Muskellunge
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The muskellunge (''Esox masquinongy''), often shortened to muskie, musky, ski, or lunge, is a
species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...
of large
freshwater Fresh water or freshwater is any naturally occurring liquid or frozen water containing low concentrations of dissolved salts and other total dissolved solids. The term excludes seawater and brackish water, but it does include non-salty mi ...
predatory fish Predatory fish are hypercarnivorous fish that actively prey upon other fish or aquatic animals, with examples including shark, billfish, barracuda, alligator gar, tuna, dolphinfish, walleye, perch and salmon. Some omnivorous fish, such as t ...
native to
North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...
. It is the largest member of the pike family,
Esocidae Esocidae is a family of ray-finned fish in the order Salmoniformes, which contains pike, pickerel, and mudminnows. While the family traditionally only contained the genus ''Esox'', recent genetic and paleontological research have recovered ''Novu ...
.


Origin of name

The name "muskellunge" originates from the
Ojibwe The Ojibwe (; Ojibwe writing systems#Ojibwe syllabics, syll.: ᐅᒋᐺ; plural: ''Ojibweg'' ᐅᒋᐺᒃ) are an Anishinaabe people whose homeland (''Ojibwewaki'' ᐅᒋᐺᐘᑭ) covers much of the Great Lakes region and the Great Plains, n ...
words ''maashkinoozhe'' meaning "great fish", ''mji-gnoozhe,'' ''maskinoše'', or ''mashkinonge,'' meaning "bad pike", "big pike", or "ugly pike" respectively. The Algonquin word ''maskinunga'' is borrowed into the
Canadian French Canadian French (, ) is the French language as it is spoken in Canada. It includes multiple varieties, the most prominent of which is Québécois (Quebec French). Formerly ''Canadian French'' referred solely to Quebec French and the closely re ...
words ''masquinongé'' or ''maskinongé''. In English, before settling on the common name "muskellunge", there were at least 94 common names applied to this species, including but not limited to: ''muskelunge'', ''muscallonge'', ''muskallonge'', ''milliganong'', ''maskinonge'', ''maskalonge'', ''mascalonge'', ''maskalung'', ''muskinunge'' and ''masquenongez''.


Description

Muskellunge closely resemble other esocids such as the
northern pike The northern pike (''Esox lucius'') is a species of carnivorous fish of the genus ''Esox'' (pikes). They are commonly found in brackish water, moderately salty and fresh waters of the Northern Hemisphere (''i.e.'' holarctic in distribution). T ...
(''Esox lucius'') and American pickerel (''E. americanus'') in both appearance and behavior. Like the northern pike and other aggressive pikes, the body plan is typical of
ambush predator Ambush predators or sit-and-wait predators are carnivorous animals that capture their prey via stealth, luring or by (typically instinctive) strategies utilizing an element of surprise. Unlike pursuit predators, who chase to capture prey u ...
s with an elongated body, flat head, and dorsal, pelvic, and anal fins set far back on the body. Muskellunge are typically long and weigh , though some have reached up to and almost . Martin Arthur Williamson caught a muskellunge with a weight of in November 2000 on
Georgian Bay The Georgian Bay () is a large bay of Lake Huron, in the Laurentia bioregion. It is located entirely within the borders of Ontario, Canada. The main body of the bay lies east of the Bruce Peninsula and Manitoulin Island. To its northwest is t ...
. The fish are a light silver, brown, or green, with dark vertical stripes on the flank, which may tend to break up into spots. In some cases, markings may be absent altogether, especially in fish from
turbid Turbidity is the cloudiness or haziness of a fluid caused by large numbers of individual particles that are generally invisible to the naked eye, similar to smoke in air. The measurement of turbidity is a key test of both water clarity and wate ...
waters. This is in contrast to northern pike, which have dark bodies with light markings. A reliable method to distinguish the two similar species is by counting the sensory pores on the underside of the
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
. A muskie will have seven or more per side, while the northern pike never has more than six. The lobes of the caudal (tail) fin in muskellunge come to a sharper point, while those of northern pike are more generally rounded. In addition, unlike pike, muskies have no scales on the lower half of their opercula. Anglers seek large muskies as trophies or for sport. In places where muskie are not native, such as in Maine, anglers are encouraged not to release the fish back into the water because of their negative impact on native populations of trout and other smaller fish species.


Habitat

Muskellunge are found in oligotrophic and
mesotrophic lake The Trophic State Index (TSI) is a classification system designed to rate water bodies based on the amount of biological productivity they sustain. Although the term "trophic index" is commonly applied to lakes, any surface water body may be ind ...
s and large
river A river is a natural stream of fresh water that flows on land or inside Subterranean river, caves towards another body of water at a lower elevation, such as an ocean, lake, or another river. A river may run dry before reaching the end of ...
s from northern
Michigan Michigan ( ) is a peninsular U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest, Upper Midwestern United States. It shares water and land boundaries with Minnesota to the northwest, Wisconsin to the west, ...
, northern
Wisconsin Wisconsin ( ) is a U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest of the United States. It borders Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michig ...
, and northern
Minnesota Minnesota ( ) is a U.S. state, state in the Upper Midwestern region of the United States. It is bordered by the Canadian provinces of Manitoba and Ontario to the north and east and by the U.S. states of Wisconsin to the east, Iowa to the so ...
through the
Great Lakes region The Great Lakes region of Northern America is a binational Canadian– American region centered on the Great Lakes that includes the U.S. states of Illinois, Indiana, Michigan, Minnesota, New York, Ohio, Pennsylvania, and Wisconsin and the Ca ...
, Chautauqua Lake in western New York, north into
Canada Canada is a country in North America. Its Provinces and territories of Canada, ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, making it the world's List of coun ...
, throughout most of the
St Lawrence River The St. Lawrence River (, ) is a large international river in the middle latitudes of North America connecting the Great Lakes to the North Atlantic Ocean. Its waters flow in a northeasterly direction from Lake Ontario to the Gulf of St. Lawren ...
drainage, and northward throughout the upper
Mississippi Mississippi ( ) is a U.S. state, state in the Southeastern United States, Southeastern and Deep South regions of the United States. It borders Tennessee to the north, Alabama to the east, the Gulf of Mexico to the south, Louisiana to the s ...
valley, although the species also extends as far south as
Chattanooga Chattanooga ( ) is a city in Hamilton County, Tennessee, United States, and its county seat. It is located along the Tennessee River and borders Georgia (U.S. state), Georgia to the south. With a population of 181,099 in 2020, it is Tennessee ...
in the
Tennessee River The Tennessee River is a long river located in the Southern United States, southeastern United States in the Tennessee Valley. Flowing through the states of Tennessee, Alabama, Mississippi, and Kentucky, it begins at the confluence of Fren ...
valley. Also, a small population is found in the Broad River in
South Carolina South Carolina ( ) is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. It borders North Carolina to the north and northeast, the Atlantic Ocean to the southeast, and Georgia (U.S. state), Georg ...
. Several North Georgia reservoirs also have healthy stocked populations of muskie. They are also found in the Red River drainage of the
Hudson Bay Hudson Bay, sometimes called Hudson's Bay (usually historically), is a large body of Saline water, saltwater in northeastern Canada with a surface area of . It is located north of Ontario, west of Quebec, northeast of Manitoba, and southeast o ...
basin. Muskie were introduced to western Saint John River in the late 1960s and have now spread to many connecting waterways in northern
Maine Maine ( ) is a U.S. state, state in the New England region of the United States, and the northeasternmost state in the Contiguous United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Provinces and ...
. The Pineview Reservoir in Utah is one of three Utah locations where the hybrid Tiger muskellunge is found. They prefer clear waters where they lurk along weed edges, rock outcrops, or other structures to rest. A fish forms two distinct home ranges in summer: a shallow range and a deeper one. The shallow range is generally much smaller than the deeper range due to shallow water heating up. A muskie continually patrols the ranges in search of available food in the appropriate conditions of water temperature.


Diet

Muskies are ambush predators who will swiftly bite their prey and then swallow it head first. Muskellunge are the top predator in any body of water where they occur and they will eat larger prey than most other freshwater fish. They eat all varieties of fish present in their ecosystem (including other muskellunge), along with the occasional
insect Insects (from Latin ') are Hexapoda, hexapod invertebrates of the class (biology), class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body (Insect morphology#Head, head, ...
,
muskrat The muskrat or common muskrat (''Ondatra zibethicus'') is a medium-sized semiaquatic rodent native to North America and an introduced species in parts of Europe, Asia, and South America. The muskrat is found in wetlands over various climates ...
,
rat Rats are various medium-sized, long-tailed rodents. Species of rats are found throughout the order Rodentia, but stereotypical rats are found in the genus ''Rattus''. Other rat genera include '' Neotoma'' (pack rats), '' Bandicota'' (bandicoo ...
,
mouse A mouse (: mice) is a small rodent. Characteristically, mice are known to have a pointed snout, small rounded ears, a body-length scaly tail, and a high breeding rate. The best known mouse species is the common house mouse (''Mus musculus'' ...
,
frog A frog is any member of a diverse and largely semiaquatic group of short-bodied, tailless amphibian vertebrates composing the order (biology), order Anura (coming from the Ancient Greek , literally 'without tail'). Frog species with rough ski ...
, or
duck Duck is the common name for numerous species of waterfowl in the family (biology), family Anatidae. Ducks are generally smaller and shorter-necked than swans and goose, geese, which are members of the same family. Divided among several subfam ...
. They are capable of taking prey up to two-thirds of their body length due to their large stomachs. There have even been reports of large muskellunge attacking small
dogs The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers ...
and even humans, although most of these reports are greatly exaggerated.


Length and weight

As muskellunge grow longer they increase in weight, but the relationship between length and weight is not linear. The relationship between them can be expressed by a power-law equation: :W = cL^b\!\, The exponent b is close to 3.0 for all species, and c is a constant for each species. For muskellunge, b = 3.325, higher than for many common species, and c = . According to the International Game Fish Association (IGFA) the largest muskellunge on record was
caught Caught is a method of dismissing a batsman in cricket. A batsman is out caught if the batsman hits the ball, from a legitimate delivery, with the bat, and the ball is caught by the bowler or a fielder before it hits the ground. If the catch ...
by Cal Johnson in Lac Courte Oreilles (recognized as Lake Courte Oreilles by the association), Hayward,
Wisconsin Wisconsin ( ) is a U.S. state, state in the Great Lakes region, Great Lakes region of the Upper Midwest of the United States. It borders Minnesota to the west, Iowa to the southwest, Illinois to the south, Lake Michigan to the east, Michig ...
,
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
, on July 24, 1949. The
fish A fish (: fish or fishes) is an aquatic animal, aquatic, Anamniotes, anamniotic, gill-bearing vertebrate animal with swimming fish fin, fins and craniate, a hard skull, but lacking limb (anatomy), limbs with digit (anatomy), digits. Fish can ...
weighed and was in length, and in girth.


Behavior

Muskellunge are sometimes
gregarious Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother was ...
, forming small
schools A school is the educational institution (and, in the case of in-person learning, the building) designed to provide learning environments for the teaching of students, usually under the direction of teachers. Most countries have systems of ...
in distinct territories. Muskellunge feeding behavior is directly synchronized with the lunar cycle. During both full and new moons, an increase in feeding activity can be attributed to the increase of moonlight, as it most similarly simulates daytime feeding. They spawn in mid- to late spring, somewhat later than northern pike, over shallow, vegetated areas. A rock or sand bottom is preferred for spawning so the eggs do not sink into the mud and suffocate. The males arrive first and attempt to establish dominance over a territory. Spawning may last from five to 10 days and occurs mainly at night. The eggs are negatively buoyant and slightly adhesive; they adhere to plants and the bottom of the lake. Soon afterward, they are abandoned by the adults. Those
embryo An embryo ( ) is the initial stage of development for a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male sp ...
s which are not eaten by fish, insects, or crayfish hatch within two weeks. The larvae live on
yolk Among animals which produce eggs, the yolk (; also known as the vitellus) is the nutrient-bearing portion of the egg whose primary function is to supply food for the development of the embryo. Some types of egg contain no yolk, for example bec ...
until the mouth is fully developed, when they begin to feed on
copepod Copepods (; meaning 'oar-feet') are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (living in the water column), some are benthos, benthic (living on the sedimen ...
s and other
zooplankton Zooplankton are the heterotrophic component of the planktonic community (the " zoo-" prefix comes from ), having to consume other organisms to thrive. Plankton are aquatic organisms that are unable to swim effectively against currents. Consequent ...
. They soon begin to prey upon fish. Juveniles generally attain a length of by November of their first year.


Predators

Adult muskellunge are
apex predator An apex predator, also known as a top predator or superpredator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the hig ...
s where they occur naturally. Only humans and (rarely) large birds of prey such as
bald eagle The bald eagle (''Haliaeetus leucocephalus'') is a bird of prey found in North America. A sea eagle, it has two known subspecies and forms a species pair with the white-tailed eagle (''Haliaeetus albicilla''), which occupies the same niche ...
s (''Haliaeetus leucocephalus'') pose a threat to an adult. But juveniles are consumed by other muskies, northern pike,
bass Bass or Basses may refer to: Fish * Bass (fish), various saltwater and freshwater species Wood * Bass or basswood, the wood of the tilia americana tree Music * Bass (sound), describing low-frequency sound or one of several instruments in th ...
,
trout Trout (: trout) is a generic common name for numerous species of carnivorous freshwater ray-finned fishes belonging to the genera '' Oncorhynchus'', ''Salmo'' and ''Salvelinus'', all of which are members of the subfamily Salmoninae in the ...
, and occasionally birds of prey. The muskellunge's low reproductive rate and slow growth render populations highly vulnerable to
overfishing Overfishing is the removal of a species of fish (i.e. fishing) from a body of water at a rate greater than that the species can replenish its population naturally (i.e. the overexploitation of the fishery's existing Fish stocks, fish stock), resu ...
. This has prompted some jurisdictions to institute
artificial propagation Plant propagation is the process by which new plants grow from various sources, including seeds, cuttings, and other plant parts. Plant propagation can refer to both man-made and natural processes. Propagation typically occurs as a step in the o ...
programs in an attempt to maintain otherwise unsustainably high rates of angling effort and
habitat destruction Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...
.


Subspecies and hybrids

Though interbreeding with other pike species can complicate the classification of some individuals,
zoologists This is a list of notable zoologists who have published names of new taxa under the International Code of Zoological Nomenclature. A * Abe – Tokiharu Abe (1911–1996) * Abeille de Perrin, Ab. – Elzéar Abeille de Perrin (1843–1910) * ...
usually recognize up to three
subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
of muskellunge. * The Great Lakes Muskellunge or Spotted Muskellunge (''E. m. masquinongy'') is the most common variety in the Great Lakes basin and surrounding area. The spots on the body form oblique rows. * The Chautauqua Muskellunge or Barred Muskellunge (''E. m. ohioensis'') is known from the Ohio River system, Chautauqua Lake,
Lake Ontario Lake Ontario is one of the five Great Lakes of North America. It is bounded on the north, west, and southwest by the Canadian province of Ontario, and on the south and east by the U.S. state of New York (state), New York. The Canada–United Sta ...
, and the St Lawrence River. * The Clear Muskellunge (''E. m. immaculatus'') is most common in the inland lakes of Wisconsin, Minnesota, northwestern Ontario, and southeastern
Manitoba Manitoba is a Provinces and territories of Canada, province of Canada at the Centre of Canada, longitudinal centre of the country. It is Canada's Population of Canada by province and territory, fifth-most populous province, with a population ...
. The tiger muskellunge (''E. masquinongy'' × ''lucius'' or ''E. lucius'' × ''masquinongy'') is a hybrid of the muskie and northern pike. Hybrids are sterile, although females sometimes unsuccessfully engage in spawning motions. Some hybrids are artificially produced and planted for anglers to catch. Tiger muskies grow faster than pure muskies, but do not attain the ultimate size of their pure relatives, as the tiger muskie does not live as long.


Attacks on humans

Although very rare, muskie attacks on humans do occur on occasion.


References


External links

{{DEFAULTSORT:Muskellunge Apex predators Esox Freshwater fish of the United States Fish of Canada Fish of the Eastern United States Fish of the Great Lakes Fish described in 1824 Taxa named by Samuel L. Mitchill Symbols of Wisconsin