Mesoblattinidae
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Mesoblattinidae is an extinct, problematic family of cockroaches known from the Mesozoic. It was formerly considered a
wastebasket taxon Wastebasket taxon (also called a wastebin taxon, dustbin taxon or catch-all taxon) is a term used by some taxonomists to refer to a taxon that has the purpose of classifying organisms that do not fit anywhere else. They are typically defined by e ...
for Mesozoic cockroaches, but the family has subsequently been better defined, with many taxa transferred to Caloblattinidae. It is considered to have close affinities with
Blattidae Blattidae is a cockroach family in the order Blattodea containing several of the most common household cockroaches. Notable species include: * '' Blatta orientalis'': Oriental cockroach, * Common shining cockroach: (''Drymaplaneta communis'' ...
and
Ectobiidae Ectobiidae (formerly Blattellidae) is a family of the order Blattodea (cockroaches). This family contains many of the smaller common household pest cockroaches, among others. They are sometimes called wood cockroaches. A few notable species inclu ...
, as well as possibly
Blaberidae Giant cockroaches, or blaberids (family (biology), family Blaberidae), are the second-largest cockroach family by number of species. Mostly distributed in warmer climates worldwide, this family is based on the American genus ''Blaberus'', but mu ...
. The family first appeared by the Early
Jurassic The Jurassic ( ) is a Geological period, geologic period and System (stratigraphy), stratigraphic system that spanned from the end of the Triassic Period million years ago (Mya) to the beginning of the Cretaceous Period, approximately 143.1 Mya. ...
. They are considered to represent amongst the oldest groups of modern cockroaches, and like them are thought to have laid oothecae. Due to the poor ability of forewing venation to correctly classify cockroaches to extensive
homoplasy Homoplasy, in biology and phylogenetics, is the term used to describe a feature that has been gained or lost independently in separate lineages over the course of evolution. This is different from homology, which is the term used to characterize ...
, the value of this family as a taxonomic unit has been strongly questioned, with some authors considering the family a ''
nomen dubium In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application. Zoology In case of a ''nomen dubium,'' it may be impossible to determine whether a ...
''.


Systematics

Based on * †'' Actinoblattula'' Handlirsch 1906 Çakrazboz Formation, Turkey, Late Triassic ( Rhetian) * †'' Archimesoblatta'' Vršanský 2003 Kota Formation, India, Early-Middle Jurassic, Doronino Formation, Russia, Early Cretaceous (
Barremian The Barremian is an age in the geologic timescale (or a chronostratigraphic stage) between 125.77 Ma (million years ago) and 121.4 ± 1.0 Ma (Historically, this stage was placed at 129.4 million to approximately 125 million years ago) It is a ...
) Dzun-Bain Formation, Mongolia,
Zaza Formation The Zaza Formation is a geological formation located in Buryatia (Russia). It dates to the Lower Cretaceous. The age of the formation is disputed, and is considered likely to be Valanginian-Hauterivian, or Aptian in age. It comprises sandstones, ...
, Russia Early Cretaceous (
Aptian The Aptian is an age (geology), age in the geologic timescale or a stage (stratigraphy), stage in the stratigraphic column. It is a subdivision of the Early Cretaceous, Early or Lower Cretaceous epoch (geology), Epoch or series (stratigraphy), S ...
) * †'' Artitocoblatta'' Handlirsch 1906
Purbeck Group The Purbeck Group is an Upper Jurassic to Lower Cretaceous lithostratigraphy, lithostratigraphic group (stratigraphy), group (a sequence of rock strata) in south-east England. The name is derived from the district known as the Isle of Purbeck in ...
, United Kingdom, Early Cretaceous (
Berriasian In the geological timescale, the Berriasian is an age/ stage of the Early/Lower Cretaceous. It is the oldest subdivision in the entire Cretaceous. It has been taken to span the time between 143.1 ±0.6 Ma and 137.05 ± 0.2 (million years ago) ...
), La Pedrera de Rúbies Formation, Spain, Barremian * †'' Austroblattula'' Tillyard 1919, Blackstone Formation, Australia, Late Triassic (
Norian The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
) * †'' Basiblattina'' Zhang 1997 Dalazi Formation, China, Aptian * †'' Breviblattina'' Vršanský 2004 Shar-Teg, Mongolia, Late Jurassic (
Tithonian In the geological timescale, the Tithonian is the latest age (geology), age of the Late Jurassic Epoch and the uppermost stage (stratigraphy), stage of the Upper Jurassic Series. It spans the time between 149.2 ±0.7 annum, Ma and 143.1 ±0.6 (mi ...
) * †'' Durdlestoneia'' Handlirsch 1906 Purbeck Group, United Kingdom, Berriasian * †'' Gondwablatta'' Vršanský 2004 Tayasir Formation, Israel, Berriasian * †'' Grandocularis'' Kaddumi 2005 Jordanian amber, Early Cretaceous (
Albian The Albian is both an age (geology), age of the geologic timescale and a stage (stratigraphy), stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early Cretaceous, Early/Lower Cretaceous epoch (geology), Epoch/s ...
) * †'' Hispanoblatta'' Martínez-Delclòs 1993, Las Hoyas, Spain, Barremian * †'' Hongaya'' Handlirsch 1908 Vietnam, Rhaetian * †'' Jingyuanoblatta'' Lin 1982 China,
Middle Jurassic The Middle Jurassic is the second Epoch (geology), epoch of the Jurassic Period (geology), Period. It lasted from about 174.1 to 161.5 million years ago. Fossils of land-dwelling animals, such as dinosaurs, from the Middle Jurassic are relativel ...
* †'' Karatavoblatta'' Vishnyakova 1968 Karabastau Formation, Kazakhstan, Late Jurassic
Yixian Formation The Yixian Formation (; formerly Romanization of Chinese, transcribed as Yihsien Formation or Yixiang Formation) is a geological formation in Jinzhou, Liaoning, People's Republic of China, that spans the Barremian stage of the Early Cretaceous. I ...
, China, Aptian * †'' Laiyangia'' Grabau 1923 Laiyang Formation, China, Aptian * †'' Lithoblatta'' Handlirsch 1906 Solnhofen Limestone, Germany, Tithonian * †'' Malmoblattina'' Handlirsch 1906 Purbeck Group, United Kingdom, Berriasian * †'' Mesoblatta'' Hinkelman 2020,
Burmese amber Burmese amber, also known as Burmite or Kachin amber, is amber from the Hukawng Valley in northern Myanmar. The amber is dated to around 100 million years ago, during the latest Albian to earliest Cenomanian ages of the mid-Cretaceous period. Th ...
, Myanmar, Late Cretaceous (
Cenomanian The Cenomanian is, in the International Commission on Stratigraphy's (ICS) geological timescale, the oldest or earliest age (geology), age of the Late Cretaceous epoch (geology), Epoch or the lowest stage (stratigraphy), stage of the Upper Cretace ...
) * †'' Mesoblattellina'' Mendes 2000
Crato Formation The Crato Formation is a geologic formation (stratigraphy), formation of Early Cretaceous (Aptian) age in northeastern Brazil's Araripe Basin. It is an important Lagerstätten, Lagerstätte (undisturbed fossil accumulation) for palaeontology, pa ...
, Brazil, Aptian * †'' Mesoblattina'' Geinitz 1880 Guanyintan Formation, China Early Jurassic (Hettangian) Xiangxi Formation, China Early Jurassic (
Pliensbachian The Pliensbachian is an age of the geologic timescale and stage in the stratigraphic column. It is part of the Early or Lower Jurassic Epoch or Series and spans the time between 192.9 ±0.3 Ma and 184.2 ±0.3 Ma (million years ago). The Plie ...
), Dashankou Formation, Middle Jurassic ( Aalenian) Hanshan Formation, China, Middle Jurassic (
Bajocian In the geologic timescale, the Bajocian is an age and stage in the Middle Jurassic. It lasted from approximately 170.9 ±0.8 Ma to around 168.2 ±1.2 Ma (million years ago). The Bajocian Age succeeds the Aalenian Age and precedes the Bathonia ...
), Houcheng Formation, Houjiatun Formation, Haifanggou Formation, China, Middle Jurassic (
Callovian In the geologic timescale, the Callovian is an age and stage in the Middle Jurassic, lasting between 165.3 ± 1.1 Ma (million years ago) and 161.5 ± 1.0 Ma. It is the last stage of the Middle Jurassic, following the Bathonian and preceding the ...
) Green Series, Germany, Early Jurassic (
Toarcian The Toarcian is, in the International Commission on Stratigraphy, ICS' geologic timescale, an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic. It spans the time between 184.2 Megaannum, Ma (million ...
), Kyzyl-Kiya, Kyrgyzstan, Pliensbachian Zhargalant Formation, Mongolia, Toarcian, Insektenmergel Formation, Switzerland, Hettangian, Xiagou Formation, China, Aptian Jinju Formation, South Korea, Albian La Pedrera de Rúbies Formation, Spain, Barremian, Purbeck Group, United Kingdom, Berriasian * †'' Blattulopsis'' Pinto 1990 Anfiteatro de Ticó Formation, Argentina, Aptian * †'' Brachymesoblatta'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Aptian * †'' Latiblatta'' Vishnyakova 1968 Karabastau Formation, Kazakhstan, Late Jurassic * †'' Nannoblattina'' Scudder 1886
Charmouth Mudstone Formation The Charmouth Mudstone Formation is a Formation (geology), geological formation in England, dating to the Early Jurassic (Sinemurian–Pliensbachian). It forms part of the lower Lias Group. It is most prominently exposed at its type locality in cl ...
, United Kingdom, Early Jurassic (
Sinemurian In the geologic timescale, the Sinemurian is an age (geology), age and stage (stratigraphy), stage in the Early Jurassic, Early or Lower Jurassic epoch (geology), Epoch or series (stratigraphy), Series. It spans the time between 199.5 ±0.3 annu ...
) Purbeck Group, United Kingdom, Berriasian * †'' Rhipidoblattinopsis'' Vishnyakova 1968 Karabastau Formation, Kazakhstan, Late Jurassic * †'' Tarakanula'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Aptian * †'' Mesoblattinopsis'' Mendes 2000 Crato Formation, Brazil, Aptian * †'' Mieroblattina'' Vršanský and Makhoul 2013
Sannine Formation The Sannine Formation, also called the Sannine Limestone, is a Cretaceous geologic formation in Lebanon. It is a Konservat-Lagerstätte that contains a high diversity of well-preserved fish, reptiles, and invertebrates from the Tethys Ocean wit ...
, Lebanon, Cenomanian * †'' Mongolblatta'' Vršanský 2004 Shar-Teg, Mongolia, Tithonian, Doronino Formation, Russia, Barremian * †'' Nipponoblatta'' Fujiyama 1974 Guanyintan Formation, China,
Hettangian The Hettangian is the earliest age and lowest stage of the Jurassic Period of the geologic timescale. It spans the time between 201.3 ± 0.2 Ma and 199.3 ± 0.3 Ma (million years ago). The Hettangian follows the Rhaetian (part of the Triass ...
, Nishinakayama Formation, Japan, Toarcian, Yixian Formation, China, Aptian * †'' Nogueroblatta'' Martínez-Delclòs 1993 La Pedrera de Rúbies Formation, Las Hoyas, Spain, Barremian * †'' Nymphoblatta'' Vršanský and Grimaldi 2004
Lebanese amber Lebanese amber is fossilized resin found in Lebanon and its surroundings. It dates back approximately 130-125 million years to the Barremian of the Early Cretaceous. It formed on what was then the northern coast of Gondwana, believed to be a tropic ...
, Barremian * †'' Perlucipecta'' Wei and Ren 2013 Crato Formation, Brazil, Aptian Yixian Formation, China, Aptian * †'' Praeblattella'' Vršanský 2003 Dzun-Bain Formation, Mongolia, Aptian Zaza Formation, Russia, Aptian * †'' Pulchellablatta'' Martins-Neto and Gallego 2005 Los Rastros Formation, Argentina, Late Triassic (
Carnian The Carnian (less commonly, Karnian) is the lowermost stage (stratigraphy), stage of the Upper Triassic series (stratigraphy), Series (or earliest age (geology), age of the Late Triassic Epoch (reference date), Epoch). It lasted from 237 to 227.3 ...
) * †'' Rhaetoblattina'' Handlirsch 1908 Vietnam, Rhaetian * †'' Schambeloblattina'' Handlirsch 1906 Insektenmergel Formation, Switzerland, Hettangian * †'' Sivis'' Vršanský 2009 Charentese amber, France, Albian * †'' Spinaeblattina'' Hinkelman 2019 Yixian Formation, China, Aptian, Burmese amber, Myanmar, Cenomanian * †'' Stantoniella'' Handlirsch 1907 Judith River Formation, Montana,
Campanian The Campanian is the fifth of six ages of the Late Cretaceous epoch on the geologic timescale of the International Commission on Stratigraphy (ICS). In chronostratigraphy, it is the fifth of six stages in the Upper Cretaceous Series. Campa ...
* †'' Stenotegmina'' Lin 1980 Funing Group, China,
Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...
* †'' Summatiblatta'' Lin 1986 Guanyintan Formation, China, Hettangian * †'' Turoniblatta'' Vršanský 2004 Ora Formation, Israel, Late Jurassic (
Turonian The Turonian is, in the International Commission on Stratigraphy, ICS' geologic timescale, the second age (geology), age in the Late Cretaceous epoch (geology), Epoch, or a stage (stratigraphy), stage in the Upper Cretaceous series (stratigraphy), ...
) * †'' Voltziablatta'' Papier and Grauvogel-Stamm 1995 Grès à Voltzia, France,
Anisian In the geologic timescale, the Anisian is the lower stage (stratigraphy), stage or earliest geologic age, age of the Middle Triassic series (stratigraphy), series or geologic epoch, epoch and lasted from million years ago until million years ag ...


References

{{Taxonbar, from=Q4044051 † Cockroaches Prehistoric insect families