance included the Cordilleran Ice Sheet, which nucleated in the northern North American Cord ...
, reached its maximum extent approximately 21,000 years ago, and ended approximately 12,000 years ago. A warm period, known as the
Holocene climatic optimum The Holocene Climate Optimum (HCO) was a warm period that occurred in the interval roughly 9,000 to 5,000 years ago BP, with a thermal maximum around 8000 years BP. It has also been known by many other names, such as Altithermal, Climatic Optimu ...
, followed the ice age. Food crops, including
wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
,
chickpea The chickpea or chick pea (''Cicer arietinum'') is an annual legume of the family Fabaceae, subfamily Faboideae. Its different types are variously known as gram" or Bengal gram, garbanzo or garbanzo bean, or Egyptian pea. Chickpea seeds are h ...
s, and
olive The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' ' ...
s, along with
sheep Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticate ...
and
goat The goat or domestic goat (''Capra hircus'') is a domesticated species of goat-antelope typically kept as livestock. It was domesticated from the wild goat (''C. aegagrus'') of Southwest Asia and Eastern Europe. The goat is a member of the a ...
s, were domesticated in the eastern Mediterranean in the 9th millennium BCE, which allowed for the establishment of agricultural settlements. Near Eastern crops spread to southeastern Europe in the 7th millennium BCE. Poppy and oats were domesticated in Europe from the 6th to the 3rd millennium BCE. Agricultural settlements spread around the Mediterranean Basin.
Megalith A megalith is a large stone that has been used to construct a prehistoric structure or monument, either alone or together with other stones. There are over 35,000 in Europe alone, located widely from Sweden to the Mediterranean sea. The ...
s were constructed in Europe from 4500 – 1500 BCE. A strengthening of the summer
monsoon A monsoon () is traditionally a seasonal reversing wind accompanied by corresponding changes in precipitation but is now used to describe seasonal changes in atmospheric circulation and precipitation associated with annual latitudinal oscil ...
9000–7000 years ago increased rainfall across the
Sahara , photo = Sahara real color.jpg , photo_caption = The Sahara taken by Apollo 17 astronauts, 1972 , map = , map_image = , location = , country = , country1 = , ...
, which became a
grassland A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
, with lakes, rivers, and wetlands. After a period of climatic instability, the Sahara settled into a desert state by the 4th millennium BCE.


Historiography

One of the earliest modern studies of the Mediterranean was
Fernand Braudel Fernand Braudel (; 24 August 1902 – 27 November 1985) was a French historian and leader of the Annales School. His scholarship focused on three main projects: ''The Mediterranean'' (1923–49, then 1949–66), ''Civilization and Capitalism'' ...
's ''La Méditerranéee et le monde méditerranéen à l époque de Philippe II'' (The Mediterranean and the Mediterranean World in the Age of Philip II), published in 1949.
S.D. Goitein Shelomo Dov Goitein (April 3, 1900 – February 6, 1985) was a German-Jewish ethnographer, historian and Arabist known for his research on Jewish life in the Islamic Middle Ages, and particularly on the Cairo Geniza. Biography Shelomo Dov (Fritz ...
's multivolume study of the
Cairo Geniza The Cairo Geniza, alternatively spelled Genizah, is a collection of some 400,000 Jewish manuscript fragments and Fatimid administrative documents that were kept in the ''genizah'' or storeroom of the Ben Ezra Synagogue in Fustat or Old Cairo, ...
documents was another important contribution in the area of Mediterranean Jewish culture.


Agriculture

Wheat Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
is the dominant grain grown around the Mediterranean Basin.
Pulses In medicine, a pulse represents the tactile arterial palpation of the cardiac cycle (heartbeat) by trained fingertips. The pulse may be palpated in any place that allows an artery to be compressed near the surface of the body, such as at the nec ...
and
vegetable Vegetables are parts of plants that are consumed by humans or other animals as food. The original meaning is still commonly used and is applied to plants collectively to refer to all edible plant matter, including the edible flower, flowers, ...
s are also grown. The characteristic tree crop is the
olive The olive, botanical name ''Olea europaea'', meaning 'European olive' in Latin, is a species of small tree or shrub in the family Oleaceae, found traditionally in the Mediterranean Basin. When in shrub form, it is known as ''Olea europaea'' ' ...
. Figs are another important fruit tree, and
citrus ''Citrus'' is a genus of flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, lemons, grapefruits, pomelos, and limes. The genus ''Citrus'' is native to ...
, especially
lemon The lemon (''Citrus limon'') is a species of small evergreen trees in the flowering plant family Rutaceae, native to Asia, primarily Northeast India (Assam), Northern Myanmar or China. The tree's ellipsoidal yellow fruit is used for culin ...
s, are grown where irrigation is present.
Grape A grape is a fruit, botanically a berry, of the deciduous woody vines of the flowering plant genus '' Vitis''. Grapes are a non- climacteric type of fruit, generally occurring in clusters. The cultivation of grapes began perhaps 8,000 years a ...
s are an important
vine A vine (Latin ''vīnea'' "grapevine", "vineyard", from ''vīnum'' "wine") is any plant with a growth habit of trailing or scandent (that is, climbing) stems, lianas or runners. The word ''vine'' can also refer to such stems or runners themsel ...
crop, grown for fruit and to make
wine Wine is an alcoholic drink typically made from Fermentation in winemaking, fermented grapes. Yeast in winemaking, Yeast consumes the sugar in the grapes and converts it to ethanol and carbon dioxide, releasing heat in the process. Different ...
.
Rice Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and '' Porteresia'', both wild and domesticat ...
and summer vegetables are grown in irrigated areas.


See also

* Ancient Egypt *
Ancient Greece Ancient Greece ( el, Ἑλλάς, Hellás) was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity ( AD 600), that comprised a loose collection of cu ...
*
Life zones of the Mediterranean region The climate and ecology of land immediately surrounding the Mediterranean Sea is influenced by several factors. Overall, the land has a Mediterranean climate, with mild, rainy winters and hot, dry summers. The climate induces characteristic Medit ...
* Mediterranean wine climate *
MISTRALS MISTRALS ("Integrated Mediterranean Studies at Regional and Local Scales") is a research program dedicated to the study of the Mediterranean basin and its surroundings, with the aim to "better understand the impact of global factors on this region ...
*
Ottoman Empire The Ottoman Empire, * ; is an archaic version. The definite article forms and were synonymous * and el, Оθωμανική Αυτοκρατορία, Othōmanikē Avtokratoria, label=none * info page on book at Martin Luther University ...
*
Phoenicia Phoenicia () was an ancient thalassocratic civilization originating in the Levant region of the eastern Mediterranean, primarily located in modern Lebanon. The territory of the Phoenician city-states extended and shrank throughout their his ...
*
Roman Empire The Roman Empire ( la, Imperium Romanum ; grc-gre, Βασιλεία τῶν Ῥωμαίων, Basileía tôn Rhōmaíōn) was the post-Roman Republic, Republican period of ancient Rome. As a polity, it included large territorial holdings aro ...
* Zanclean flood


References


Further reading

* * Borutta, Manuel,
Mediterraneum
'
EGO - European History Online
Mainz
Institute of European History
2021, retrieved: March 8, 2021. * * *


External links


Mediterranean Basin biodiversity hotspot (Conservation International)Are wildfires a disaster in the Mediterranean basin? – A reviewMedTrees: Trees and large shrubs of the Mediterranean Basin.
{{Authority control Mediterranean forests, woodlands, and scrub Palearctic realm Palearctic ecoregions Natural history of Europe Levant Floristic regions Regions of Europe Geography of North Africa Geography of Southern Europe Geography of Western Asia Geography of the Middle East Regions of Eurasia Regions of Africa