Margaret Formation
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The Margaret Formation is a
geologic formation A geological formation, or simply formation, is a body of rock having a consistent set of physical characteristics ( lithology) that distinguishes it from adjacent bodies of rock, and which occupies a particular position in the layers of rock exp ...
of the Eureka Sound Group in the Sverdrup Basin in
Northwest Territories The Northwest Territories is a federal Provinces and territories of Canada, territory of Canada. At a land area of approximately and a 2021 census population of 41,070, it is the second-largest and the most populous of Provinces and territorie ...
and
Nunavut Nunavut is the largest and northernmost Provinces and territories of Canada#Territories, territory of Canada. It was separated officially from the Northwest Territories on April 1, 1999, via the ''Nunavut Act'' and the Nunavut Land Claims Agr ...
, Canada. The unit belonging to the Eureka Sound Group which crops out at
Ellesmere Island Ellesmere Island (; ) is Canada's northernmost and List of Canadian islands by area, third largest island, and the List of islands by area, tenth largest in the world. It comprises an area of , slightly smaller than Great Britain, and the total ...
preserves
fossils A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
dating back to the
Early Eocene In the geologic timescale the Ypresian is the oldest age (geology), age or lowest stage (stratigraphy), stratigraphic stage of the Eocene. It spans the time between , is preceded by the Thanetian Age (part of the Paleocene) and is followed by th ...
period Period may refer to: Common uses * Period (punctuation) * Era, a length or span of time *Menstruation, commonly referred to as a "period" Arts, entertainment, and media * Period (music), a concept in musical composition * Periodic sentence (o ...
, or
Wasatchian The Wasatchian North American Stage on the geologic timescale is the North American faunal stage according to the North American Land Mammal Ages chronology (NALMA), typically set from 55,400,000 to 50,300,000 years Before Present, BP lasting . I ...
in the NALMA classification.Margaret Formation
at
Fossilworks Fossilworks was a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was cr ...
.org
The Margaret Formation comprises
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
s, sandy
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
s, clay-rich sandstones, coal seams and clay-rich
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal i ...
seams and volcanic ash beds. The thickness of the formation, which overlies the Mount Moore Formation, reaches about . Radiometric dating of the formation provided ages of 52.6 ± 1.9 Ma (2010) and 53.7 ± 0.6 Ma (2017). The area where the formation was deposited in the Early Eocene experienced a much warmer climate than the High Arctic today, with mean annual temperatures ranging from and warmest month mean temperatures from . The deltaic to swamp environment of the Margaret Formation has provided a diverse fauna of various groups of mammals, birds (''
Presbyornis ''Presbyornis'' is an extinct genus of presbyornithid bird from North America during the Paleogene period, between the Late Paleocene and Early Eocene. History of discovery The fossil record of ''P. pervetus'' includes many complete skeletons ...
'' and ''
Gastornis ''Gastornis'' is an extinct genus of large, flightless birds that lived during the mid-Paleocene to mid-Eocene epochs of the Paleogene period. Most fossils have been found in Europe, and some species typically referred to the genus are known fr ...
''), reptiles (turtles, snakes, lizards and crocodiles) and fish.


Description

The Margaret Formation comprises
sandstone Sandstone is a Clastic rock#Sedimentary clastic rocks, clastic sedimentary rock composed mainly of grain size, sand-sized (0.0625 to 2 mm) silicate mineral, silicate grains, Cementation (geology), cemented together by another mineral. Sand ...
s, sandy
siltstone Siltstone, also known as aleurolite, is a clastic sedimentary rock that is composed mostly of silt. It is a form of mudrock with a low clay mineral content, which can be distinguished from shale by its lack of fissility. Although its permeabil ...
s, clay-rich sandstones, coal seams and clay-rich
coal Coal is a combustible black or brownish-black sedimentary rock, formed as rock strata called coal seams. Coal is mostly carbon with variable amounts of other Chemical element, elements, chiefly hydrogen, sulfur, oxygen, and nitrogen. Coal i ...
seams and volcanic ash beds. The thickness of the formation, which overlies the Mount Moore Formation, reaches about . The formation was probably deposited in a lush proximal delta front to delta plain environment, with abundant channels and coal
swamp A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
s.Bay Fiord
at
Fossilworks Fossilworks was a portal which provides query, download, and analysis tools to facilitate access to the Paleobiology Database, a large relational database assembled by hundreds of paleontologists from around the world. History Fossilworks was cr ...
.org


Dating

A volcanic ash layer containing crandallite in the middle of the formation was dated using U-Pb radiometric dating in 2017 to 53.7 ± 0.6 Ma.Sudermann, 2017 In 2010, ashes of the formation were dated to 52.6 ± 1.9 Ma.Reinhardt et al., 2010, p.2


Arctic climate of the Early Eocene

During the Early Eocene, the climate of much of northern North America was warm and wet, with mean annual temperatures (MAT) as high as , mean annual precipitation (MAP) of , mild frost-free winters (coldest month mean temperature >), and climatic conditions that supported extensive temperate forest ecosystems.West et al., 2020, p.1387 Ensemble estimates of mean annual temperatures for the high-latitude fossil localities in Arctic Canada ranged from , with the range of coldest month mean temperature from and warmest month mean temperatures from . Mean annual precipitation estimates for the Margaret Formation ranged between .West et al., 2020, p.1394 The mean summer precipitation has been estimated at and mean winter precipitation at .Schubert et al., 2012, p.525 The fossils and sedimentology indicate a lush, rain forest community on a coastal delta plain. Multiple palaeoclimate proxies, ranging from oxygen isotope analysis of vertebrate bones and teeth to palaeofloral analyses, estimate a mild temperate climate for the Eocene High Arctic, where winters remained at or just above freezing and summer temperatures extended to or higher. These temperatures are a far cry from today's High Arctic, where central Ellesmere Island experiences a mean annual temperature of , a warm month mean temperature of about and a cold month mean temperature of or colder.Stidham & Eberle, 2016, p.5 Despite the mild Eocene Arctic climate, the vertebrate fauna would have experienced months of total darkness and cooler temperatures during the winter. Recent isotopic work suggests that some mammals, including the hippo-like ''
Coryphodon ''Coryphodon'' (from Greek , "point", and , "tooth", meaning ''peaked tooth'', referring to "the development of the angles of the ridges into points n the molars") is an extinct genus of pantodonts of the family Coryphodontidae. ''Coryphodo ...
'', were year-round residents in the High Arctic. Given that ''
Gastornis ''Gastornis'' is an extinct genus of large, flightless birds that lived during the mid-Paleocene to mid-Eocene epochs of the Paleogene period. Most fossils have been found in Europe, and some species typically referred to the genus are known fr ...
'' was large (approaching ) and flightless, it likely also was a year-round resident of the Arctic. In contrast, the volant ''
Presbyornis ''Presbyornis'' is an extinct genus of presbyornithid bird from North America during the Paleogene period, between the Late Paleocene and Early Eocene. History of discovery The fossil record of ''P. pervetus'' includes many complete skeletons ...
'' might have been a seasonal migrant to the Arctic.


Fossil content

The formation has provided the following fossils:


Mammals

;Primates * Paromomyidae indet.West et al., 1977 ;Acreodi * '' Pachyaena sp.'' ;Carnivora * '' Miacis sp.'' * '' cf. Vulpavus sp.'' ;Dermoptera * '' Plagiomene sp.'' ;Eutheria * '' Palaeosinopa nunavutensis''Eberle & McKenna, 2002 * '' Anacodon sp.'' ;Ferae * '' Arcticanodon dawsonae''Rose et al., 2004 * '' Palaeonictis sp.'' * '' Viverravus sp.''McKenna, 1980 ;Glires * '' Microparamys bayi'' * '' Paramys hunti''Dawson, 2001 * '' Strathcona major'' * '' S. minor'' * ?Paramyinae indet. ;Hyaenodonta * '' Prolimnocyon sp.''Dawson, 1990 ;Leptictida * '' Prodiacodon sp.''Eberle, 2001 ;Multituberculata * '' Ectypodus arctos''Beard & Dawson, 2014 * Neoplagiaulacidae indet. ;Pantodonta * '' Coryphodon pisuqti''Dawson, 2012 ;Perissodactyla * '' Lambdotherium sp.'' * '' Thuliadanta mayri''Eberle, 2005 * Equidae indet.Alroy, 2002


Birds

;Anseriformes * '' Presbyornis cf. pervetus''Stidham & Eberle, 2016, p.3 ;Gastornithiformes * '' Gastornis sp.''


Reptiles

;Turtles * '' Geochelone sp.''Estes & Hutchison, 1980 * Cryptodira indet. * Emydinae indet. * Kinosterninae indet. * Trionychidae indet.Alroy, 2006 ;Crocodiles * '' Allognathosuchus sp.'' ;Lizards * Anguidae indet. * Varanidae indet. ;Snakes * Erycidae indet.


Amphibians

;Caudata * '' Piceoerpeton willwoodense''Gardner, 2012


Fish

;Teleostei * '' Esox sp.'' ;Amiiformes * '' Amia fragosa'' * '' Amia cf. uintaensis'' ;Lepisosteiformes * '' ?Lepisosteus sp.''


Correlations

The formation has been correlated with the
Allenby Formation The Allenby formation is a sedimentary rock formation in British Columbia which was deposited during the Ypresian stage of the Early Eocene. It consists of conglomerates, sandstones with interbedded shales and coal. The shales contain an ...
and Kamloops Group of
British Columbia British Columbia is the westernmost Provinces and territories of Canada, province of Canada. Situated in the Pacific Northwest between the Pacific Ocean and the Rocky Mountains, the province has a diverse geography, with rugged landscapes that ...
, the Chickaloon Formation of Alaska,West et al., 2020, p.1390 and with the Wishbone, Chuckanut and Iceberg Bay Formations.West et al., 2020, p.1391 The upper Margaret Formation also has been correlated with the Buchanan Lake Formation of the Eastern Arctic Archipelago.Eberle & Greenwood, 2012, p.6


See also

*
List of fossiliferous stratigraphic units in Northwest Territories This is a list of fossiliferous stratigraphic units in the Northwest Territories, Canada. References * {{Lists of fossiliferous stratigraphic units in Canada * Northwest Territories The Northwest Territories is a federal Provinces a ...
* List of fossiliferous stratigraphic units in Nunavut


References


Bibliography

;Geology and climate * * * * * ;Paleontology * * * * * * * * * * * * * * {{citation , last1=West , first1=R. M. , first2=M. R. , last2=Dawson , first3=J. H. , last3=Hutchison , year=1977 , title=Fossils from the Paleogene Eureka Sound Formation, N.W.T., Canada; occurrence, climatic and paleogeographic implications. In West, R. M. (ed), Paleontology and plate tectonics with special reference to the history of the Atlantic Ocean , journal= Milwaukee Public Museum Contributions in Biology and Geology , volume=2 , pages=77–93 Geologic formations of Nunavut Eocene Series of North America Paleogene Northwest Territories Paleogene Nunavut Ypresian Stage Wasatchian Sandstone formations of Canada Siltstone formations of Canada Coal formations Coal in Canada Tuff formations of Canada Deltaic deposits Paludal deposits Fossiliferous stratigraphic units of North America Paleontology in the Northwest Territories Paleontology in Nunavut Formations Geologic formations of the Northwest Territories