Microcleididae is an

extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...

family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...

of basal plesiosauroid

plesiosaur The Plesiosauria or plesiosaurs are an Order (biology), order or clade of extinct Mesozoic marine reptiles, belonging to the Sauropterygia. Plesiosaurs first appeared in the latest Triassic Period (geology), Period, possibly in the Rhaetian st ...

s from the

Early Jurassic The Early Jurassic Epoch (geology), Epoch (in chronostratigraphy corresponding to the Lower Jurassic series (stratigraphy), Series) is the earliest of three epochs of the Jurassic Period. The Early Jurassic starts immediately after the Triassic� ...

(middle

Sinemurian In the geologic timescale, the Sinemurian is an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between 199.5 ±0.3 annu ...

to late

Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...

stages) of

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...

Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...

and the

United Kingdom The United Kingdom of Great Britain and Northern Ireland, commonly known as the United Kingdom (UK) or Britain, is a country in Northwestern Europe, off the coast of European mainland, the continental mainland. It comprises England, Scotlan ...

. Currently, the oldest and the most known microcleidid is '' Eretmosaurus'' from the middle Sinemurian of the United Kingdom. Microcleididae was formally named and described by Roger B. J. Benson, Mark Evans and Patrick S. Druckenmiller in 2012.

History and Phylogeny

Großman (2007) referred informally to "microcleidid

elasmosaur Elasmosauridae, often called elasmosaurs or elasmosaurids, is an extinct family (biology), family of plesiosaurs that lived from the Hauterivian Stage (stratigraphy), stage of the Early Cretaceous to the Maastrichtian stage of the Late Cretaceou ...

s", a clade comprising the three taxa referred to the Microcleididae by Benson ''et al.'' (2012). Ketchum and Benson (2010) found the traditional "microcleidid" genera to nest within a monophyletic Plesiosauridae, in a close position to ''

Plesiosaurus ''Plesiosaurus'' (Greek: ' ('), near to + ' ('), lizard) is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias of England. It is distinguishable by ...

''. However, Ketchum and Benson (2011) performed a phylogenetic analysis which found a monospecific Plesiosauridae, and a monophyletic Microcleididae. Smith ''et al.'' (in press) described a new "microcleidid" '' Lusonectes'', and obtained similar results in a different phylogenetic analysis. However, the family has not formally been erected until Benson ''et al.'' (2012) also found the same results in their analysis. Microcleidus1DB

To avoid further confusion between Plesiosauridae and Microcleididae, Benson ''et al.'' (2012) proposed a stem-based definition for this taxon, utilizing ''Plesiosaurus'' as an external specifier. Microcleididae was defined as "'' Microcleidus homalospondylus'' and all taxa more closely related to it than to ''

Plesiosaurus dolichodeirus ''Plesiosaurus'' (Greek: ' ('), near to + ' ('), lizard) is a genus of extinct, large marine sauropterygian reptile that lived during the Early Jurassic. It is known by nearly complete skeletons from the Lias Group, Lias of England. It is disting ...

'', '' Cryptoclidus eurymerus'', '' Elasmosaurus platyurus'', '' Leptocleidus superstes'', '' Pliosaurus brachydeirus'' or '' Polycotylus latipinnis''." Benson ''et al.'' (2012) found strong support for Microcleididae, however their diagnosis focused on unambiguous

postcrania The postcranium ("behind the cranium"; plural: postcrania) or postcranial skeleton in zoology and vertebrate paleontology is the skeleton apart from the skull. The postcranium encompasses the axial skeleton, which includes the entirety of the verte ...

synapomorphies In phylogenetics, an apomorphy (or derived trait) is a novel character or character state that has evolved from its ancestral form (or plesiomorphy). A synapomorphy is an apomorphy shared by two or more taxa and is therefore hypothesized to ...

since cranial material of basal microcleidids is unknown. The synapomorphies are: widely separated posterior

cervical rib Cervical ribs are the ribs of the neck in many tetrapods. In most mammals, including humans, cervical ribs are not normally present as separate structures. They can, however, occur as a pathology. In humans, pathological cervical ribs are usually no ...

facets, posteriormost dorsal rib facets split between centrum and

neural arch Each vertebra (: vertebrae) is an irregular bone with a complex structure composed of bone and some hyaline cartilage, that make up the vertebral column or spine, of vertebrates. The proportions of the vertebrae differ according to their spinal ...

, medial surface of the iliac blade anteroposteriorly concave and a prominent flange extends anteriorly from the proximal half of the radius. This analysis focused on basal plesiosaurs and therefore only one derived pliosaurid and one

cryptoclidia Plesiosauroidea (; Ancient Greek, Greek: 'near, close to' and 'lizard') is an extinct clade of carnivore, carnivorous Marine (ocean), marine Reptilia, reptiles. They have the snake-like longest neck to body ratio of any reptile. Plesiosauroid ...

n were included while

elasmosaurid Elasmosauridae, often called elasmosaurs or elasmosaurids, is an extinct family of plesiosaurs that lived from the Hauterivian stage of the Early Cretaceous to the Maastrichtian stage of the Late Cretaceous period (c. 130 to 66 mya). The taxo ...

s weren't included at all. The

cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...

below follows the topology from Benson ''et al.'' (2012) analysis.

References

{{Taxonbar, from=Q6839368 Jurassic plesiosaurs Sinemurian first appearances Early Jurassic extinctions Plesiosauroids Sauropterygian families