''Morus'', a

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

of flowering plants in the family

Moraceae Moraceae is a family of flowering plants comprising about 48 genera and over 1100 species, and is commonly known as the mulberry or fig family. Most are widespread in tropical and subtropical regions, less so in temperate climates; however, their ...

, consists of 19 species of

deciduous In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed Leaf, leaves, usually in the autumn; to the shedding of petals, aft ...

trees commonly known as mulberries, growing wild and under cultivation in many

temperate In geography, the temperate climates of Earth occur in the middle latitudes (approximately 23.5° to 66.5° N/S of the Equator), which span between the tropics and the polar regions of Earth. These zones generally have wider temperature ran ...

world regions. Generally, the genus has 64 subordinate taxa, though the three most common are referred to as white, red, and black, originating from the color of their dormant buds and not necessarily the fruit color (''

Morus alba Morus may refer to: People * Alexander Morus (1616–1670), Franco-Scottish Protestant preacher * Henryk Moruś (1943–2013), Polish serial killer * Huw Morus (1622–1709), Welsh poet * Thomas More or Morus (1478–1535), English philosopher * M ...

'', '' M. rubra'', and '' M. nigra'', respectively), with numerous

cultivar A cultivar is a kind of Horticulture, cultivated plant that people have selected for desired phenotypic trait, traits and which retains those traits when Plant propagation, propagated. Methods used to propagate cultivars include division, root a ...

s and some taxa currently unchecked and awaiting taxonomic scrutiny. ''M. alba'' is native to

South Asia South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...

, but is widely distributed across

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

Southern Africa Southern Africa is the southernmost region of Africa. No definition is agreed upon, but some groupings include the United Nations geoscheme for Africa, United Nations geoscheme, the intergovernmental Southern African Development Community, and ...

South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a considerably smaller portion in the Northern Hemisphere. It can also be described as the southern Subregion#Americas, subregion o ...

, and

North America North America is a continent in the Northern Hemisphere, Northern and Western Hemisphere, Western hemispheres. North America is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South Ameri ...

. ''M. alba'' is also the species most preferred by the

silkworm ''Bombyx mori'', commonly known as the domestic silk moth, is a moth species belonging to the family Bombycidae. It is the closest relative of '' Bombyx mandarina'', the wild silk moth. Silkworms are the larvae of silk moths. The silkworm is of ...

. It is regarded as an

invasive species An invasive species is an introduced species that harms its new environment. Invasive species adversely affect habitats and bioregions, causing ecological, environmental, and/or economic damage. The term can also be used for native spec ...

in Brazil, the United States and some states of Australia. The closely related genus ''

Broussonetia ''Broussonetia'' is a genus of four species (including one hybrid species) of trees in the family Moraceae, native to eastern Asia. These four species have high-quality fiber which consist of more than 90% of cellulose. They are traditionally ap ...

'' is also commonly known as mulberry, notably the

paper mulberry The paper mulberry (''Broussonetia papyrifera'', syn. ''Morus papyrifera'' L.) is a species of flowering plant in the family Moraceae. It is native to Asia, Despite their similar appearance, mulberries are not closely related to

raspberries The raspberry is the edible fruit of several plant species in the genus ''Rubus'' of the Rosaceae, rose family, most of which are in the subgenus ''Rubus#Modern classification, Idaeobatus''. The name also applies to these plants themselves. Ras ...

blackberries BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...

. All three species belong to the

Rosales Rosales (, ) are an order of flowering plants. Peter F. Stevens (2001 onwards). "Rosales". At: Trees At: Angiosperm Phylogeny Website. At: Missouri Botanical Garden Website. (see ''External links'' below) Well-known members of Rosales include: ...

order. But while the mulberry is a tree belonging to the

family (also including the

fig The fig is the edible fruit of ''Ficus carica'', a species of tree or shrub in the flowering plant family Moraceae, native to the Mediterranean region, together with western and southern Asia. It has been cultivated since ancient times and i ...

jackfruit The jackfruit or ''nangka'' (''Artocarpus heterophyllus'') is a species of tree in the Common fig, fig, mulberry, and breadfruit family (Moraceae). The jackfruit is the largest tree fruit, reaching as much as in weight, in length, and in d ...

, and other fruits), raspberries and blackberries are

brambles ''Rubus'' is a large and diverse genus of flowering plants in the rose family, Rosaceae, subfamily Rosoideae, most commonly known as brambles. Fruits of various species are known as raspberries, blackberries, dewberries, and bristleberries. ...

and belong to the

Rosaceae Rosaceae (), the rose family, is a family of flowering plants that includes 4,828 known species in 91 genera. The name is derived from the type genus '' Rosa''. The family includes herbs, shrubs, and trees. Most species are deciduous, but som ...

family.

Description

Mulberries are fast-growing when young, and can grow to tall. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are alternately arranged, simple, and often lobed and serrated on the margin. Lobes are more common on juvenile shoots than on mature trees. The trees can be

monoecious Monoecy (; adj. monoecious ) is a sexual system in seed plants where separate male and female cones or flowers are present on the same plant. It is a monomorphic sexual system comparable with gynomonoecy, andromonoecy and trimonoecy, and contras ...

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

. The mulberry fruit is a multiple, about long. Immature fruits are white, green, or pale yellow. The fruit turns from pink to red while ripening, then dark purple or black, and has a sweet flavor when fully ripe.

Taxonomy

The taxonomy of ''Morus'' is complex and disputed.

Fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s of ''Morus'' appear in the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Netherlands , Terminology of the Low Countries, informally Holland, is a country in Northwestern Europe, with Caribbean Netherlands, overseas territories in the Caribbean. It is the largest of the four constituent countries of the Kingdom of the Nether ...

. Over 150 species names have been published, and although differing sources may cite different selections of accepted names, less than 20 are accepted by the vast majority of botanical authorities. ''Morus'' classification is even further complicated by widespread hybridisation, wherein the hybrids are fertile. The following species are accepted: *''

'' L. – white mulberry (China, Korea, Japan) *''

Morus australis Morus may refer to: People * Alexander Morus (1616–1670), Franco-Scottish Protestant preacher * Henryk Moruś (1943–2013), Polish serial killer * Huw Morus (1622–1709), Welsh poet * Thomas More or Morus (1478–1535), English philosopher * ...

'' Poir. – East and South-East Asia *'' Morus boninensis'' Koidz. *'' Morus cathayana'' Hemsl. – China, Japan, Korea *'' Morus celtidifolia'' Kunth – Texas mulberry (southwestern United States, Mexico, Central America, South America) *''

Morus indica ''Morus indica'' is a species of flowering plant in the Morus (plant), mulberry family Moraceae. As with other mulberries, it is a deciduous tree. ''Morus indica'' is native to the temperate and sub-tropical Himalayan region and is currently cul ...

'' L. – India,

Southeast Asia Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...

*''

Morus koordersiana Morus may refer to: People * Alexander Morus (1616–1670), Franco-Scottish Protestant preacher * Henryk Moruś (1943–2013), Polish serial killer * Huw Morus (1622–1709), Welsh poet * Thomas More or Morus (1478–1535), English philosopher * M ...

'' J.-F.Leroy *''

Morus liboensis Morus may refer to: People * Alexander Morus (1616–1670), Franco-Scottish Protestant preacher * Henryk Moruś (1943–2013), Polish serial killer * Huw Morus (1622–1709), Welsh poet * Thomas More or Morus (1478–1535), English philosopher * M ...

'' S.S.Chang –

Guizhou ) , image_skyline = , image_caption = , image_map = Guizhou in China (+all claims hatched).svg , mapsize = 275px , map_alt = Map showing the location of Guizhou Province , map_caption = Map s ...

Province in China *''

Morus macroura ''Morus macroura'', also known as the king white mulberry, shahtoot mulberry, Tibetan mulberry, or long mulberry is a flowering plant species in the genus '' Morus'' found in Tibet, the Himalayas, mountainous area of Indonesia, and rain forests o ...

'' Miq. – long mulberry (Tibet, Himalayas, Indochina) *'' Morus microphylla'' *'' Morus miyabeana'' Hotta *'' Morus mongolica'' (Bureau) C.K.Schneid. *''

Morus nigra ''Morus nigra'', the black mulberry, is a species of flowering plant in the family Moraceae that is native to southwestern Asia, where it has been cultivated for so long that its precise natural range is unknown. The black mulberry is known f ...

'' L. - black mulberry (Iran,

Caucasus The Caucasus () or Caucasia (), is a region spanning Eastern Europe and Western Asia. It is situated between the Black Sea and the Caspian Sea, comprising parts of Southern Russia, Georgia, Armenia, and Azerbaijan. The Caucasus Mountains, i ...

Levant The Levant ( ) is the subregion that borders the Eastern Mediterranean, Eastern Mediterranean sea to the west, and forms the core of West Asia and the political term, Middle East, ''Middle East''. In its narrowest sense, which is in use toda ...

) *''

Morus notabilis Morus may refer to: People * Alexander Morus (1616–1670), Franco-Scottish Protestant preacher * Henryk Moruś (1943–2013), Polish serial killer * Huw Morus (1622–1709), Welsh poet * Thomas More or Morus (1478–1535), English philosopher * M ...

'' C.K.Schneid. –

Yunnan Yunnan; is an inland Provinces of China, province in Southwestern China. The province spans approximately and has a population of 47.2 million (as of 2020). The capital of the province is Kunming. The province borders the Chinese provinces ...

and

Sichuan Sichuan is a province in Southwestern China, occupying the Sichuan Basin and Tibetan Plateau—between the Jinsha River to the west, the Daba Mountains to the north, and the Yunnan–Guizhou Plateau to the south. Its capital city is Cheng ...

Provinces in China *''

Morus rubra ''Morus rubra'', commonly known as the red mulberry, is a species of Morus (plant), mulberry native to Eastern United States, eastern and central North America. It is found from Ontario, Minnesota, and Vermont south to southern Florida, and west ...

'' L. – red mulberry (eastern North America) *'' Morus serrata'' Roxb. – Tibet, Nepal, northwestern India *'' Morus trilobata'' (S.S.Chang) Z.Y.Cao – Guizhou Province in China *'' Morus wittiorum'' Hand.-Mazz. – southern China

Distribution

Black, red, and white mulberries are widespread in

Southern Europe Southern Europe is also known as Mediterranean Europe, as its geography is marked by the Mediterranean Sea. Definitions of southern Europe include some or all of these countries and regions: Albania, Andorra, Bosnia and Herzegovina, Bulgaria, C ...

, the

Middle East The Middle East (term originally coined in English language) is a geopolitical region encompassing the Arabian Peninsula, the Levant, Turkey, Egypt, Iran, and Iraq. The term came into widespread usage by the United Kingdom and western Eur ...

Central Asia Central Asia is a region of Asia consisting of Kazakhstan, Kyrgyzstan, Tajikistan, Turkmenistan, and Uzbekistan. The countries as a group are also colloquially referred to as the "-stans" as all have names ending with the Persian language, Pers ...

Northern Africa North Africa (sometimes Northern Africa) is a region encompassing the northern portion of the African continent. There is no singularly accepted scope for the region. However, it is sometimes defined as stretching from the Atlantic shores of t ...

, and the

Indian subcontinent The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...

, where the tree and the fruit have names under regional

dialect A dialect is a Variety (linguistics), variety of language spoken by a particular group of people. This may include dominant and standard language, standardized varieties as well as Vernacular language, vernacular, unwritten, or non-standardize ...

s. Black mulberry was imported to Britain in the 17th century in the hopes that it would be useful in the cultivation of silkworms. It was much used in

folk medicine Traditional medicine (also known as indigenous medicine or folk medicine) refers to the knowledge, skills, and practices rooted in the cultural beliefs of various societies, especially Indigenous groups, used for maintaining health and treatin ...

, especially in the treatment of

tapeworm Eucestoda, commonly referred to as tapeworms, is the larger of the two subclasses of flatworms in the class Cestoda (the other subclass being Cestodaria). Larvae have six posterior hooks on the scolex (head), in contrast to the ten-hooked Ce ...

s. The United States has native red mulberries, as well as imported black and white mulberries. In North America, the white mulberry is considered an invasive exotic and has taken over extensive tracts from native plant species, including the red mulberry. Mulberries are also widespread in

Greece Greece, officially the Hellenic Republic, is a country in Southeast Europe. Located on the southern tip of the Balkan peninsula, it shares land borders with Albania to the northwest, North Macedonia and Bulgaria to the north, and Turkey to th ...

, particularly in the

Peloponnese The Peloponnese ( ), Peloponnesus ( ; , ) or Morea (; ) is a peninsula and geographic region in Southern Greece, and the southernmost region of the Balkans. It is connected to the central part of the country by the Isthmus of Corinth land bridg ...

, which in the

Middle Ages In the history of Europe, the Middle Ages or medieval period lasted approximately from the 5th to the late 15th centuries, similarly to the post-classical period of global history. It began with the fall of the Western Roman Empire and ...

was known as

Morea Morea ( or ) was the name of the Peloponnese peninsula in southern Greece during the Middle Ages and the early modern period. The name was used by the Principality of Achaea, the Byzantine province known as the Despotate of the Morea, by the O ...

, deriving from the Greek word for the tree (, '). Australia has two types of native mulberries: ''

Hedycarya angustifolia ''Hedycarya angustifolia'', commonly known as native mulberry, Australian mulberry, or djelwuck is a species of flowering plant in the family ''Monimiaceae'', and is endemic to eastern Australia. It is a shrub or small tree with elliptic or egg ...

'', and ''

Pipturus argenteus ''Pipturus argenteus'', known as false stinger, native mulberry, white mulberry, white nettle, amahatyan (Chamorro language, Chamorro), and ghasooso (Carolinian language, Carolinian), is a small tree native to tropical Asia, northern and eastern ...

'', which are both from different families to

. The exotic black, red and white ''Morus'' mulberries are also commonly grown in Australian backyards. White mulberry is considered an

environmental weed Environment most often refers to: __NOTOC__ * Natural environment, referring respectively to all living and non-living things occurring naturally and the physical and biological factors along with their chemical interactions that affect an organism ...

in the states of

Queensland Queensland ( , commonly abbreviated as Qld) is a States and territories of Australia, state in northeastern Australia, and is the second-largest and third-most populous state in Australia. It is bordered by the Northern Territory, South Austr ...

and

New South Wales New South Wales (commonly abbreviated as NSW) is a States and territories of Australia, state on the Eastern states of Australia, east coast of :Australia. It borders Queensland to the north, Victoria (state), Victoria to the south, and South ...

Cultivation

Canons Ashby House and Mulberry Tree - geograph

Mulberries can be grown from seed, and this is often advised, as seedling-grown trees are generally of better shape and health. Mulberry trees grown from seed can take up to ten years to bear fruit. Mulberries are most often planted from large cuttings, which root readily. The mulberry plants allowed to grow tall have a

crown A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, parti ...

height of from ground level and a stem girth of . They are specially raised with the help of well-grown

sapling In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only pla ...

s 8–10 months old of any of the varieties recommended for rainfed areas like S-13 (for red loamy soil) or S-34 (black cotton soil), which are tolerant to drought or soil-moisture stress conditions. Usually, the plantation is raised and in block formation with a spacing of , or , as plant-to-plant and row-to-row distances. The plants are usually pruned once a year during the

monsoon season The wet season (sometimes called the rainy season or monsoon season) is the time of year when most of a region's average annual rainfall occurs. Generally, the season lasts at least one month. The term ''green season'' is also sometimes used a ...

to a height of and allowed to grow with a maximum of 8–10 shoots at the crown. Mulberry tree scion wood can easily be

grafted Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...

onto other mulberry trees during the winter, when the tree is dormant. One common scenario is converting a problematic male mulberry tree to an allergy-free female tree, by grafting all-female mulberry tree scions to a male mulberry that has been pruned back to the trunk. However, any new growth from below the graft(s) must be removed, as they would be from the original male mulberry tree.

Toxicity and allergenicity

All parts of the plant besides the ripe fruit can exude a milky sap (latex) which is mildly toxic if ingested, causing digestive distress and, at larger doses, hallucinations. It is also an irritant and may cause a skin rash on contact. Unripe green fruit may cause nausea, cramps, and be

hallucinogen Hallucinogens, also known as psychedelics, entheogens, or historically as psychotomimetics, are a large and diverse class of psychoactive drugs that can produce altered states of consciousness characterized by major alterations in thought, mo ...

ic. The berries have a

laxative Laxatives, purgatives, or aperients are substances that loosen stools and increase bowel movements. They are used to treat and prevent constipation. Laxatives vary as to how they work and the side effects they may have. Certain stimulant, lubri ...

effect; too many will cause diarrhoea. Some North American cities have banned the planting of mulberries because of the large amounts of pollen they produce, posing a potential health hazard for some pollen allergy sufferers. Only the male mulberry trees produce pollen; this lightweight pollen can be inhaled deeply into the lungs, sometimes triggering

asthma Asthma is a common long-term inflammatory disease of the airways of the lungs. It is characterized by variable and recurring symptoms, reversible airflow obstruction, and easily triggered bronchospasms. Symptoms include episodes of wh ...

. Conversely, female mulberry trees produce all-female flowers, which draw pollen and dust from the air. Because of this pollen-absorbing feature, all-female mulberry trees have an OPALS allergy scale rating of just 1 (lowest level of allergy potential), and some consider it "allergy-free".

Nutrition

Raw mulberries are 88% water, 10%

carbohydrate A carbohydrate () is a biomolecule composed of carbon (C), hydrogen (H), and oxygen (O) atoms. The typical hydrogen-to-oxygen atomic ratio is 2:1, analogous to that of water, and is represented by the empirical formula (where ''m'' and ''n'' ...

s, 1%

protein Proteins are large biomolecules and macromolecules that comprise one or more long chains of amino acid residue (biochemistry), residues. Proteins perform a vast array of functions within organisms, including Enzyme catalysis, catalysing metab ...

, and less than 1%

fat In nutrition science, nutrition, biology, and chemistry, fat usually means any ester of fatty acids, or a mixture of such chemical compound, compounds, most commonly those that occur in living beings or in food. The term often refers specif ...

. In a reference amount, raw mulberries provide 43 calories, 44% of the

Daily Value In the U.S. and Canada, the Reference Daily Intake (RDI) is used in nutrition labeling on food and dietary supplement products to indicate the daily intake level of a nutrient that is considered to be sufficient to meet the requirements of 97� ...

(DV) for

vitamin C Vitamin C (also known as ascorbic acid and ascorbate) is a water-soluble vitamin found in citrus and other fruits, berries and vegetables. It is also a generic prescription medication and in some countries is sold as a non-prescription di ...

, and 14% of the DV for

iron Iron is a chemical element; it has symbol Fe () and atomic number 26. It is a metal that belongs to the first transition series and group 8 of the periodic table. It is, by mass, the most common element on Earth, forming much of Earth's o ...

; other

micronutrient Micronutrients are essential chemicals required by organisms in small quantities to perform various biogeochemical processes and regulate physiological functions of cells and organs. By enabling these processes, micronutrients support the heal ...

s are insignificant in quantity.

Uses

Agricultural

The leaves are harvested three or four times a year by a leaf-picking method under rain-fed or semi-arid conditions, depending on the monsoon. The leaves are useful as animal fodder. The tree branches pruned in the fall (autumn; after the leaves have fallen) are cut and used to make durable baskets supporting agriculture and

animal husbandry Animal husbandry is the branch of agriculture concerned with animals that are raised for meat, animal fiber, fibre, milk, or other products. It includes day-to-day care, management, production, nutrition, selective breeding, and the raising ...

Culinary

As the fruit matures, mulberries change in texture and color, becoming succulent, plump, and juicy, resembling a

blackberry BlackBerry is a discontinued brand of handheld devices and related mobile services, originally developed and maintained by the Canadian company Research In Motion (RIM, later known as BlackBerry Limited) until 2016. The first BlackBerry device ...

. The color of the fruit does not distinguish the mulberry species, as mulberries may be white, lavender or black in color. The fruit of the black mulberry (native to southwest Asia) and the red mulberry (native to eastern North America) have distinct flavors. White mulberry fruits are typically sweet, but not tart, while red mulberries are usually deep red, sweet, and juicy. Black mulberries are large and juicy, with balanced sweetness and tartness. The fruit of the East Asian white mulberry – a species extensively naturalized in urban regions of eastern North America – has a different flavor, sometimes characterized as refreshing and a little tart, with a bit of gumminess to it and a hint of

vanilla Vanilla is a spice derived from orchids of the genus ''Vanilla (genus), Vanilla'', primarily obtained from pods of the flat-leaved vanilla (''Vanilla planifolia, V. planifolia''). ''Vanilla'' is not Autogamy, autogamous, so pollination ...

. Mulberries are used in pies, tarts, wines, cordials, and

herbal tea Herbal teas, technically known as herbal infusions, and less commonly called tisanes (UK and US , US also ), are beverages made from the infusion or decoction of herbs, spices, or other plant material in hot water. Often herb tea, or the plai ...

Jam Fruit preserves are preparations of fruits whose main preserving agent is sugar and sometimes acid, often stored in glass jars and used as a condiment or spread. There are many varieties of fruit preserves globally, distinguished by the meth ...

s and sherbets are often made from the fruit in the

Old World The "Old World" () is a term for Afro-Eurasia coined by Europeans after 1493, when they became aware of the existence of the Americas. It is used to contrast the continents of Africa, Europe, and Asia in the Eastern Hemisphere, previously ...

. In spring, new tender twigs are semisweet and can be eaten raw or cooked.

Supplement

The fruit and leaves are sold in various forms as

dietary supplement A dietary supplement is a manufactured product intended to supplement a person's diet by taking a pill (pharmacy), pill, capsule (pharmacy), capsule, tablet (pharmacy), tablet, powder, or liquid. A supplement can provide nutrients eithe ...

Silk industry

Silkworm mulberry tree zetarra marugatze arbolean3

Mulberry leaves, particularly those of the white mulberry, are ecologically important as the sole food source of the

(''Bombyx mori'', named after the mulberry genus ''Morus''), the cocoon of which is used to make

silk Silk is a natural fiber, natural protein fiber, some forms of which can be weaving, woven into textiles. The protein fiber of silk is composed mainly of fibroin and is most commonly produced by certain insect larvae to form cocoon (silk), c ...

. The wild silk moth also eats mulberry. Other

Lepidoptera Lepidoptera ( ) or lepidopterans is an order (biology), order of winged insects which includes butterflies and moths. About 180,000 species of the Lepidoptera have been described, representing 10% of the total described species of living organ ...

larva A larva (; : larvae ) is a distinct juvenile form many animals undergo before metamorphosis into their next life stage. Animals with indirect development such as insects, some arachnids, amphibians, or cnidarians typically have a larval phase ...

e—which include the

common emerald The common emerald (''Hemithea aestivaria'') is a moth of the family Geometridae. The species is found throughout the Nearctic and Palearctic regions and the Near East. It is mostly commonly found in the southern half of the Ireland and Britain. ...

, lime hawk-moth, sycamore moth, and

fall webworm The fall webworm (''Hyphantria cunea'') is a moth in the family Erebidae known principally for its larval stage, which creates the characteristic webbed nests on the tree limbs of a wide variety of hardwoods in the late summer and fall. It is con ...

—also eat the plant. The Ancient Greeks and Romans cultivated the mulberry for silkworms; at least as early as 220 AD, Emperor

Elagabalus Marcus Aurelius Antoninus (born Sextus Varius Avitus Bassianus, 204 – 13 March 222), better known by his posthumous nicknames Elagabalus ( ) and Heliogabalus ( ), was Roman emperor from 218 to 222, while he was still a teenager. His short r ...

wore a silk robe. English clergy wore silk vestments from about 1500 onwards. Mulberry and the silk industry played a role in

colonial Virginia The Colony of Virginia was a British Empire, British colonial settlement in North America from 1606 to 1776. The first effort to create an English settlement in the area was chartered in 1584 and established in 1585; the resulting Roanoke Colo ...

Pigment

Mulberry fruit color derives from

anthocyanin Anthocyanins (), also called anthocyans, are solubility, water-soluble vacuole, vacuolar pigments that, depending on their pH, may appear red, purple, blue, or black. In 1835, the German pharmacist Ludwig Clamor Marquart named a chemical compou ...

s, which have unknown effects in humans.Direct link
/ref> Anthocyanins are responsible for the attractive colors of fresh plant foods, including orange, red, purple, black, and blue. These colors are water-soluble and easily extractable, yielding natural

food colorant Food coloring, color additive or colorant is any dye, pigment, or substance that imparts color when it is added to food or beverages. Colorants can be supplied as liquids, powders, gels, or pastes. Food coloring is commonly used in commercial ...

s. Due to a growing demand for natural food colorants, they have numerous applications in the food industry. A cheap and industrially feasible method has been developed to extract anthocyanins from mulberry fruit that could be used as a fabric dye or food colorant of high

color value Lightness is a visual perception of the luminance (L) of an object. It is often judged relative to a similarly lit object. In colorimetry and color appearance models, lightness is a prediction of how an illuminated color will appear to a stand ...

. Scientists found that, of 31 Chinese mulberry cultivars tested, the total anthocyanin yield varied from 148 to 2725 mg/L of fruit juice. Sugars, acids, and vitamins of the fruit remained intact in the residual juice after removal of the anthocyanins, indicating that the juice may be used for other food products. Mulberry

germplasm Germplasm refers to genetic resources such as seeds, tissues, and DNA sequences that are maintained for the purpose of animal and plant breeding, conservation efforts, agriculture, and other research uses. These resources may take the form of s ...

resources may be used for: * exploration and collection of fruit yielding mulberry species * their characterization, cataloging, and evaluation for anthocyanin content by using traditional, as well as modern, means and

biotechnology Biotechnology is a multidisciplinary field that involves the integration of natural sciences and Engineering Science, engineering sciences in order to achieve the application of organisms and parts thereof for products and services. Specialists ...

tools * developing an information system about these cultivars and varieties * training and global coordination of genetic stocks * evolving suitable breeding strategies to improve the anthocyanin content in potential breeds by collaboration with various research stations in the field of sericulture, plant genetics, and breeding, biotechnology and

pharmacology Pharmacology is the science of drugs and medications, including a substance's origin, composition, pharmacokinetics, pharmacodynamics, therapeutic use, and toxicology. More specifically, it is the study of the interactions that occur betwee ...

Paper

During the Angkorian age of the

Khmer Empire The Khmer Empire was an empire in Southeast Asia, centered on Hydraulic empire, hydraulic cities in what is now northern Cambodia. Known as Kambuja (; ) by its inhabitants, it grew out of the former civilization of Chenla and lasted from 802 t ...

, monks at Buddhist temples made paper from the bark of mulberry trees. The paper was used to make books, known as '' kraing''.

Tengujo is a specialist Japanese paper. It is an extremely thin kōzo paper that is almost transparent. One of its uses is for archival conservation. It has also been used for lighting design. The paper is produced in the Kochi prefecture of Japan by th ...

is the thinnest paper in the world. It is produced in Japan and made with kozo (stems of mulberry trees). Traditional Japanese washi paper is often created from parts of the mulberry tree.

Wood

The wood of mulberry trees is used for

barrel aging A barrel or cask is a hollow cylindrical container with a bulging center, longer than it is wide. They are traditionally made of wooden staves and bound by wooden or metal hoops. The word vat is often used for large containers for liquids, ...

Țuică Țuică () is a traditional Romanian Distilled beverage, spirit that contains ~ 24–86% alcohol by volume (usually 40–55%), prepared only from plums. Other spirits that are produced from other fruit or from a cereal grain are called "rachiu" ...

, a traditional Romanian plum brandy.

Culture

Babylonia Babylonia (; , ) was an Ancient history, ancient Akkadian language, Akkadian-speaking state and cultural area based in the city of Babylon in central-southern Mesopotamia (present-day Iraq and parts of Kuwait, Syria and Iran). It emerged as a ...

etiological myth An origin myth is a type of myth that explains the beginnings of a natural or social aspect of the world. Creation myths are a type of origin myth narrating the formation of the universe. However, numerous cultures have stories that take place af ...

, which

Ovid Publius Ovidius Naso (; 20 March 43 BC – AD 17/18), known in English as Ovid ( ), was a Augustan literature (ancient Rome), Roman poet who lived during the reign of Augustus. He was a younger contemporary of Virgil and Horace, with whom he i ...

incorporated in his ''

Metamorphoses The ''Metamorphoses'' (, , ) is a Latin Narrative poetry, narrative poem from 8 Common Era, CE by the Ancient Rome, Roman poet Ovid. It is considered his ''Masterpiece, magnum opus''. The poem chronicles the history of the world from its Cre ...

'', attributes the reddish-purple color of the mulberry fruits to the tragic deaths of the lovers

Pyramus and Thisbe In Greek mythology, Pyramus and Thisbe () are a pair of ill-fated lovers from Babylon, whose story is best known from Ovid's narrative poem ''Metamorphoses''. The tragic myth has been retold by many authors. Pyramus and Thisbe's parents, drive ...

. Meeting under a mulberry tree (probably the native ''

''), Thisbe dies by suicide by sword after Pyramus does the same, he having believed, on finding her bloodstained cloak, that she was killed by a lion. Their splashed blood stained the previously white fruit, and the gods forever changed the mulberry's colour to honour their forbidden love. In the Old Testament's

1 Maccabees 1 Maccabees, also known as the First Book of Maccabees, First Maccabees, and abbreviated as 1 Macc., is a deuterocanonical book which details the history of the Maccabean Revolt against the Seleucid Empire as well as the founding and earliest hi ...

, the

Seleucids The Seleucid Empire ( ) was a Greek state in West Asia during the Hellenistic period. It was founded in 312 BC by the Macedonian general Seleucus I Nicator, following the division of the Macedonian Empire founded by Alexander the Great, ...

used the "blood of grapes and mulberries" to provoke their

war elephant A war elephant is an elephant that is Animal training, trained and guided by humans for combat purposes. Historically, the war elephant's main use was to charge (warfare), charge the enemy, break their ranks, and instill terror and fear. Elep ...

s in preparation for battle against

Jewish rebels Jews (, , ), or the Jewish people, are an ethnoreligious group and nation, originating from the Israelites of History of ancient Israel and Judah, ancient Israel and Judah. They also traditionally adhere to Judaism. Jewish ethnicity, rel ...

. The

nursery rhyme A nursery rhyme is a traditional poem or song for children in Britain and other European countries, but usage of the term dates only from the late 18th/early 19th century. The term Mother Goose rhymes is interchangeable with nursery rhymes. Fr ...

Here We Go Round the Mulberry Bush "Here We Go Round the Mulberry Bush" (also titled "Mulberry Bush" or "This Is the Way") is an English nursery rhyme and singing game. It has a Roud Folk Song Index number of 7882. It uses the tune which Nancy Dawson danced into fame in ''The ...

" uses the tree in the refrain, as do some contemporary American versions of the nursery rhyme " Pop Goes the Weasel".

Vincent van Gogh Vincent Willem van Gogh (; 30 March 185329 July 1890) was a Dutch Post-Impressionist painter who is among the most famous and influential figures in the history of Western art. In just over a decade, he created approximately 2,100 artworks ...

featured the mulberry tree in some of his paintings, notably ''Mulberry Tree'' (, 1889, now in

Pasadena Pasadena ( ) is a city in Los Angeles County, California, United States, northeast of downtown Los Angeles. It is the most populous city and the primary cultural center of the San Gabriel Valley. Old Pasadena is the city's original commercial d ...

Norton Simon Museum The Norton Simon Museum is an art museum located in Pasadena, California. It was previously known as the Pasadena Art Institute and the Pasadena Art Museum and displays numerous sculptures on its grounds. Overview The Norton Simon collections ...

). He painted it after a stay at an asylum, and he considered it a technical success.

References

The Times of India ''The Times of India'' (''TOI'') is an Indian English-language daily newspaper and digital news media owned and managed by the Times Group. It is the List of newspapers in India by circulation, third-largest newspaper in India by circulation an ...

''
The Morus Londinium project - Mulberry tree heritage in London, UK
{{Authority control Berries Fruit trees Medicinal plants Moraceae genera Taxa named by Carl Linnaeus Pyramus and Thisbe