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Teeth are common to most
vertebrates Vertebrates () comprise all animal taxa within the subphylum Vertebrata () ( chordates with backbones), including all mammals, birds, reptiles, amphibians, and fish. Vertebrates represent the overwhelming majority of the phylum Chordata, ...
, but mammalian teeth are distinctive in having a variety of shapes and functions. This feature first arose among early
therapsid Therapsida is a major group of eupelycosaurian synapsids that includes mammals, their ancestors and relatives. Many of the traits today seen as unique to mammals had their origin within early therapsids, including limbs that were oriented more ...
s during the
Permian The Permian ( ) is a geologic period and stratigraphic system which spans 47 million years from the end of the Carboniferous Period million years ago (Mya), to the beginning of the Triassic Period 251.9 Mya. It is the last period of the Paleoz ...
, and has continued to the present day. All therapsid groups with the exception of the mammals are now extinct, but each of these groups possessed different tooth patterns, which aids with the classification of fossils. Most extant mammals including humans are
diphyodont A diphyodont is any animal with two ss of teeth, initially the ''deciduous'' set and consecutively the '' permanent'' set. Most mammals are diphyodonts—as to chew their food they need a strong, durable and complete set of teeth. Diphyodonts contr ...
s, i.e. they have an early set of
deciduous teeth Deciduous teeth or primary teeth, also informally known as baby teeth, milk teeth, or temporary teeth,Illustrated Dental Embryology, Histology, and Anatomy, Bath-Balogh and Fehrenbach, Elsevier, 2011, page 255 are the first set of teeth in the ...
and a later set of permanent or "adult" teeth. Notable exceptions are elephants, kangaroos, and manatees, all of which are
polyphyodont A polyphyodont is any animal whose teeth are continually replaced. In contrast, diphyodonts are characterized by having only two successive sets of teeth. Polyphyodonts include most toothed fishes, many reptiles such as crocodiles and geckos, an ...
s, i.e. having teeth that are continuously being replaced. Mammal teeth include
incisor Incisors (from Latin ''incidere'', "to cut") are the front teeth present in most mammals. They are located in the premaxilla above and on the mandible below. Humans have a total of eight (two on each side, top and bottom). Opossums have 18, wher ...
s,
canines Canine may refer to: Zoology and anatomy * a dog-like Canid animal in the subfamily Caninae ** ''Canis'', a genus including dogs, wolves, coyotes, and jackals ** Dog, the domestic dog * Canine tooth, in mammalian oral anatomy People with the sur ...
,
premolar The premolars, also called premolar teeth, or bicuspids, are transitional teeth located between the canine and molar teeth. In humans, there are two premolars per quadrant in the permanent set of teeth, making eight premolars total in the mou ...
s, and
molars The molars or molar teeth are large, flat teeth at the back of the mouth. They are more developed in mammals. They are used primarily to grind food during chewing. The name ''molar'' derives from Latin, ''molaris dens'', meaning "millstone to ...
, not all of which are present in all mammals. Various evolutionary modifications have occurred, such as the lack of canines in
Glires Glires (, Latin ''glīrēs'' 'dormice') is a clade (sometimes ranked as a grandorder) consisting of rodents and lagomorphs (rabbits, hares, and pikas). The hypothesis that these form a monophyletic group has been long debated based on morphologi ...
, the development of
tusk Tusks are elongated, continuously growing front teeth that protrude well beyond the mouth of certain mammal species. They are most commonly canine teeth, as with pigs and walruses, or, in the case of elephants, elongated incisors. Tusks share co ...
s from either incisors (elephants) or canines (pigs and walruses), the adaptation of molars into flesh-shearing
carnassial Carnassials are paired upper and lower teeth modified in such a way as to allow enlarged and often self-sharpening edges to pass by each other in a shearing manner. This adaptation is found in carnivorans, where the carnassials are the modified ...
s in
Carnivora Carnivora is a monophyletic order of placental mammals consisting of the most recent common ancestor of all cat-like and dog-like animals, and all descendants of that ancestor. Members of this group are formally referred to as carnivorans, ...
, and others.


Diversity

The extant mammalian infraclasses each have a set
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolog ...
; the
Eutheria Eutheria (; from Greek , 'good, right' and , 'beast'; ) is the clade consisting of all therian mammals that are more closely related to placentals than to marsupials. Eutherians are distinguished from noneutherians by various phenotypic tra ...
(placental mammals) commonly have three pairs of molars and four premolars per jaw, whereas the
Metatheria Metatheria is a mammalian clade that includes all mammals more closely related to marsupials than to placentals. First proposed by Thomas Henry Huxley in 1880, it is a more inclusive group than the marsupials; it contains all marsupials as well ...
(marsupials) generally have four pairs of molars and between three or two premolars. For example, the
tiger quoll The tiger quoll (''Dasyurus maculatus''), also known as the spotted-tail quoll, the spotted quoll, the spotted-tail dasyure, native cat or the tiger cat, is a carnivorous marsupial of the quoll genus ''Dasyurus'' native to Australia. With males ...
(''Dasyurus maculatus'') is a
dasyurid The Dasyuridae are a family of marsupials native to Australia and New Guinea, including 71 extant species divided into 17 genera. Many are small and mouse-like or shrew-like, giving some of them the name marsupial mice or marsupial shrews, but th ...
marsupial Marsupials are any members of the mammalian infraclass Marsupialia. All extant marsupials are endemic to Australasia, Wallacea and the Americas. A distinctive characteristic common to most of these species is that the young are carried in a ...
native to Australia. The quoll possesses four upper incisors and three lower incisors per left and right-hand side = 14 two upper premolars Mand two lower premolars per side M = 8 and four upper and four lower molars per side = 16 giving the animal a complement of thirty-eight teeth. The tiger quoll's
dental formula Dentition pertains to the development of teeth and their arrangement in the mouth. In particular, it is the characteristic arrangement, kind, and number of teeth in a given species at a given age. That is, the number, type, and morpho-physiolog ...
is therefore as follows: .


Rabbits

Rabbit Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit specie ...
s and other
lagomorph The lagomorphs are the members of the taxonomic order Lagomorpha, of which there are two living families: the Leporidae (hares and rabbits) and the Ochotonidae (pikas). The name of the order is derived from the Ancient Greek ''lagos'' (λαγ ...
s usually shed their deciduous teeth before (or very shortly after) their birth, and are usually born with their permanent teeth. The teeth of rabbits complement their diet, which consist of a wide range of vegetation. Since many of the foods are abrasive enough to cause attrition, rabbit teeth grow continuously throughout life. Rabbits have a dental formula of . There are no canines. Three to four millimeters of tooth is worn away by incisors every week, whereas the posterior teeth require a month to wear away the same amount. Anatomy of rabbit teeth The incisors and cheek teeth of rabbits are called aradicular hypsodont teeth. Aradicular teeth never form a true root with an apex, and
hypsodont Hypsodont is a pattern of dentition with high-crowned teeth and enamel extending past the gum line, providing extra material for wear and tear. Some examples of animals with hypsodont dentition are cows and horses; all animals that feed on gritty ...
teeth have a high crown to root ratio (providing more room for wear and tear). This is sometimes referred to as an elodont dentition, meaning ever-growing. These teeth grow or erupt continuously. The growth or eruption is held in balance by dental abrasion from chewing a diet high in fiber.


Rodents

Like those of rabbits,
rodent Rodents (from Latin , 'to gnaw') are mammals of the order Rodentia (), which are characterized by a single pair of continuously growing incisors in each of the upper and lower jaws. About 40% of all mammal species are rodents. They are nat ...
s' incisors grow continuously throughout their lives, and are both aradicular and hypsodont. Unlike humans whose ameloblasts die after
tooth development Tooth development or odontogenesis is the complex process by which teeth form from embryonic cells, grow, and erupt into the mouth. For human teeth to have a healthy oral environment, all parts of the tooth must develop during appropriate stage ...
, rodents continually produce enamel and must wear down their teeth by gnawing on various materials. These teeth are used for cutting wood, biting through the skin of fruit, or for defense. The teeth have enamel on the outside and exposed
dentin Dentin () (American English) or dentine ( or ) (British English) ( la, substantia eburnea) is a calcified tissue of the body and, along with enamel, cementum, and pulp, is one of the four major components of teeth. It is usually covered by ena ...
on the inside, so they self-sharpen during
gnawing {{Short pages monitor