Marine habitats are
habitat
In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
s that support
marine life
Marine life, sea life, or ocean life is the plants, animals and other organisms that live in the salt water of seas or oceans, or the brackish water of coastal estuaries. At a fundamental level, marine life affects the nature of the planet. ...
. Marine life depends in some way on the
saltwater that is in the sea (the term ''marine'' comes from the
Latin
Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
''mare'', meaning sea or ocean). A habitat is an
ecological
Ecology () is the study of the relationships between living organisms, including humans, and their physical environment. Ecology considers organisms at the individual, population, community, ecosystem, and biosphere level. Ecology overlaps wi ...
or
environmental
A biophysical environment is a biotic and abiotic surrounding of an organism or population, and consequently includes the factors that have an influence in their survival, development, and evolution. A biophysical environment can vary in scal ...
area inhabited by one or more living
species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
.
[Abercrombie, M., Hickman, C.J. and Johnson, M.L. 1966.''A Dictionary of Biology.'' Penguin Reference Books, London] The marine environment supports many kinds of these habitats. Marine habitats can be divided into
coastal
The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in ...
and
open ocean habitats. Coastal habitats are found in the area that extends from as far as the tide comes in on the
shoreline
A shore or a shoreline is the fringe of land at the edge of a large body of water, such as an ocean, sea, or lake. In physical oceanography, a shore is the wider fringe that is geologically modified by the action of the body of water past a ...
out to the edge of the
continental shelf
A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
. Most marine life is found in coastal habitats, even though the shelf area occupies only seven percent of the total ocean area. Open ocean habitats are found in the deep ocean beyond the edge of the continental shelf.
Alternatively, marine habitats can be divided into
pelagic
The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
and
demersal zone
The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer of ...
s. Pelagic habitats are found near the surface or in the open
water column
A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
, away from the bottom of the ocean.
Demersal
The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer ...
habitats are near or on the bottom of the ocean. An organism living in a pelagic habitat is said to be a pelagic organism, as in
pelagic fish
Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral re ...
. Similarly, an organism living in a demersal habitat is said to be a demersal organism, as in
demersal fish
Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
. Pelagic habitats are intrinsically shifting and ephemeral, depending on what
ocean current
An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth conto ...
s are doing.
Marine habitats can be modified by their inhabitants. Some marine organisms, like
coral
Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...
s,
kelp
Kelps are large brown algae seaweeds that make up the order Laminariales. There are about 30 different genera. Despite its appearance, kelp is not a plant - it is a heterokont, a completely unrelated group of organisms.
Kelp grows in "underwa ...
,
mangrove
A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...
s and
seagrass
Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families ( Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the ...
es, are
ecosystem engineer
An ecosystem engineer is any species that creates, significantly modifies, maintains or destroys a habitat. These organisms can have a large impact on species richness and landscape-level heterogeneity of an area. As a result, ecosystem engin ...
s which reshape the marine environment to the point where they create further habitat for other organisms. By volume the ocean provides most of the habitable space on the planet.
__TOC__
Overview
In contrast to terrestrial habitats, marine habitats are shifting and
ephemeral
Ephemerality (from the Greek word , meaning 'lasting only one day') is the concept of things being transitory, existing only briefly. Academically, the term ephemeral constitutionally describes a diverse assortment of things and experiences, f ...
. Swimming organisms find areas by the edge of a
continental shelf
A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
a good habitat, but only while
upwelling
Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted ...
s bring nutrient rich water to the surface. Shellfish find habitat on sandy beaches, but storms, tides and currents mean their habitat continually reinvents itself.
The presence of
seawater
Seawater, or salt water, is water from a sea or ocean. On average, seawater in the world's oceans has a salinity of about 3.5% (35 g/L, 35 ppt, 600 mM). This means that every kilogram (roughly one liter by volume) of seawater has appro ...
is common to all marine habitats. Beyond that many other things determine whether a marine area makes a good habitat and the type of habitat it makes. For example:
* temperature – is affected by geographical
latitude
In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north ...
,
ocean currents
An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth contou ...
,
weather
Weather is the state of the atmosphere, describing for example the degree to which it is hot or cold, wet or dry, calm or stormy, clear or cloudy. On Earth, most weather phenomena occur in the lowest layer of the planet's atmosphere, the ...
, the discharge of rivers, and by the presence of
hydrothermal vent
A hydrothermal vent is a fissure on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hotspo ...
s or
cold seep
A cold seep (sometimes called a cold vent) is an area of the ocean floor where hydrogen sulfide, methane and other hydrocarbon-rich fluid seepage occurs, often in the form of a brine pool. ''Cold'' does not mean that the temperature of the see ...
s
* sunlight –
photosynthetic
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
processes depend on how deep and
turbid
Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality.
Fluids c ...
the water is
* nutrients – are transported by ocean currents to different marine habitats from
land runoff, or by upwellings from the deep sea, or they sink through the sea as
marine snow
* salinity – varies, particularly in
estuaries
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
or near
river delta
A river delta is a landform shaped like a triangle, created by deposition of sediment that is carried by a river and enters slower-moving or stagnant water. This occurs where a river enters an ocean, sea, estuary, lake, reservoir, or (more rare ...
s, or by hydrothermal vents
* dissolved gases – oxygen levels in particular, can be increased by wave actions and decreased during
algal blooms
* acidity – this is partly to do with dissolved gases above, since the acidity of the ocean is largely controlled by how much carbon dioxide is in the water.
* turbulence –
ocean wave
In fluid dynamics, a wind wave, water wave, or wind-generated water wave, is a surface wave that occurs on the free surface of bodies of water as a result from the wind blowing over the water surface. The contact distance in the direction of t ...
s, fast currents and the agitation of water affect the nature of habitats
* cover – the availability of cover such as the adjacency of the
sea bottom, or the presence of
floating objects
* substrate – The slope, orientation, profile and
rugosity of hard substrates, and particle size,
sorting
Sorting refers to ordering data in an increasing or decreasing manner according to some linear relationship among the data items.
# ordering: arranging items in a sequence ordered by some criterion;
# categorizing: grouping items with similar pro ...
and density of
unconsolidated
Soil consolidation refers to the mechanical process by which soil changes volume gradually in response to a change in pressure. This happens because soil is a two-phase material, comprising soil grains and pore fluid, usually groundwater. Whe ...
sediment bottoms can make a big difference to the life forms that can settle on it.
* the occupying organisms themselves – since organisms modify their habitats by the act of occupying them, and some, like corals, kelp, mangroves and seagrasses, create further habitats for other organisms.
There are five major oceans, of which the
Pacific Ocean
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the conti ...
is nearly as large as the rest put together. Coastlines fringe the land for nearly 380,000 kilometres.
Altogether, the ocean occupies 71 percent of the world surface, averaging nearly four kilometres in depth. By volume, the ocean contains more than 99 percent of the Earth's liquid water.
[Where is Earth's water?](_blank)
United States Geological Survey
The United States Geological Survey (USGS), formerly simply known as the Geological Survey, is a scientific agency of the United States government. The scientists of the USGS study the landscape of the United States, its natural resources, ...
. The science fiction writer
Arthur C. Clarke
Sir Arthur Charles Clarke (16 December 191719 March 2008) was an English science-fiction writer, science writer, futurist, inventor, undersea explorer, and television series host.
He co-wrote the screenplay for the 1968 film '' 2001: A Spac ...
has pointed out it would be more appropriate to refer to the planet Earth as the planet Sea or the planet
Ocean.
Marine habitats can be broadly divided into
pelagic
The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
and
demersal
The demersal zone is the part of the sea or ocean (or deep lake) consisting of the part of the water column near to (and significantly affected by) the seabed and the benthos. The demersal zone is just above the benthic zone and forms a layer ...
habitats.
Pelagic
The pelagic zone consists of the water column of the open ocean, and can be further divided into regions by depth (as illustrated on the right). The word ''pelagic'' is derived . The pelagic zone can be thought of as an imaginary cylinder or w ...
habitats are the habitats of the open
water column
A water column is a conceptual column of water from the surface of a sea, river or lake to the bottom sediment.Munson, B.H., Axler, R., Hagley C., Host G., Merrick G., Richards C. (2004).Glossary. ''Water on the Web''. University of Minnesota-D ...
, away from the bottom of the ocean. Demersal habitats are the habitats that are near or on the bottom of the ocean. An organism living in a pelagic habitat is said to be a pelagic organism, as in
pelagic fish
Pelagic fish live in the pelagic zone of ocean or lake waters—being neither close to the bottom nor near the shore—in contrast with demersal fish that do live on or near the bottom, and reef fish that are associated with coral re ...
. Similarly, an organism living in a demersal habitat is said to be a demersal organism, as in
demersal fish
Demersal fish, also known as groundfish, live and feed on or near the bottom of seas or lakes (the demersal zone).Walrond Carl . "Coastal fish - Fish of the open sea floor"Te Ara - the Encyclopedia of New Zealand. Updated 2 March 2009 They oc ...
. Pelagic habitats are intrinsically ephemeral, depending on what
ocean current
An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth conto ...
s are doing.
The
land-based ecosystem depends on topsoil and fresh water, while the
marine ecosystem
Marine ecosystems are the largest of Earth's aquatic ecosystems and exist in waters that have a high salt content. These systems contrast with freshwater ecosystems, which have a lower salt content. Marine waters cover more than 70% of the sur ...
depends on dissolved nutrients washed down from the land.
Ocean deoxygenation poses a threat to marine habitats, due to the growth of low oxygen zones.
Ocean currents
In marine systems,
ocean current
An ocean current is a continuous, directed movement of sea water generated by a number of forces acting upon the water, including wind, the Coriolis effect, breaking waves, cabbeling, and temperature and salinity differences. Depth conto ...
s have a key role determining which areas are effective as habitats, since ocean currents transport the basic nutrients needed to support marine life.
[Ocean Habitats](_blank)
''Marietta College''. Retrieved 17 April 2011. Plankton
Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...
are the life forms that inhabit the ocean that are so small (less than 2 mm) that they cannot effectively propel themselves through the water, but must drift instead with the currents. If the current carries the right nutrients, and if it also flows at a suitably shallow depth where there is plenty of sunlight, then such a current itself can become a suitable habitat for photosynthesizing tiny algae called
phytoplankton
Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'.
...
. These tiny plants are the
primary producers in the ocean, at the start of the
food chain. In turn, as the population of drifting phytoplankton grows, the water becomes a suitable habitat for
zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
, which feed on the phytoplankton. While phytoplankton are tiny drifting plants,
zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
are tiny drifting animals, such as the
larvae
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
T ...
of
fish
Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
and
marine invertebrates. If sufficient zooplankton establish themselves, the current becomes a candidate habitat for the
forage fish
Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the f ...
that feed on them. And then if sufficient forage fish move to the area, it becomes a candidate habitat for larger
predatory fish
Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, pike/ muskellunge, walleye, perch and salmon. Some omnivorous fish, such as the red-bellie ...
and other marine animals that feed on the forage fish. In this dynamic way, the current itself can, over time, become a moving habitat for multiple types of marine life.
Ocean currents can be generated by differences in the density of the water. How dense water is depends on how
saline or warm it is. If water contains differences in salt content or temperature, then the different densities will initiate a current. Water that is saltier or cooler will be denser, and will sink in relation to the surrounding water. Conversely, warmer and less salty water will float to the surface. Atmospheric winds and pressure differences also produces surface currents,
waves
Waves most often refers to:
* Waves, oscillations accompanied by a transfer of energy that travel through space or mass.
* Wind waves, surface waves that occur on the free surface of bodies of water.
Waves may also refer to:
Music
*Waves (ban ...
and
seiche
A seiche ( ) is a standing wave in an enclosed or partially enclosed body of water. Seiches and seiche-related phenomena have been observed on lakes, reservoirs, swimming pools, bays, harbors, caves and seas. The key requirement for formation of ...
s. Ocean currents are also generated by the gravitational pull of the sun and moon (
tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables ...
s), and seismic activity (
tsunami
A tsunami ( ; from ja, 津波, lit=harbour wave, ) is a series of waves in a water body caused by the displacement of a large volume of water, generally in an ocean or a large lake. Earthquakes, volcanic eruptions and other underwater exp ...
).
The rotation of the Earth affects the direction ocean currents take, and explains which way the large circular
ocean gyre
In oceanography, a gyre () is any large system of circulating ocean currents, particularly those involved with large wind movements. Gyres are caused by the Coriolis effect; planetary vorticity, horizontal friction and vertical friction determi ...
s rotate in the image above left. Suppose a current at the equator is heading north. The Earth rotates eastward, so the water possesses that rotational momentum. But the further the water moves north, the slower the earth moves eastward. If the current could get to the North Pole, the earth wouldn't be moving eastward at all. To conserve its rotational momentum, the further the current travels north the faster it must move eastward. So the effect is that the current curves to the right. This is the
Coriolis effect
In physics, the Coriolis force is an inertial or fictitious force that acts on objects in motion within a frame of reference that rotates with respect to an inertial frame. In a reference frame with clockwise rotation, the force acts to the ...
. It is weakest at the equator and strongest at the poles. The effect is opposite south of the equator, where currents curve left.
Topography
Biomass
One measure of the relative importance of different marine habitats is the rate at which they produce
biomass
Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bio ...
.
Coastal
Marine coasts are dynamic environments which constantly change, like the ocean which partially shape them. The Earth's natural processes, including
weather
Weather is the state of the atmosphere, describing for example the degree to which it is hot or cold, wet or dry, calm or stormy, clear or cloudy. On Earth, most weather phenomena occur in the lowest layer of the planet's atmosphere, the ...
and
sea level change, result in the
erosion
Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is d ...
,
accretion
Accretion may refer to:
Science
* Accretion (astrophysics), the formation of planets and other bodies by collection of material through gravity
* Accretion (meteorology), the process by which water vapor in clouds forms water droplets around nucl ...
and resculpturing of coasts as well as the flooding and creation of
continental shelves
A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
and
drowned river valleys.
The main agents responsible for deposition and
erosion
Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is d ...
along coastlines are
waves
Waves most often refers to:
* Waves, oscillations accompanied by a transfer of energy that travel through space or mass.
* Wind waves, surface waves that occur on the free surface of bodies of water.
Waves may also refer to:
Music
*Waves (ban ...
,
tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables ...
s and
currents. The formation of coasts also depends on the
nature of the rocks they are made of – the harder the rocks the less likely they are to erode, so variations in rock hardness result in coastlines with different shapes.
Tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables ...
s often determine the range over which
sediment
Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand ...
is deposited or eroded. Areas with high tidal ranges allow waves to reach farther up the shore, and areas with lower tidal ranges produce deposition at a smaller elevation interval. The tidal range is influenced by the size and shape of the coastline. Tides do not typically cause erosion by themselves; however,
tidal bore
Tidal is the adjectival form of tide.
Tidal may also refer to:
* ''Tidal'' (album), a 1996 album by Fiona Apple
* Tidal (king), a king involved in the Battle of the Vale of Siddim
* TidalCycles, a live coding environment for music
* Tidal (se ...
s can erode as the waves surge up river
estuaries
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
from the ocean.
Waves erode coastline as they break on shore releasing their energy; the larger the wave the more energy it releases and the more sediment it moves. Sediment deposited by waves comes from eroded cliff faces and is moved along the coastline by the waves. Sediment deposited by rivers is the dominant influence on the amount of sediment located on a coastline.
The
sedimentologist
Sedimentology encompasses the study of modern sediments such as sand, silt, and clay, and the processes that result in their formation (erosion and weathering), transport, deposition and diagenesis. Sedimentologists apply their understanding of mo ...
Francis Shepard classified coasts as ''primary'' or ''secondary''.
[Shepard FP (1937]
Revised "Classification of Marine Shorelines"
''The Journal of Geology,'' 45(6): 602–624.
* Primary coasts are shaped by non-marine processes, by changes in the land form. If a coast is in much the same condition as it was when sea level was stabilised after the last ice age, it is called a primary coast.
"Primary coasts are created by erosion (the wearing away of soil or rock), deposition (the buildup of sediment or sand) or tectonic activity (changes in the structure of the rock and soil because of earthquakes). Many of these coastlines were formed as the sea level rose during the last 18,000 years, submerging river and glacial valleys to form bays and fjords."
''Office of Naval Research
The Office of Naval Research (ONR) is an organization within the United States Department of the Navy responsible for the science and technology programs of the U.S. Navy and Marine Corps. Established by Congress in 1946, its mission is to pl ...
''. Retrieved 17 April 2011. An example of a primary coast is a
river delta
A river delta is a landform shaped like a triangle, created by deposition of sediment that is carried by a river and enters slower-moving or stagnant water. This occurs where a river enters an ocean, sea, estuary, lake, reservoir, or (more rare ...
, which forms when a river deposits soil and other material as it enters the sea.
* Secondary coasts are produced by marine processes, such as the action of the sea or by creatures that live in it. Secondary coastlines include
sea cliffs,
barrier island
Barrier islands are coastal landforms and a type of dune system that are exceptionally flat or lumpy areas of sand that form by wave and tidal action parallel to the mainland coast. They usually occur in chains, consisting of anything from a ...
s,
mud flats,
coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s,
mangrove swamp
Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangroves cannot withstand fre ...
s and
salt marsh
A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is domin ...
es.
Continental coastlines usually have a
continental shelf
A continental shelf is a portion of a continent that is submerged under an area of relatively shallow water, known as a shelf sea. Much of these shelves were exposed by drops in sea level during glacial periods. The shelf surrounding an island ...
, a shelf of relatively shallow water, less than 200 metres deep, which extends 68 km on average beyond the coast. Worldwide, continental shelves occupy a total area of about 24 million km
2 (9 million sq mi), 8% of the ocean's total area and nearly 5% of the world's total area.
[World]
The World Factbook, CIA. Retrieved 26 February 2010. Since the continental shelf is usually less than 200 metres deep, it follows that coastal habitats are generally
photic
The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological pro ...
, situated in the sunlit
epipelagic zone. This means the conditions for
photosynthetic
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
processes so important for
primary production
In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through ...
, are available to coastal marine habitats. Because land is nearby, there are large discharges of nutrient rich
land runoff into coastal waters. Further, periodic
upwelling
Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted ...
s from the deep ocean can provide cool and nutrient rich currents along the edge of the continental shelf.
As a result, coastal marine life is the most abundant in the world. It is found in
tidal pools,
fjord
In physical geography, a fjord or fiord () is a long, narrow inlet with steep sides or cliffs, created by a glacier. Fjords exist on the coasts of Alaska, Antarctica, British Columbia, Chile, Denmark, Germany, Greenland, the Faroe Islands, Icel ...
s and
estuaries
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
, near sandy shores and rocky coastlines, around
coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s and on or above the continental shelf.
Coastal fish include small forage fish as well as the larger
predator fish
Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, pike/ muskellunge, walleye, perch and salmon. Some omnivorous fish, such as the red-belli ...
that feed on them.
Forage fish
Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the f ...
thrive in inshore waters where high
productivity
Productivity is the efficiency of production of goods or services expressed by some measure. Measurements of productivity are often expressed as a ratio of an aggregate output to a single input or an aggregate input used in a production proces ...
results from
upwelling
Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted ...
and shoreline run off of nutrients. Some are partial residents that spawn in streams, estuaries and bays, but most complete their life cycle in the zone.
[Moyle and Cech, 2004, page 572] There can also be a
mutualism between species that occupy adjacent marine habitats. For example,
fringing reef
A fringing reef is one of the three main types of coral reef. It is distinguished from the other main types, barrier reefs and atolls, in that it has either an entirely shallow backreef zone (lagoon) or none at all. If a fringing reef grows dire ...
s just below
low tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables c ...
level have a mutually beneficial relationship with
mangrove
A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...
forests at high tide level and
sea grass
Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the or ...
meadows in between: the reefs protect the mangroves and seagrass from strong currents and waves that would damage them or
erode
Erode () is a city in the Indian state of Tamil Nadu. Erode is the seventh largest urban agglomeration in the state, after Chennai, Coimbatore, Madurai, Tiruchirapalli, Tiruppur and Salem. It is also the administrative headquarters of the ...
the sediments in which they are rooted, while the mangroves and seagrass protect the coral from large influxes of
silt
Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel ...
, fresh water and
pollutants
A pollutant or novel entity is a substance or energy introduced into the environment that has undesired effects, or adversely affects the usefulness of a resource. These can be both naturally forming (i.e. minerals or extracted compounds like o ...
. This additional level of variety in the environment is beneficial to many types of coral reef animals, which for example may feed in the sea grass and use the reefs for protection or breeding.
Coastal habitats are the most visible marine habitats, but they are not the only important marine habitats. Coastlines run for 380,000 kilometres, and the total volume of the ocean is 1,370 million cu km. This means that for each metre of coast, there is 3.6 cu km of ocean space available somewhere for marine habitats.
Intertidal
Intertidal zone
The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
s, those areas close to shore, are constantly being exposed and covered by the ocean's
tides
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables c ...
. A huge array of life lives within this zone.
Shore habitats range from the upper intertidal zones to the area where land vegetation takes prominence. It can be underwater anywhere from daily to very infrequently. Many species here are scavengers, living off of sea life that is washed up on the shore. Many land animals also make much use of the shore and intertidal habitats. A subgroup of organisms in this habitat bores and grinds exposed rock through the process of
bioerosion
Bioerosion describes the breakdown of hard ocean substrates – and less often terrestrial substrates – by living organisms. Marine bioerosion can be caused by mollusks, polychaete worms, phoronids, sponges, crustaceans, echinoids, and ...
.
Sandy shores
Sandy shores, also called
beach
A beach is a landform alongside a body of water which consists of loose particles. The particles composing a beach are typically made from rock, such as sand, gravel, shingle, pebbles, etc., or biological sources, such as mollusc s ...
es, are coastal shorelines where
sand
Sand is a granular material composed of finely divided mineral particles. Sand has various compositions but is defined by its grain size. Sand grains are smaller than gravel and coarser than silt. Sand can also refer to a textural class ...
accumulates. Waves and currents shift the sand, continually building and eroding the shoreline.
Longshore currents flow parallel to the beaches, making waves break obliquely on the sand. These currents transport large amounts of sand along coasts, forming
spits,
barrier island
Barrier islands are coastal landforms and a type of dune system that are exceptionally flat or lumpy areas of sand that form by wave and tidal action parallel to the mainland coast. They usually occur in chains, consisting of anything from a ...
s and
tombolo
A tombolo is a sandy or shingle isthmus. A tombolo, from the Italian ', meaning 'pillow' or 'cushion', and sometimes translated incorrectly as ''ayre'' (an ayre is a shingle beach of any kind), is a deposition landform by which an island becom ...
s. Longshore currents also commonly create
offshore bars, which give beaches some stability by reducing erosion.
[Habitats: Beaches - Characteristics](_blank)
''Office of Naval Research
The Office of Naval Research (ONR) is an organization within the United States Department of the Navy responsible for the science and technology programs of the U.S. Navy and Marine Corps. Established by Congress in 1946, its mission is to pl ...
''. Retrieved 17 April 2011.
Sandy shores are full of life, The grains of sand host
diatom
A diatom ( Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising se ...
s,
bacteria
Bacteria (; singular: bacterium) are ubiquitous, mostly free-living organisms often consisting of one biological cell. They constitute a large domain of prokaryotic microorganisms. Typically a few micrometres in length, bacteria were am ...
and other
microscopic
The microscopic scale () is the scale of objects and events smaller than those that can easily be seen by the naked eye, requiring a lens or microscope to see them clearly. In physics, the microscopic scale is sometimes regarded as the scale be ...
creatures. Some fish and turtles return to certain beaches and
spawn
Spawn or spawning may refer to:
* Spawn (biology), the eggs and sperm of aquatic animals
Arts, entertainment, and media
* Spawn (character), a fictional character in the comic series of the same name and in the associated franchise
** '' Spawn: A ...
eggs in the sand. Birds habitat beaches, like
gull
Gulls, or colloquially seagulls, are seabirds of the family Laridae in the suborder Lari. They are most closely related to the terns and skimmers and only distantly related to auks, and even more distantly to waders. Until the 21st century ...
s,
loon
Loons ( North American English) or divers ( British / Irish English) are a group of aquatic birds found in much of North America and northern Eurasia. All living species of loons are members of the genus ''Gavia'', family Gaviidae and order ...
s,
sandpiper
Sandpipers are a large family, Scolopacidae, of waders. They include many species called sandpipers, as well as those called by names such as curlew and snipe. The majority of these species eat small invertebrates picked out of the mud or soil ...
s,
terns and
pelican
Pelicans (genus ''Pelecanus'') are a genus of large water birds that make up the family Pelecanidae. They are characterized by a long beak and a large throat pouch used for catching prey and draining water from the scooped-up contents before ...
s.
Aquatic mammal
Aquatic and semiaquatic mammals are a diverse group of mammals that dwell partly or entirely in bodies of water. They include the various marine mammals who dwell in oceans, as well as various freshwater species, such as the European otter. They ...
s, such sea lions, recuperate on them.
Clam
Clam is a common name for several kinds of bivalve molluscs. The word is often applied only to those that are edible and live as infauna, spending most of their lives halfway buried in the sand of the seafloor or riverbeds. Clams have two shel ...
s,
periwinkles,
crab
Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all th ...
s,
shrimp
Shrimp are crustaceans (a form of shellfish) with elongated bodies and a primarily swimming mode of locomotion – most commonly Caridea and Dendrobranchiata of the decapod order, although some crustaceans outside of this order are ref ...
,
starfish
Starfish or sea stars are star-shaped echinoderms belonging to the class Asteroidea (). Common usage frequently finds these names being also applied to ophiuroids, which are correctly referred to as brittle stars or basket stars. Starfish a ...
and
sea urchin
Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) o ...
s are found on most beaches.
[Habitats: Beaches - Animal & Plant Life](_blank)
''Office of Naval Research
The Office of Naval Research (ONR) is an organization within the United States Department of the Navy responsible for the science and technology programs of the U.S. Navy and Marine Corps. Established by Congress in 1946, its mission is to pl ...
''. Retrieved 17 April 2011.
Sand is a
sediment
Sediment is a naturally occurring material that is broken down by processes of weathering and erosion, and is subsequently transported by the action of wind, water, or ice or by the force of gravity acting on the particles. For example, sand ...
made from small grains or
particles with diameters between about 60 µm and 2 mm. Mud (see
mudflats below) is a sediment made from particles finer than sand. This small particle size means that mud particles tend to stick together, whereas sand particles do not. Mud is not easily shifted by waves and currents, and when it dries out, cakes into a solid. By contrast, sand is easily shifted by waves and currents, and when sand dries out it can be blown in the wind, accumulating into shifting
sand dune
A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, f ...
s. Beyond the high tide mark, if the beach is low-lying, the wind can form rolling hills of sand dunes. Small dunes shift and reshape under the influence of the wind while larger dunes stabilise the sand with vegetation.
Ocean processes grade loose sediments to
particle sizes other than sand, such as
gravel
Gravel is a loose aggregation of rock fragments. Gravel occurs naturally throughout the world as a result of sedimentary and erosive geologic processes; it is also produced in large quantities commercially as crushed stone.
Gravel is classifi ...
or
cobbles. Waves breaking on a beach can leave a
berm
A berm is a level space, shelf, or raised barrier (usually made of compacted soil) separating areas in a vertical way, especially partway up a long slope. It can serve as a terrace road, track, path, a fortification line, a border/ separation ...
, which is a raised ridge of coarser pebbles or sand, at the high tide mark.
Shingle beach
A shingle beach (also referred to as rocky beach or pebble beach) is a beach which is armoured with pebbles or small- to medium-sized cobbles (as opposed to fine sand). Typically, the stone composition may grade from characteristic sizes rangi ...
es are made of particles larger than sand, such as cobbles, or small stones. These beaches make poor habitats. Little life survives because the stones are churned and pounded together by waves and currents.
Rocky shores
The relative solidity of
rocky shores seems to give them a permanence compared to the shifting nature of sandy shores. This apparent stability is not real over even quite short geological time scales, but it is real enough over the short life of an organism. In contrast to sandy shores, plants and animals can anchor themselves to the rocks.
[Tour of Rocky Shoreline Habitats](_blank)
''Marietta College''. Retrieved 17 April 2011.
Competition can develop for the rocky spaces. For example,
barnacle
A barnacle is a type of arthropod constituting the subclass Cirripedia in the subphylum Crustacea, and is hence related to crabs and lobsters. Barnacles are exclusively marine, and tend to live in shallow and tidal waters, typically in erosiv ...
s can compete successfully on open intertidal rock faces to the point where the rock surface is covered with them. Barnacles resist desiccation and grip well to exposed rock faces. However, in the crevices of the same rocks, the inhabitants are different. Here
mussel
Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which ...
s can be the successful species, secured to the rock with their
byssal threads.
Rocky and sandy coasts are vulnerable because humans find them attractive and want to live near them. An increasing proportion of the humans live by the coast, putting pressure on coastal habitats.
Mudflats
Mudflat
Mudflats or mud flats, also known as tidal flats or, in Ireland, slob or slobs, are coastal wetlands that form in intertidal areas where sediments have been deposited by tides or rivers. A global analysis published in 2019 suggested that tidal f ...
s are coastal wetlands that form when mud is deposited by tides or rivers. They are found in sheltered areas such as
bays
A bay is a recessed, coastal body of water that directly connects to a larger main body of water, such as an ocean, a lake, or another bay. A large bay is usually called a gulf, sea, sound, or bight. A cove is a small, circular bay with a na ...
,
bayous
In usage in the Southern United States, a bayou () is a body of water typically found in a flat, low-lying area. It may refer to an extremely slow-moving stream, river (often with a poorly defined shoreline), marshy lake, wetland, or creek. They ...
,
lagoons, and
estuaries
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environme ...
. Mudflats may be viewed
geologic
Geology () is a branch of natural science concerned with Earth and other astronomical objects, the features or rocks of which it is composed, and the processes by which they change over time. Modern geology significantly overlaps all other E ...
ally as exposed layers of
bay mud
Bay mud consists of thick deposits of soft, unconsolidated silty clay, which is saturated with water; these soil layers are situated at the bottom of certain estuaries, which are normally in temperate regions that have experienced cyclical glac ...
, resulting from deposition of
estuarine
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environmen ...
silt
Silt is granular material of a size between sand and clay and composed mostly of broken grains of quartz. Silt may occur as a soil (often mixed with sand or clay) or as sediment mixed in suspension with water. Silt usually has a floury feel ...
s,
clay
Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4).
Clays develop plasticity when wet, due to a molecular film of water surrounding the clay pa ...
s and marine animal
detritus
In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts comm ...
. Most of the sediment within a mudflat is within the
intertidal zone
The intertidal zone, also known as the foreshore, is the area above water level at low tide and underwater at high tide (in other words, the area within the tidal range). This area can include several types of habitats with various species ...
, and thus the flat is submerged and exposed approximately twice daily.
Mangrove forests and salt marshes
Mangrove swamp
Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangroves cannot withstand fre ...
s and
salt marsh
A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is domin ...
es form important coastal habitats in tropical and temperate areas respectively.
Mangroves are species of shrubs and medium size trees that grow in
saline coastal sediment habitats in the
tropics
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also refer ...
and
subtropics
The subtropical zones or subtropics are geographical and climate zones to the north and south of the tropics. Geographically part of the temperate zones of both hemispheres, they cover the middle latitudes from to approximately 35° nort ...
– mainly between
latitude
In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north ...
s ° N and ° S. The saline conditions tolerated by various species range from
brackish water
Brackish water, sometimes termed brack water, is water occurring in a natural environment that has more salinity than freshwater, but not as much as seawater. It may result from mixing seawater (salt water) and fresh water together, as in estuari ...
, through pure
seawater
Seawater, or salt water, is water from a sea or ocean. On average, seawater in the world's oceans has a salinity of about 3.5% (35 g/L, 35 ppt, 600 mM). This means that every kilogram (roughly one liter by volume) of seawater has appro ...
(30 to 40
ppt), to water concentrated by
evaporation
Evaporation is a type of vaporization that occurs on the surface of a liquid as it changes into the gas phase. High concentration of the evaporating substance in the surrounding gas significantly slows down evaporation, such as when h ...
to over twice the salinity of ocean seawater (up to 90 ppt).
There are many mangrove species, not all closely related. The term "mangrove" is used generally to cover all of these species, and it can be used narrowly to cover just mangrove trees of the genus ''
Rhizophora
''Rhizophora'' is a genus of tropical mangrove trees, sometimes collectively called true mangroves. The most notable species is the red mangrove (''Rhizophora mangle'') but some other species and a few natural hybrids are known. ''Rhizophora'' ...
''.
Mangroves form a distinct characteristic saline
woodland
A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (see ...
or
shrubland
Shrubland, scrubland, scrub, brush, or bush is a plant community characterized by vegetation dominated by shrubs, often also including grasses, herbs, and geophytes. Shrubland may either occur naturally or be the result of human activity. It ...
habitat, called a ''mangrove swamp'' or ''mangrove forest''.
[Hogarth, Peter J. (1999) ''The Biology of Mangroves'' Oxford University Press, Oxford, England, What page? ] Mangrove swamps are found in
depositional coastal environments, where fine sediments (often with high organic content) collect in areas protected from high-energy wave action. Mangroves dominate three quarters of tropical coastlines.
[
]
Estuaries
An estuary
An estuary is a partially enclosed coastal body of brackish water with one or more rivers or streams flowing into it, and with a free connection to the open sea. Estuaries form a transition zone between river environments and maritime environm ...
is a partly enclosed coastal
The coast, also known as the coastline or seashore, is defined as the area where land meets the ocean, or as a line that forms the boundary between the land and the coastline. The Earth has around of coastline. Coasts are important zones in ...
body of water
Water (chemical formula ) is an inorganic, transparent, tasteless, odorless, and nearly colorless chemical substance, which is the main constituent of Earth's hydrosphere and the fluids of all known living organisms (in which it acts as ...
with one or more river
A river is a natural flowing watercourse, usually freshwater, flowing towards an ocean, sea, lake or another river. In some cases, a river flows into the ground and becomes dry at the end of its course without reaching another body of ...
s or stream
A stream is a continuous body of surface water flowing within the bed and banks of a channel. Depending on its location or certain characteristics, a stream may be referred to by a variety of local or regional names. Long large streams ...
s flowing into it, and with a free connection to the open sea
The sea, connected as the world ocean or simply the ocean, is the body of salty water that covers approximately 71% of the Earth's surface. The word sea is also used to denote second-order sections of the sea, such as the Mediterranean Sea, ...
.[Pritchard, D. W. (1967) ''What is an estuary: physical viewpoint''. p. 3–5 ''in:'' G. H. Lauf (ed.) ''Estuaries'', A.A.A.S. Publ. No. 83, Washington, D.C.] Estuaries form a transition zone between river environments and ocean environments and are subject to both marine influences, such as tides, waves, and the influx of saline water; and riverine influences, such as flows of fresh water and sediment. The inflow of both seawater and freshwater provide high levels of nutrients in both the water column and sediment, making estuaries among the most productive natural habitats in the world.[McLusky, D.S. and Elliott, M. (2004) "The Estuarine Ecosystem: ecology, threats and management." New York: Oxford University Press Inc. ]
Most estuaries were formed by the flooding of river-eroded or glacially scoured valleys when sea level began to rise about 10,000-12,000 years ago.[Wolanski, E. (2007) "Estuarine Ecohydrology." Amsterdam, The Netherlands: Elsevier. ] They are amongst the most heavily populated areas throughout the world, with about 60% of the world's population living along estuaries and the coast. As a result, estuaries are suffering degradation by many factors, including sedimentation from soil erosion from deforestation; overgrazing and other poor farming practices; overfishing; drainage and filling of wetlands; eutrophication due to excessive nutrients from sewage and animal wastes; pollutants including heavy metals, PCBs, radionuclides and hydrocarbons from sewage inputs; and diking or damming for flood control or water diversion.
Estuaries provide habitats for a large number of organisms and support very high productivity. Estuaries provide habitats for salmon
Salmon () is the common name for several commercially important species of euryhaline ray-finned fish from the family Salmonidae, which are native to tributaries of the North Atlantic (genus '' Salmo'') and North Pacific (genus '' Onco ...
and sea trout
Sea trout is the common name usually applied to anadromous (sea-run) forms of brown trout (''Salmo trutta''), and is often referred to as ''Salmo trutta'' morpha ''trutta''. Other names for anadromous brown trout are sewin (Wales), peel or peal ...
nurseries, as well as migratory bird populations. Two of the main characteristics of estuarine life are the variability in salinity
Salinity () is the saltiness or amount of salt (chemistry), salt dissolved in a body of water, called saline water (see also soil salinity). It is usually measured in g/L or g/kg (grams of salt per liter/kilogram of water; the latter is dimensio ...
and sedimentation
Sedimentation is the deposition of sediments. It takes place when particles in suspension settle out of the fluid in which they are entrained and come to rest against a barrier. This is due to their motion through the fluid in response to the ...
. Many species of fish
Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
and invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
s have various methods to control or conform to the shifts in salt concentrations and are termed osmoconformer
Osmoconformers are marine organisms that maintain an internal environment which is isotonic to their external environment. This means that the osmotic pressure of the organism's cells is equal to the osmotic pressure of their surrounding environm ...
s and osmoregulators. Many animals also burrow
An Eastern chipmunk at the entrance of its burrow
A burrow is a hole or tunnel excavated into the ground by an animal to construct a space suitable for habitation or temporary refuge, or as a byproduct of locomotion. Burrows provide a form of s ...
to avoid predation
Predation is a biological interaction where one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
and to live in the more stable sedimental environment. However, large numbers of bacteria are found within the sediment which have a very high oxygen demand. This reduces the levels of oxygen within the sediment often resulting in partially anoxic
The term anoxia means a total depletion in the level of oxygen, an extreme form of hypoxia or "low oxygen". The terms anoxia and hypoxia are used in various contexts:
* Anoxic waters, sea water, fresh water or groundwater that are depleted of diss ...
conditions, which can be further exacerbated by limited water flux. Phytoplankton
Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'.
...
are key primary producers in estuaries. They move with the water bodies and can be flushed in and out with the tide
Tides are the rise and fall of sea levels caused by the combined effects of the gravitational forces exerted by the Moon (and to a much lesser extent, the Sun) and are also caused by the Earth and Moon orbiting one another.
Tide tables ...
s. Their productivity is largely dependent on the turbidity
Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of water quality.
Fluids ...
of the water. The main phytoplankton present are diatoms
A diatom (Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising sev ...
and dinoflagellates
The dinoflagellates ( Greek δῖνος ''dinos'' "whirling" and Latin ''flagellum'' "whip, scourge") are a monophyletic group of single-celled eukaryotes constituting the phylum Dinoflagellata and are usually considered algae. Dinoflagellates ...
which are abundant in the sediment.
Kelp forests
Kelp forest
Kelp forests are underwater areas with a high density of kelp, which covers a large part of the world's coastlines. Smaller areas of anchored kelp are called kelp beds. They are recognized as one of the most productive and dynamic ecosystems on Ea ...
s are underwater areas with a high density of kelp
Kelps are large brown algae seaweeds that make up the order Laminariales. There are about 30 different genera. Despite its appearance, kelp is not a plant - it is a heterokont, a completely unrelated group of organisms.
Kelp grows in "underwa ...
. They form some of the most productive and dynamic ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s on Earth.[Mann, K.H. 1973. Seaweeds: their productivity and strategy for growth. Science 182: 975-981.] Smaller areas of anchored kelp are called ''kelp beds''. Kelp forests occur worldwide throughout temperate
In geography, the temperate climates of Earth occur in the middle latitudes (23.5° to 66.5° N/S of Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ranges throughout ...
and polar
Polar may refer to:
Geography
Polar may refer to:
* Geographical pole, either of two fixed points on the surface of a rotating body or planet, at 90 degrees from the equator, based on the axis around which a body rotates
*Polar climate, the cli ...
coastal oceans.
Kelp forests provide a unique three-dimensional habitat for marine organisms and are a source for understanding many ecological processes. Over the last century, they have been the focus of extensive research, particularly in trophic
Trophic, from Ancient Greek τροφικός (''trophikos'') "pertaining to food or nourishment", may refer to:
* Trophic cascade
* Trophic coherence
* Trophic egg
* Trophic function
* Trophic hormone
* Trophic level index
* Trophic level
...
ecology, and continue to provoke important ideas that are relevant beyond this unique ecosystem. For example, kelp forests can influence coastal oceanographic
Oceanography (), also known as oceanology and ocean science, is the scientific study of the oceans. It is an Earth science, which covers a wide range of topics, including ecosystem dynamics; ocean currents, waves, and geophysical fluid dynamic ...
patterns and provide many ecosystem services
Ecosystem services are the many and varied benefits to humans provided by the natural environment and healthy ecosystems. Such ecosystems include, for example, agroecosystems, forest ecosystem, grassland ecosystems, and aquatic ecosystems. ...
.[Steneck, R.S., M.H. Graham, B.J. Bourque, D. Corbett, J.M. Erlandson, J.A. Estes and M.J. Tegner. 2002. Kelp forest ecosystems: biodiversity, stability, resilience and future. Environmental Conservation 29: 436-459.]
However, humans have contributed to kelp forest degradation
Forest degradation is a process in which the biological wealth of a forest area is permanently diminished by some factor or by a combination of factors. "This does not involve a reduction of the forest area, but rather a quality decrease in its c ...
. Of particular concern are the effects of overfishing
Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing fish stock), resulting in t ...
nearshore ecosystems, which can release herbivores
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
from their normal population regulation and result in the over-grazing of kelp and other algae.[Sala, E., C.F. Bourdouresque and M. Harmelin-Vivien. 1998. Fishing, trophic cascades, and the structure of algal assemblages: evaluation of an old but untested paradigm. Oikos 82: 425-439.] This can rapidly result in transitions to barren landscapes where relatively few species persist.[Dayton, P.K. 1985a. Ecology of kelp communities. Annual Review of Ecology and Systematics 16: 215-245.]
Frequently considered an ecosystem engineer
An ecosystem engineer is any species that creates, significantly modifies, maintains or destroys a habitat. These organisms can have a large impact on species richness and landscape-level heterogeneity of an area. As a result, ecosystem engin ...
, kelp provides a physical substrate and habitat for kelp forest communities. In algae (Kingdom: Protista
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the e ...
), the body of an individual organism is known as a thallus
Thallus (plural: thalli), from Latinized Greek (), meaning "a green shoot" or "twig", is the vegetative tissue of some organisms in diverse groups such as algae, fungi, some liverworts, lichens, and the Myxogastria. Many of these organisms ...
rather than as a plant (Kingdom: Plantae
Plants are predominantly photosynthetic eukaryotes of the kingdom Plantae. Historically, the plant kingdom encompassed all living things that were not animals, and included algae and fungi; however, all current definitions of Plantae exclude ...
). The morphological structure of a kelp thallus is defined by three basic structural units:
:* The holdfast is a root-like mass that anchors the thallus to the sea floor, though unlike true roots it is not responsible for absorbing and delivering nutrients to the rest of the thallus;
:* The stipe is analogous to a plant stalk, extending vertically from the holdfast and providing a support framework for other morphological features;
:* The fronds are leaf- or blade-like attachments extending from the stipe, sometimes along its full length, and are the sites of nutrient uptake and photosynthetic activity.
In addition, many kelp species have pneumatocyst
In phycology, a pneumatocyst is a floating structure that contains gas found on brown seaweed. A seaweed's thallus may have more than one. They provide buoyancy
Buoyancy (), or upthrust, is an upward force exerted by a fluid that oppose ...
s, or gas-filled bladders, usually located at the base of fronds near the stipe. These structures provide the necessary buoyancy for kelp to maintain an upright position in the water column.
The environmental factors necessary for kelp to survive include hard substrate (usually rock), high nutrients (e.g., nitrogen, phosphorus), and light (minimum annual irradiance In radiometry, irradiance is the radiant flux ''received'' by a ''surface'' per unit area. The SI unit of irradiance is the watt per square metre (W⋅m−2). The CGS unit erg per square centimetre per second (erg⋅cm−2⋅s−1) is often used ...
dose > 50 E m−2[Druehl, L.D. 1981. The distribution of Laminariales in the North Pacific with reference to environmental influences. Proceedings of the International Congress on Systematic Evolution and Biology 2: 248-256.]). Especially productive kelp forests tend to be associated with areas of significant oceanographic upwelling
Upwelling is an physical oceanography, oceanographic phenomenon that involves wind-driven motion of dense, cooler, and usually nutrient-rich water from deep water towards the ocean surface. It replaces the warmer and usually nutrient-depleted ...
, a process that delivers cool nutrient-rich water from depth to the ocean's mixed surface layer. Water flow and turbulence facilitate nutrient assimilation across kelp fronds throughout the water column. Water clarity affects the depth to which sufficient light can be transmitted. In ideal conditions, giant kelp (''Macrocystis spp.'') can grow as much as 30-60 centimetres vertically per day. Some species such as ''Nereocystis'' are annual
Annual may refer to:
*Annual publication, periodical publications appearing regularly once per year
** Yearbook
** Literary annual
*Annual plant
*Annual report
*Annual giving
*Annual, Morocco, a settlement in northeastern Morocco
*Annuals (band), ...
while others like ''Eisenia'' are perennial
A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
, living for more than 20 years. In perennial kelp forests, maximum growth rates occur during upwelling months (typically spring and summer) and die-backs correspond to reduced nutrient availability, shorter photoperiods and increased storm frequency.
Seagrass meadows
Seagrass
Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families ( Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the ...
es are flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s from one of four plant families which grow in marine environments. They are called ''seagrasses'' because the leaves are long and narrow and are very often green, and because the plants often grow in large meadows which look like grassland. Since seagrasses photosynthesize and are submerged, they must grow submerged in the photic zone
The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological pro ...
, where there is enough sunlight. For this reason, most occur in shallow and sheltered coastal waters anchored in sand or mud bottoms.
Seagrasses form extensive beds or meadows, which can be either monospecific (made up of one species) or multispecific (where more than one species co-exist). Seagrass beds make highly diverse and productive ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s. They are home to phyla Phyla, the plural of ''phylum'', may refer to:
* Phylum, a biological taxon between Kingdom and Class
* by analogy, in linguistics, a large division of possibly related languages, or a major language family which is not subordinate to another
Phy ...
such as juvenile and adult fish, epiphytic and free-living macroalgae
Seaweed, or macroalgae, refers to thousands of species of macroscopic, multicellular, marine algae. The term includes some types of '' Rhodophyta'' (red), ''Phaeophyta'' (brown) and ''Chlorophyta'' (green) macroalgae. Seaweed species such as ...
and microalgae
Microalgae or microphytes are microscopic algae invisible to the naked eye. They are phytoplankton typically found in freshwater and marine systems, living in both the water column and sediment. They are unicellular species which exist indiv ...
, mollusk
Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000 extant species of molluscs are recognized. The number of fossil species is e ...
s, bristle worms, and nematode
The nematodes ( or grc-gre, Νηματώδη; la, Nematoda) or roundworms constitute the phylum Nematoda (also called Nemathelminthes), with plant- parasitic nematodes also known as eelworms. They are a diverse animal phylum inhabiting a bro ...
s. Few species were originally considered to feed directly on seagrass leaves
A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
(partly because of their low nutritional content), but scientific review
A review is an evaluation of a publication, product, service, or company or a critical take on current affairs in literature, politics or culture. In addition to a critical evaluation, the review's author may assign the work a rating to indi ...
s and improved working methods have shown that seagrass herbivory
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthpart ...
is a highly important link in the food chain, with hundreds of species feeding on seagrasses worldwide, including green turtle
The green sea turtle (''Chelonia mydas''), also known as the green turtle, black (sea) turtle or Pacific green turtle, is a species of large sea turtle of the family Cheloniidae. It is the only species in the genus ''Chelonia''. Its range exten ...
s, dugong
The dugong (; ''Dugong dugon'') is a marine mammal. It is one of four living species of the order Sirenia, which also includes three species of manatees. It is the only living representative of the once-diverse family Dugongidae; its closest m ...
s, manatee
Manatees (family Trichechidae, genus ''Trichechus'') are large, fully aquatic, mostly herbivorous marine mammals sometimes known as sea cows. There are three accepted living species of Trichechidae, representing three of the four living speci ...
s, fish
Fish are Aquatic animal, aquatic, craniate, gill-bearing animals that lack Limb (anatomy), limbs with Digit (anatomy), digits. Included in this definition are the living hagfish, lampreys, and Chondrichthyes, cartilaginous and bony fish as we ...
, geese, swan
Swans are birds of the family Anatidae within the genus ''Cygnus''. The swans' closest relatives include the geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe Cygnini. Som ...
s, sea urchin
Sea urchins () are spiny, globular echinoderms in the class Echinoidea. About 950 species of sea urchin live on the seabed of every ocean and inhabit every depth zone from the intertidal seashore down to . The spherical, hard shells (tests) o ...
s and crab
Crabs are decapod crustaceans of the infraorder Brachyura, which typically have a very short projecting "tail" (abdomen) ( el, βραχύς , translit=brachys = short, / = tail), usually hidden entirely under the thorax. They live in all th ...
s.
Seagrasses are ecosystem engineer
An ecosystem engineer is any species that creates, significantly modifies, maintains or destroys a habitat. These organisms can have a large impact on species richness and landscape-level heterogeneity of an area. As a result, ecosystem engin ...
s in the sense that they partly create their own habitat. The leaves slow down water-currents increasing sedimentation
Sedimentation is the deposition of sediments. It takes place when particles in suspension settle out of the fluid in which they are entrained and come to rest against a barrier. This is due to their motion through the fluid in response to the ...
, and the seagrass root
In vascular plants, the roots are the organs of a plant that are modified to provide anchorage for the plant and take in water and nutrients into the plant body, which allows plants to grow taller and faster. They are most often below the su ...
s and rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...
s stabilize the seabed. Their importance to associated species is mainly due to provision of shelter (through their three-dimensional structure in the water column), and due to their extraordinarily high rate of primary production
In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through ...
. As a result, seagrasses provide coastal zones with ecosystem services
Ecosystem services are the many and varied benefits to humans provided by the natural environment and healthy ecosystems. Such ecosystems include, for example, agroecosystems, forest ecosystem, grassland ecosystems, and aquatic ecosystems. ...
, such as fishing grounds, wave
In physics, mathematics, and related fields, a wave is a propagating dynamic disturbance (change from equilibrium) of one or more quantities. Waves can be periodic, in which case those quantities oscillate repeatedly about an equilibrium (re ...
protection, oxygen
Oxygen is the chemical element with the symbol O and atomic number 8. It is a member of the chalcogen group in the periodic table, a highly reactive nonmetal, and an oxidizing agent that readily forms oxides with most elements ...
production and protection against coastal erosion
Erosion is the action of surface processes (such as water flow or wind) that removes soil, rock, or dissolved material from one location on the Earth's crust, and then transports it to another location where it is deposited. Erosion is d ...
. Seagrass meadows account for 15% of the ocean's total carbon storage.
Reefs
A reef is a ridge or shoal
In oceanography, geomorphology, and geoscience, a shoal is a natural submerged ridge, bank, or bar that consists of, or is covered by, sand or other unconsolidated material and rises from the bed of a body of water to near the surface. It ...
of rock, coral
Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...
or similar relatively stable material, lying beneath the surface of a natural body of water. Many reefs result from natural, abiotic processes but there are also reefs such as the coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s of tropical waters formed by biotic processes dominated by corals and coralline algae
Coralline algae are red algae in the order Corallinales. They are characterized by a thallus that is hard because of calcareous deposits contained within the cell walls. The colors of these algae are most typically pink, or some other shade of re ...
. Artificial reef
An artificial reef is a human-created underwater structure, typically built to promote marine life in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of trawling nets, or improve surfing.
Many ...
s such as shipwrecks and other anthropogenic underwater structures may occur intentionally or as the result of an accident, and sometimes have a designed role in enhancing the physical complexity of featureless sand bottoms, thereby attracting a more diverse assemblage of organisms. Reefs are often quite near to the surface, but not all definitions require this. Fringing reefs, the most common type of reef, are found close to shorelines and surrounding islands.
Rocky reefs
Coral reefs
Coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s comprise some of the densest and most diverse habitats in the world. The best-known types of reefs are tropical
The tropics are the regions of Earth surrounding the Equator. They are defined in latitude by the Tropic of Cancer in the Northern Hemisphere at N and the Tropic of Capricorn in
the Southern Hemisphere at S. The tropics are also referred to ...
coral reef
A coral reef is an underwater ecosystem characterized by reef-building corals. Reefs are formed of Colony (biology), colonies of coral polyp (zoology), polyps held together by calcium carbonate. Most coral reefs are built from stony corals, wh ...
s which exist in most tropical waters; however, coral reefs can also exist in cold water. Reefs are built up by coral
Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and se ...
s and other calcium
Calcium is a chemical element with the symbol Ca and atomic number 20. As an alkaline earth metal, calcium is a reactive metal that forms a dark oxide-nitride layer when exposed to air. Its physical and chemical properties are most similar ...
-depositing animals, usually on top of a rocky outcrop on the ocean floor. Reefs can also grow on other surfaces, which has made it possible to create artificial reef
An artificial reef is a human-created underwater structure, typically built to promote marine life in areas with a generally featureless bottom, to control erosion, block ship passage, block the use of trawling nets, or improve surfing.
Many ...
s. Coral reefs also support a huge community of life, including the corals themselves, their symbiotic zooxanthellae
Zooxanthellae is a colloquial term for single-celled dinoflagellates that are able to live in symbiosis with diverse marine invertebrates including demosponges, corals, jellyfish, and nudibranchs. Most known zooxanthellae are in the genus ''S ...
, tropical fish and many other organisms.
Much attention in marine biology is focused on coral reefs and the El Niño
El Niño (; ; ) is the warm phase of the El Niño–Southern Oscillation (ENSO) and is associated with a band of warm ocean water that develops in the central and east-central equatorial Pacific (approximately between the International Date ...
weather phenomenon. In 1998, coral reefs experienced the most severe mass bleaching events on record, when vast expanses of reefs across the world died because sea surface temperatures rose well above normal. Some reefs are recovering, but scientists say that between 50% and 70% of the world's coral reefs are now endangered and predict that global warming
In common usage, climate change describes global warming—the ongoing increase in global average temperature—and its effects on Earth's climate system. Climate change in a broader sense also includes previous long-term changes to ...
could exacerbate this trend.
Surface waters
Surface microlayer
The surface microlayer of the ocean serves as the transitional area between the atmosphere and the ocean. It covers around 70% of the Earth's surface as it covers most of the ocean waters on the planet. The microlayer is known for its unique biological and chemical properties which give it a small ecosystem of its own and serves as a distinct habitat from the deeper ocean waters.
The surface microlayer is not in fact entirely aqueous like the rest of the ocean, but is closer to a kind of hydrated gel composed of concentrated nutrients forming a biological film over the water it covers. This film is rich in microbes which mediate the interactions between the sun, the atmosphere, and the waters below.
Although thin, the surface microlayer is critical for life beneath it. Because of the environment rich in microbes and nutrients, larvae of fish and other aquatic animals are often laid in the microlayer to incubate. The plankton in the microlayer are distinctly adapted to withstand high levels of radiation, and serve as buffers to prevent this potentially harmful radiation from reaching the deeper water. Environmental changes such as aerosols or dust storms can cause these surface plankton to become overproductive, leading to blooms.
Because of the unique properties of the microlayer, pollutants often accumulate within and use it to reach other parts of the ocean. Hydrophobic compounds, such as petroleum
Petroleum, also known as crude oil, or simply oil, is a naturally occurring yellowish-black liquid mixture of mainly hydrocarbons, and is found in geological formations. The name ''petroleum'' covers both naturally occurring unprocessed crud ...
, flame retardants, and heavy metals, have a particular affinity for the surface microlayer. Recently, the abundance of aerosols and microplastics
Microplastics are fragments of any type of plastic less than in length, according to the U.S. National Oceanic and Atmospheric Administration (NOAA) and the European Chemicals Agency. They cause pollution by entering natural ecosystems from a v ...
has also had an impact on the SML and their accumulation has led to many problems, such as animal ingestion of these compounds leading to widespread disruption of balance and spread of these compounds among marine communities.
The surface microlayer is also critical to gas exchange between the atmosphere and the ocean. Because the microlayer is filled with microbes, it is widely theorized that it plays a critical role in gas exchange and uptake of nutrients, but relatively little data on this has been collected. The central feature of the microlayer is the temperature, as it is an indicator of how pollutants and human activity affects the ocean.
Epipelagic zone
The surface waters are sunlit. The waters down to about 200 metres are said to be in the epipelagic zone. Enough sunlight enters the epipelagic zone to allow photosynthesis
Photosynthesis is a process used by plants and other organisms to convert light energy into chemical energy that, through cellular respiration, can later be released to fuel the organism's activities. Some of this chemical energy is stored in ...
by phytoplankton
Phytoplankton () are the autotrophic (self-feeding) components of the plankton community and a key part of ocean and freshwater ecosystems. The name comes from the Greek words (), meaning 'plant', and (), meaning 'wanderer' or 'drifter'.
...
. The epipelagic zone is usually low in nutrients. This partially because the organic debris produced in the zone, such as excrement and dead animals, sink to the depths and are lost to the upper zone. Photosynthesis can happen only if both sunlight and nutrients are present.[Ocean ecology: sunlit surface waters]
''WWF''. Retrieved 17 May 2011.
In some places, like at the edge of continental shelves, nutrients can upwell from the ocean depth, or land runoff can be distributed by storms and ocean currents. In these areas, given that both sunlight and nutrients are now present, phytoplankton can rapidly establish itself, multiplying so fast that the water turns green from the chlorophyll, resulting in an algal bloom. These nutrient rich surface waters are among the most biologically productive in the world, supporting billions of tonnes of biomass
Biomass is plant-based material used as a fuel for heat or electricity production. It can be in the form of wood, wood residues, energy crops, agricultural residues, and waste from industry, farms, and households. Some people use the terms bio ...
.
"Phytoplankton are eaten by zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
- small animals which, like phytoplankton, drift in the ocean currents. The most abundant zooplankton species are copepod
Copepods (; meaning "oar-feet") are a group of small crustaceans found in nearly every freshwater and saltwater habitat. Some species are planktonic (inhabiting sea waters), some are benthic (living on the ocean floor), a number of species have ...
s and krill
Krill are small crustaceans of the order Euphausiacea, and are found in all the world's oceans. The name "krill" comes from the Norwegian word ', meaning "small fry of fish", which is also often attributed to species of fish.
Krill are consi ...
: tiny crustacean
Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
s that are the most numerous animals on Earth. Other types of zooplankton include jelly fish and the larvae
A larva (; plural larvae ) is a distinct juvenile form many animals undergo before metamorphosis into adults. Animals with indirect development such as insects, amphibians, or cnidarians typically have a larval phase of their life cycle.
T ...
of fish, marine worms, starfish
Starfish or sea stars are star-shaped echinoderms belonging to the class Asteroidea (). Common usage frequently finds these names being also applied to ophiuroids, which are correctly referred to as brittle stars or basket stars. Starfish a ...
, and other marine organisms". In turn, the zooplankton are eaten by filter-feeding animals, including some seabird
Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s, small forage fish
Forage fish, also called prey fish or bait fish, are small pelagic fish which are preyed on by larger predators for food. Predators include other larger fish, seabirds and marine mammals. Typical ocean forage fish feed near the base of the f ...
like herrings and sardines, whale shark
The whale shark (''Rhincodon typus'') is a slow-moving, filter feeder, filter-feeding carpet shark and the largest known Extant taxon, extant fish species. The largest confirmed individual had a length of .McClain CR, Balk MA, Benfield MC, Bran ...
s, manta ray
Manta rays are large rays belonging to the genus ''Mobula'' (formerly its own genus ''Manta''). The larger species, '' M. birostris'', reaches in width, while the smaller, '' M. alfredi'', reaches . Both have triangular pectoral fins, horn-s ...
s, and the largest animal in the world, the blue whale
The blue whale (''Balaenoptera musculus'') is a marine mammal and a baleen whale. Reaching a maximum confirmed length of and weighing up to , it is the largest animal known to have ever existed. The blue whale's long and slender body can ...
. Yet again, moving up the foodchain, the small forage fish are in turn eaten by larger predators, such as tuna, marlin, sharks, large squid, seabirds, dolphins, and toothed whale
The toothed whales (also called odontocetes, systematic name Odontoceti) are a parvorder of cetaceans that includes dolphins, porpoises, and all other whales possessing teeth, such as the beaked whales and sperm whales. Seventy-three species of t ...
s.
Open ocean
The open ocean is relatively unproductive because of a lack of nutrients, yet because it is so vast, it has more overall primary production
In ecology, primary production is the synthesis of organic compounds from atmospheric or aqueous carbon dioxide. It principally occurs through the process of photosynthesis, which uses light as its source of energy, but it also occurs through ...
than any other marine habitat. Only about 10 percent of marine species live in the open ocean. But among them are the largest and fastest of all marine animals, as well as the animals that dive the deepest and migrate the longest. In the depths lurk animal that, to our eyes, appear hugely alien.
Deep sea
The deep sea
The deep sea is broadly defined as the ocean depth where light begins to fade, at an approximate depth of 200 metres (656 feet) or the point of transition from continental shelves to continental slopes. Conditions within the deep sea are a combin ...
starts at the aphotic zone, the point where sunlight loses most of its energy in the water. Many life forms that live at these depths have the ability to create their own light a unique evolution known as bio-luminescence.
In the deep ocean, the waters extend far below the epipelagic zone, and support very different types of pelagic life forms adapted to living in these deeper zones.[Moyle and Cech, 2004, page 585]
Much of the aphotic zone's energy is supplied by the open ocean in the form of detritus
In biology, detritus () is dead particulate organic material, as distinguished from dissolved organic material. Detritus typically includes the bodies or fragments of bodies of dead organisms, and fecal material. Detritus typically hosts comm ...
. In deep water, marine snow is a continuous shower of mostly organic detritus falling from the upper layers of the water column. Its origin lies in activities within the productive photic zone
The photic zone, euphotic zone, epipelagic zone, or sunlight zone is the uppermost layer of a body of water that receives sunlight, allowing phytoplankton to perform photosynthesis. It undergoes a series of physical, chemical, and biological pro ...
. Marine snow includes dead or dying plankton
Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...
, protist
A protist () is any eukaryotic organism (that is, an organism whose cells contain a cell nucleus) that is not an animal, plant, or fungus. While it is likely that protists share a common ancestor (the last eukaryotic common ancestor), the e ...
s (diatom
A diatom ( Neo-Latin ''diatoma''), "a cutting through, a severance", from el, διάτομος, diátomos, "cut in half, divided equally" from el, διατέμνω, diatémno, "to cut in twain". is any member of a large group comprising se ...
s), fecal matter, sand, soot and other inorganic dust. The "snowflakes" grow over time and may reach several centimetres in diameter, travelling for weeks before reaching the ocean floor. However, most organic components of marine snow are consumed by microbe
A microorganism, or microbe,, ''mikros'', "small") and ''organism'' from the el, ὀργανισμός, ''organismós'', "organism"). It is usually written as a single word but is sometimes hyphenated (''micro-organism''), especially in olde ...
s, zooplankton
Zooplankton are the animal component of the planktonic community ("zoo" comes from the Greek word for ''animal''). Plankton are aquatic organisms that are unable to swim effectively against currents, and consequently drift or are carried along by ...
and other filter-feeding animals within the first 1,000 metres of their journey, that is, within the epipelagic zone. In this way marine snow may be considered the foundation of deep-sea mesopelagic and benthic
The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning " ...
ecosystem
An ecosystem (or ecological system) consists of all the organisms and the physical environment with which they interact. These biotic and abiotic components are linked together through nutrient cycles and energy flows. Energy enters the syst ...
s: As sunlight cannot reach them, deep-sea organisms rely heavily on marine snow as an energy source.
Some deep-sea pelagic groups, such as the lanternfish, ridgehead, marine hatchetfish, and lightfish families are sometimes termed ''pseudoceanic'' because, rather than having an even distribution in open water, they occur in significantly higher abundances around structural oases, notably seamount
A seamount is a large geologic landform that rises from the ocean floor that does not reach to the water's surface (sea level), and thus is not an island, islet or cliff-rock. Seamounts are typically formed from extinct volcanoes that rise a ...
s and over continental slope
A continental margin is the outer edge of continental crust abutting oceanic crust under coastal waters. It is one of the three major zones of the ocean floor, the other two being deep-ocean basins and mid-ocean ridges. The continental margin ...
s. The phenomenon is explained by the likewise abundance of prey species which are also attracted to the structures.
The fish in the different pelagic and deep water benthic zones are physically structured, and behave in ways, that differ markedly from each other. Groups of coexisting species within each zone all seem to operate in similar ways, such as the small mesopelagic vertically migrating plankton-feeders, the bathypelagic anglerfish
The anglerfish are fish of the teleost order Lophiiformes (). They are bony fish named for their characteristic mode of predation, in which a modified luminescent fin ray (the esca or illicium) acts as a lure for other fish. The luminescence ...
es, and the deep water benthic rattail
Grenadiers or rattails are generally large, brown to black gadiform marine fish of the subfamily Macrourinae, the largest subfamily of the family Macrouridae. Found at great depths from the Arctic to Antarctic, members of this subfamily are amon ...
s. "[Moyle and Cech, 2004, p. 591]
Ray finned species, with spiny fins, are rare among deep sea fishes, which suggests that deep sea fish are ancient and so well adapted to their environment that invasions by more modern fishes have been unsuccessful.[Haedrich RL (1996]
"Deep-water fishes: evolution and adaptation in the earth's largest living spaces"
''Journal of Fish Biology'' 49(sA):40-53. The few ray fins that do exist are mainly in the Beryciformes and Lampriformes, which are also ancient forms. Most deep sea pelagic fishes belong to their own orders, suggesting a long evolution in deep sea environments. In contrast, deep water benthic species, are in orders that include many related shallow water fishes.[Moyle and Cech, 2004, page 586]
The umbrella mouth gulper is a deep sea eel with an enormous loosely hinged mouth. It can open its mouth wide enough to swallow a fish much larger than itself, and then expand its stomach to accommodate its catch.
Sea floor
Vents and seeps
Hydrothermal vent
A hydrothermal vent is a fissure on the seabed from which geothermally heated water discharges. They are commonly found near volcanically active places, areas where tectonic plates are moving apart at mid-ocean ridges, ocean basins, and hotspo ...
s along the mid-ocean ridge
A mid-ocean ridge (MOR) is a seafloor mountain system formed by plate tectonics. It typically has a depth of about and rises about above the deepest portion of an ocean basin. This feature is where seafloor spreading takes place along a div ...
spreading centers act as oases
In ecology, an oasis (; ) is a fertile area of a desert or semi-desert environment'ksar''with its surrounding feeding source, the palm grove, within a relational and circulatory nomadic system.”
The location of oases has been of critical im ...
, as do their opposites, cold seeps. Such places support unique marine biomes and many new marine microorganisms and other lifeforms have been discovered at these locations.
Trenches
The deepest recorded oceanic trench
Oceanic trenches are prominent long, narrow topographic depressions of the ocean floor. They are typically wide and below the level of the surrounding oceanic floor, but can be thousands of kilometers in length. There are about of oceanic tren ...
es measure to date is the Mariana Trench
The Mariana Trench is an oceanic trench located in the western Pacific Ocean, about east of the Mariana Islands; it is the deepest oceanic trench on Earth. It is crescent-shaped and measures about in length and in width. The maximum known ...
, near the Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
, in the Pacific Ocean
The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the conti ...
at 10,924 m (35,838 ft). At such depths, water pressure is extreme and there is no sunlight, but some life still exists. A white flatfish
A flatfish is a member of the ray-finned demersal fish order Pleuronectiformes, also called the Heterosomata, sometimes classified as a suborder of Perciformes. In many species, both eyes lie on one side of the head, one or the other migrating ...
, a shrimp and a jellyfish were seen by the American crew of the bathyscaphe
A bathyscaphe ( or ) is a free-diving self-propelled deep-sea submersible, consisting of a crew cabin similar to a bathysphere, but suspended below a float rather than from a surface cable, as in the classic bathysphere design.
The float is ...
''Trieste
Trieste ( , ; sl, Trst ; german: Triest ) is a city and seaport in northeastern Italy. It is the capital city, and largest city, of the autonomous region of Friuli Venezia Giulia, one of two autonomous regions which are not subdivided into pr ...
'' when it dove to the bottom in 1960.
Seamounts
Marine life also flourishes around seamount
A seamount is a large geologic landform that rises from the ocean floor that does not reach to the water's surface (sea level), and thus is not an island, islet or cliff-rock. Seamounts are typically formed from extinct volcanoes that rise a ...
s that rise from the depths, where fish and other sea life congregate to spawn and feed.
Anthropogenic impacts
Mudflats are typically important regions for wildlife, supporting a large population, although levels of biodiversity are not particularly high. They are of particular importance to migratory birds as well as crabs, shrimp, and shellfish. These areas along the coast act as a nursery for these animals by providing an area for reproduction and feeding. However, this can pose as an issue due to the high trafficking of the birds migrating for nesting, then leaving to return to their seasonal homes. Whatever pollutants the birds take in while breeding are brought back with them to their next location, thus polluting that area as well. In the United Kingdom
The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Europe, off the north-western coast of the continental mainland. It comprises England, Scotland, Wales and ...
mudflats have been classified as a Biodiversity Action Plan priority habitat. European countries such as France have also found it beneficial to use the Marine Influence Index (MII) to be able to monitor the responses to pollution the local plant and animal species may have as well as monitor any type of deviation from the natural patterns displayed previously.
Although many parts of the seafloor have yet to be explored, researchers have found that parts of it have been greatly affected by human activity. Bottom trawling, microplastic pollution, and industrial metals have slowly changed and altered the composition of the sea floor. Bottom trawling refers to a commercial deep sea fishing technique in which the equipment drags across the sea floor. This has had an adverse effect on the seafloor as it changes the surface structure and composition. In addition, microplastic
Microplastics are fragments of any type of plastic less than in length, according to the U.S. National Oceanic and Atmospheric Administration (NOAA) and the European Chemicals Agency. They cause pollution by entering natural ecosystems from a v ...
pollution has become an increasing problem to the seafloor as plastics and other debris are found in many of the sediments. Due to the build up of litter, the habitats and environments of organisms on the seafloor are being impacted and changed. This includes industrial facilities dumping new metals and minerals, such as cadmium
Cadmium is a chemical element with the Symbol (chemistry), symbol Cd and atomic number 48. This soft, silvery-white metal is chemically similar to the two other stable metals in group 12 element, group 12, zinc and mercury (element), mercury. Li ...
, onto the seafloor that change the chemical composition of the water and poison the inhabitants.
File:Microplastic.jpg, Microplastics found in sediments on the seafloor
File:Eurythenes plasticus (10.11646-zootaxa.4748.1.9) Figure 3.jpg, 640 µm microplastic found in the deep sea amphipod ''Eurythenes plasticus''
There are also negative anthropogenic impacts on deep sea habitats, including trash pollution and chemical pollution. Plastic pollution
Plastic pollution is the accumulation of plastic objects and particles (e.g. plastic bottles, bags and microbeads) in the Earth's environment that adversely affects humans, wildlife and their habitat. Plastics that act as pollutants are cate ...
in particular, is one of the greatest forms of uncontrolled human activity that is visible in our oceans today. Researchers in the Northwestern south China Sea recorded large plastic-dominated litter piles in submarine canyons. These durable plastics can diffuse into smaller organisms and are then inadvertently consumed by humans in the food we eat and water we drink. Another threat to organisms lurking in the deep ocean is ghost fishing, and bycatch
Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
. Ghost fishing is the term that refers to any abandoned fishing gear in the ocean that continues to entangle and trap marine organisms. Gill nets for example, have been recorded tangled around deep sea corals and continue ghost fishing for extended periods of time.
Gallery
Mushroom corals.jpg, Deepsea mushroom coral
The Fungiidae () are a family of Cnidaria, commonly known as mushroom corals or plate corals. The family contains thirteen extant genera. They range from solitary corals to colonial species. Some genera such as ''Cycloseris'' and '' Fungia'' ar ...
s growing on the chimney of an Explorer Ridge hydrothermal vent
Gandalfus yunohana by OpenCage.jpg, Deep sea crab found on hydrothermal vents in the Philippine Sea
The Philippine Sea is a marginal sea of the Western Pacific Ocean east of the Philippine archipelago (hence the name), the largest in the world, occupying an estimated surface area of . The Philippine Sea Plate forms the floor of the sea. I ...
Deepsea anemone.jpg, Deep sea anemone on Blake Ridge
The Blake Plateau lies in the western Atlantic Ocean off the southeastern United States coasts of North Carolina, South Carolina, Georgia and Florida. The Blake Plateau lies between the North American continental shelf and the deep ocean basin e ...
Wreckdiver 300.jpg, Submerged wrecks create artificial reef habitat
Greenspotrock 100.jpg, Versatile rockfish can be found living in almost any habitat from rocky bottoms to sand and mud, and from vertical faces to horizontal plains.
Otter in kelp field.jpg, Marine life within the kelp forest and rocky reef habitat
Monterey Bay STS090-705-5.jpg, Monterey Bay, the largest marine sanctuary in the United States, is home to the world's largest group of marine research institutions
Orcas off Santa Rosa island.jpg, The killer whale
The orca or killer whale (''Orcinus orca'') is a toothed whale belonging to the oceanic dolphin family, of which it is the largest member. It is the only extant species in the genus ''Orcinus'' and is recognizable by its black-and-white pat ...
, apex predator of the ocean, cruises a huge range of different marine habitats
Starr 080603-5639 Boerhavia repens.jpg, Laysan albatross chick in a contemporary modified habitat, surrounded by human marine debris
Marine debris, also known as marine litter, is human-created waste that has deliberately or accidentally been released in a sea or ocean. Floating oceanic debris tends to accumulate at the center of gyres and on coastlines, frequently washing ...
File:Explorer Ridge Zooarium chimney.jpg, Zooarium chimney provides a habitat for vent biota
See also
* Effects of climate change on oceans
Among the effects of climate change on oceans are: an increase in sea surface temperature as well as ocean temperatures at greater depths, more frequent marine heatwaves, a reduction in pH value, a rise in sea level from ocean warming and ic ...
* Future of Marine Animal Populations
* Maritime forest
* Seashore wildlife Seashore wildlife habitats exist from the Tropics to the Arctic and Antarctic. Seashores and beaches provide varied habitats in different parts of the world, and even within the same beach. Phytoplankton is at the bottom of some food chains, whil ...
* Underwater habitat
References
Sources
* Kritzer JP and Sale PF (2006
''Marine metapopulations''
Academic Press. .
* Moyle, PB and Cech, JJ (2004) ''Fishes, An Introduction to Ichthyology.'' 5th Ed, Benjamin Cummings.
* Nybakken JW and Bertness MD (2005
''Marine biology: an ecological approach''
Sixth edition, Pearson/Benjamin Cummings. – organized by habitat, not classification
*
External links
Missouri Botanical Garden: Marine ecosystems
{{DEFAULTSORT:Marine Biology
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Biological oceanography
Fisheries science
Oceanographical terminology
Oceanography