The moonfish of the genus ''Mene'', the sole extant genus of the family Menidae, are disk-shaped fish which bear a vague resemblance to

gourami Gouramis, or gouramies , are a group of fresh water, freshwater Anabantiformes, anabantiform fish that comprise the family (biology), family Osphronemidae. The fish are native to Asia—from the Indian Subcontinent to Southeast Asia and northeas ...

, thanks to their thread-like pelvic fins. Today, the genus is represented only by ''

Mene maculata ''Mene maculata'', the moonfish, is the only extant member of the genus ''Mene'' and of the family Menidae. The body is highly compressed laterally and very deep vertically. The ventral profile is steep, with a sharp ventral edge. The caudal (tai ...

'' of the

Indo-Pacific The Indo-Pacific is a vast biogeographic region of Earth. In a narrow sense, sometimes known as the Indo-West Pacific or Indo-Pacific Asia, it comprises the tropical waters of the Indian Ocean, the western and central Pacific Ocean, and the ...

, where it is a popular food fish, especially in the

Philippines The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...

, where it is known as ''bilong-bilong'', ''chabita'', ''hiwas'' or ''tahas''.

Taxonomy

Anatomical and recent molecular studies strongly suggest a relationship with the

pomfret Pomfrets are scombriform fish belonging to the family Bramidae. The family currently includes 20 species across seven genera. Several species are important food sources for humans, especially ''Brama brama'' in South Asia. The earlier form of ...

dolphinfish ''Coryphaena'' is a genus of marine ray-finned fishes known as the dolphinfishes, and is currently the only known genus in the family Coryphaenidae. The generic name is from Greek κορυφή (''koryphē'', "crown, top") and -αινα (-''aina ...

es,

remora The remora (), sometimes called suckerfish or sharksucker, is any of a family (Echeneidae) of ray-finned fish in the order Carangiformes. Depending on species, they grow to long. Their distinctive first dorsal fins take the form of a modified ...

s and the jacks in the

order Order, ORDER or Orders may refer to: * A socio-political or established or existing order, e.g. World order, Ancien Regime, Pax Britannica * Categorization, the process in which ideas and objects are recognized, differentiated, and understood ...

Carangiformes Carangiformes is a large, diverse order of ray-finned fishes within the clade Percomorpha. It is part of a sister clade to the Ovalentaria, alongside its sister group, the Anabantaria (including Anabantiformes and Synbranchiformes). The Cara ...

Fossil record

As a genus, ''Mene'' has a long fossil history, with species found in marine sediments throughout the

Cenozoic Era The Cenozoic Era ( ; ) is Earth's current geological Era (geology), era, representing the last 66million years of Earth's history. It is characterized by the dominance of mammals, Insect, insects, birds and flowering plant, angiosperms (floweri ...

.Matt Friedman, and G. David Johnson. “A New Species of Mene (Perciformes: Menidae) from the Paleocene of South America, with Notes on Paleoenvironment and a Brief Review of Menid Fishes.” Journal of Vertebrate Paleontology, vol. 25, no. 4, 2005, pp. 770–83. JSTOR, http://www.jstor.org/stable/4524505. The earliest accepted species, '' M. purdyi'' from the

Paleocene The Paleocene ( ), or Palaeocene, is a geological epoch (geology), epoch that lasted from about 66 to 56 mya (unit), million years ago (mya). It is the first epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), ...

of Peru, resemble later species, such as '' M. rhombea'' of the Monte

Bolca Bolca is a village in the Veneto, on the southern margin of the Italian Alps. It is a of the ''comune'' of Vestenanova, in the province of Verona. The area is famous for the marine fossils from the lagerstätte of '' Monte Bolca''. It was on ...

lagerstätte A Fossil-Lagerstätte (, from ''Lager'' 'storage, lair' '' Stätte'' 'place'; plural ''Lagerstätten'') is a sedimentary deposit that preserves an exceptionally high amount of palaeontological information. ''Konzentrat-Lagerstätten'' preserv ...

, and even the living species, ''M. maculata''. Experts remain undecided whether the

Tunisia Tunisia, officially the Republic of Tunisia, is a country in the Maghreb region of North Africa. It is bordered by Algeria to the west and southwest, Libya to the southeast, and the Mediterranean Sea to the north and east. Tunisia also shares m ...

n species, '' M. phosphatica'' is from the Lower Paleocene, thus making it older than ''M. purdyi'', or whether it is from the

Ypresian In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...

epoch of the Eocene. Almost all of the species are known primarily from the

Paleogene The Paleogene Period ( ; also spelled Palaeogene or Palæogene) is a geologic period and system that spans 43 million years from the end of the Cretaceous Period Ma (million years ago) to the beginning of the Neogene Period Ma. It is the fir ...

; the

Neogene The Neogene ( ,) is a geologic period and system that spans 20.45 million years from the end of the Paleogene Period million years ago ( Mya) to the beginning of the present Quaternary Period million years ago. It is the second period of th ...

record is rather sparse, if not totally nonexistent, with some

otolith An otolith (, ' ear + , ', a stone), also called otoconium, statolith, or statoconium, is a calcium carbonate structure in the saccule or utricle (ear), utricle of the inner ear, specifically in the vestibular system of vertebrates. The saccule ...

s found in

Miocene The Miocene ( ) is the first epoch (geology), geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and mea ...

strata In geology and related fields, a stratum (: strata) is a layer of Rock (geology), rock or sediment characterized by certain Lithology, lithologic properties or attributes that distinguish it from adjacent layers from which it is separated by v ...

, and no whole or even partial specimens known from

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...

strata. The first species of the genus to be described was not the extant ''M. maculata'', but rather the extinct fossil species '' Mene rhombea''; however, ''M. rhombea'' was not initially classified in ''Mene'', with the genus name being coined for the extant species. File:Mene_rhombea_and_Mene_oblonga.jpg, Restoration comparing †'' Mene rhombea'' and †'' Mene oblonga'' File:Mene oblonga 443.JPG, †''Mene oblonga'' File:Menidae - Mene rhombea 001.jpg, †''Mene rhombea'', from

File:Mene sp. from the Early Eocene Denmark.jpg, ''Mene sp'', from

Fur Formation The Fur Formation is a marine geological formation of Ypresian ( Lower Eocene Epoch, c. 56.0-54.5 Ma) age which crops out in the Limfjord region of northern Denmark from Silstrup via Mors and Fur to Ertebølle, and can be seen in many cliffs and ...

Denmark Denmark is a Nordic countries, Nordic country in Northern Europe. It is the metropole and most populous constituent of the Kingdom of Denmark,, . also known as the Danish Realm, a constitutionally unitary state that includes the Autonomous a ...

Timeline

ImageSize = width:1000px height:auto barincrement:15px PlotArea = left:10px bottom:50px top:10px right:10px Period = from:-65.5 till:10 TimeAxis = orientation:horizontal ScaleMajor = unit:year increment:5 start:-65.5 ScaleMinor = unit:year increment:1 start:-65.5 TimeAxis = orientation:hor AlignBars = justify Colors = #legends id:CAR value:claret id:ANK value:rgb(0.4,0.3,0.196) id:HER value:teal id:HAD value:green id:OMN value:blue id:black value:black id:white value:white id:cenozoic value:rgb(0.54,0.54,0.258) id:paleogene value:rgb(0.99,0.6,0.32) id:paleocene value:rgb(0.99,0.65,0.37) id:eocene value:rgb(0.99,0.71,0.42) id:oligocene value:rgb(0.99,0.75,0.48) id:neogene value:rgb(0.999999,0.9,0.1) id:miocene value:rgb(0.999999,0.999999,0) id:pliocene value:rgb(0.97,0.98,0.68) id:quaternary value:rgb(0.98,0.98,0.5) id:pleistocene value:rgb(0.999999,0.95,0.68) id:holocene value:rgb(0.999,0.95,0.88) BarData= bar:eratop bar:space bar:periodtop bar:space bar:NAM1 bar:space bar:period bar:space bar:era PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 shift:(7,-4) bar:periodtop from: -65.5 till: -55.8 color:paleocene text:

from: -55.8 till: -33.9 color:eocene text:

Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...

from: -33.9 till: -23.03 color:oligocene text:

Oligocene The Oligocene ( ) is a geologic epoch (geology), epoch of the Paleogene Geologic time scale, Period that extends from about 33.9 million to 23 million years before the present ( to ). As with other older geologic periods, the rock beds that defin ...

from: -23.03 till: -5.332 color:miocene text:

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:

Pleist. The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...

from: -0.0117 till: 0 color:holocene text: H. bar:eratop from: -65.5 till: -23.03 color:paleogene text:

from: -23.03 till: -2.588 color:neogene text:

from: -2.588 till: 0 color:quaternary text: Q. PlotData= align:left fontsize:M mark:(line,white) width:5 anchor:till align:left color:eocene bar:NAM1 from: -55.8 till: 0 text: Mene PlotData= align:center textcolor:black fontsize:M mark:(line,black) width:25 bar:period from: -65.5 till: -55.8 color:paleocene text:

from: -55.8 till: -33.9 color:eocene text:

from: -33.9 till: -23.03 color:oligocene text:

from: -23.03 till: -5.332 color:miocene text:

from: -5.332 till: -2.588 color:pliocene text: Plio. from: -2.588 till: -0.0117 color:pleistocene text:

from: -0.0117 till: 0 color:holocene text: H. bar:era from: -65.5 till: -23.03 color:paleogene text:

from: -23.03 till: -2.588 color:neogene text:

from: -2.588 till: 0 color:quaternary text: Q.

References

External links

"A new species of Mene (Perciformes: Menidae) from the Paleocene of South America, with notes on paleoenvironment and a brief review of menid fishes."
{{Taxonbar, from=Q325846 Menidae Paleocene fish Eocene fish Oligocene fish Miocene fish Prehistoric fish of Africa Paleogene fish of Asia Cenozoic fish of Europe Prehistoric fish of South America Extant Thanetian first appearances Thanetian genera Ypresian genera Lutetian genera Bartonian genera Priabonian genera Rupelian genera Chattian genera Aquitanian genera Burdigalian genera Langhian genera Serravallian genera Tortonian genera Messinian genera Holocene genera Fish genera with one living species