''Mandschurosaurus'' (meaning "lizard from
Manchuria
Manchuria is an exonym (derived from the endo demonym "Manchu") for a historical and geographic region in Northeast Asia encompassing the entirety of present-day Northeast China (Inner Manchuria) and parts of the Russian Far East ( Outer ...
") is an
extinct genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
of
hadrosaur
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which inclu ...
s based on material from the
Late Cretaceous
The Late Cretaceous (100.5–66 Ma) is the younger of two epochs into which the Cretaceous Period is divided in the geologic time scale. Rock strata from this epoch form the Upper Cretaceous Series. The Cretaceous is named after ''creta'', ...
of
China and possibly also the
Early Cretaceous
The Early Cretaceous (geochronological name) or the Lower Cretaceous ( chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145 Ma to 100.5 Ma.
Geology
Pr ...
of
Laos. It was the first
dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
genus
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial n ...
named from
China, and a mounted skeleton based on the holotype of ''M. amurensis'' is on display at the Central Geological and Prospecting Museum in
St. Petersburg
Saint Petersburg ( rus, links=no, Санкт-Петербург, a=Ru-Sankt Peterburg Leningrad Petrograd Piter.ogg, r=Sankt-Peterburg, p=ˈsankt pʲɪtʲɪrˈburk), formerly known as Petrograd (1914–1924) and later Leningrad (1924–1991), i ...
; however, much of the skeleton is
plaster
Plaster is a building material used for the protective or decorative coating of walls and ceilings and for moulding and casting decorative elements. In English, "plaster" usually means a material used for the interiors of buildings, while "r ...
.
Discovery and naming
The holotype of ''M. amurensis'' (IVP AS coll.), the only valid species within the genus, is based on a poorly preserved and incomplete skeleton collected by
Russia
Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eigh ...
n scientists in 1914 from the banks of the
Amur River
The Amur (russian: река́ Аму́р, ), or Heilong Jiang (, "Black Dragon River", ), is the world's tenth longest river, forming the border between the Russian Far East and Northeastern China ( Inner Manchuria). The Amur proper is long ...
,
China in a layer of the
Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the interv ...
-aged
Yuliangze Formation
The Yuliangze Formation, or Yuliangzi Formation, is a geological formation in Heilongjiang, China whose strata date back to the early-middle Maastrichtian. Dinosaur remains are among the fossils that have been recovered from the formation.Weishampe ...
and the remains represent a large hadrosaurid.
[ ussian/ref> The holotype material was initially referred to the genus ''"]Trachodon
''Trachodon'' (meaning "rough tooth") is a dubious genus of hadrosaurid dinosaur based on teeth from the Campanian-age Upper Cretaceous Judith River Formation of Montana, U.S.Leidy, J. (1856). "Notice of remains of extinct reptiles and fishes, ...
"'' (a ''nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
'') as ''T. amurense'' by Riabinin (1925),[Riabinin, A.N. (1925). A mounted skeleton of the gigantic reptile ''Trachodon amurense'', nov. sp. ''Izvest. Geol. Kom.'' 44(1):1-12. ussian/ref> but was later reassigned to a new genus by Riabinin (1930).][ The holotype of ''M. laosensis'', which is based mainly on an ilium and other fragmentary remains, comes from the ]Aptian
The Aptian is an age in the geologic timescale or a stage in the stratigraphic column. It is a subdivision of the Early or Lower Cretaceous Epoch or Series and encompasses the time from 121.4 ± 1.0 Ma to 113.0 ± 1.0 Ma (million years ag ...
-Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± ...
-aged Grès supérieurs Formation
The Grès supérieurs Formation (French for "Upper sandstone") is a geological formation in Laos whose strata date back to the Aptian to Albian stages of the Early Cretaceous. Dinosaur remains are among the fossils that have been recovered from t ...
in Laos and was named by Hoffet (1944).[J.-H. Hoffet. (1944). Description of the most characteristic bones belonging to Avipelvians from the Senonian of Lower Laos. ''Minutes of the Sessions of the Council for Scientific Research of Indochina'' 1944:179-186]
Description
''M. amurensis'' would likely have reached up to when fully grown.
Classification
There has been some debate regarding the validity of this genus. Brett-Surman (1979) first considered it a ''nomen dubium
In binomial nomenclature, a ''nomen dubium'' (Latin for "doubtful name", plural ''nomina dubia'') is a scientific name that is of unknown or doubtful application.
Zoology
In case of a ''nomen dubium'' it may be impossible to determine whether a s ...
'', though some later workers have continued to see it as a valid taxon (Chapman et Brett Surman, 1990,[Chapman R. et Brett-Surman M. K. (1990). Morphometric observations on hadrosaurid ornithopods. in Carpenter K. et Currie, P. J. (eds.), ''Dinosaur Systematics: Perspectives and Approaches'', Cambridge University Press, Cambridge 163-177.] for example). Most recently, Horner
Horner is an English and German surname that derives from the Middle English word for the occupation ''horner'', meaning horn-worker or horn-maker, or even horn-blower.
People
*Alison Horner (born 1966), British businesswoman
*Arthur Horner (disa ...
''et al.'' (2004) listed the type species as a ''nomen dubium'' in the second edition of ''The Dinosauria
''The Dinosauria'' is an extensive book on dinosaurs, compiled by David B. Weishampel, Peter Dodson, and Halszka Osmólska. It has been published in 2 editions, with the first edition published in 1990, consisting of material from 23 scientists.Be ...
''.[Horner J., Weishampel D. B., et Forster C. A. (2004). Chapter Twenty: Hadrosauridae. in ''The Dinosauria'' (2nd edition), Weishampel D. B., Dodson P., and Osmólska H., editors. University of California Press.
]
Over the years, three species have been placed within this genus: ''Mandschurosaurus amurensis'', ''M. mongoliensis'', and ''M. laosensis''. Brett-Surman (1979) considered ''M. mongoliensis'' a distinct genus, which he named ''Gilmoreosaurus
''Gilmoreosaurus'' (meaning "Charles Whitney Gilmore's lizard") is the name given to a genus of dinosaur from the Cretaceous of Asia. The type species is ''Gilmoreosaurus mongoliensis''. It is believed to be a hadrosaur or iguanodont from the Ire ...
'' and Horner ''et al.'' (2004) considered ''M. laosensis'' a ''nomen dubium''; This leaves only Riabinin's original species, ''M. amurensis'', as a possibly valid taxon.
Within Ornithopoda
Ornithopoda () is a clade of ornithischian dinosaurs, called ornithopods (), that started out as small, bipedal running grazers and grew in size and numbers until they became one of the most successful groups of herbivores in the Cretaceous wor ...
, ''Mandschurosaurus'' is most often placed within Hadrosauridae
Hadrosaurids (), or duck-billed dinosaurs, are members of the ornithischian family Hadrosauridae. This group is known as the duck-billed dinosaurs for the flat duck-bill appearance of the bones in their snouts. The ornithopod family, which incl ...
as a ''nomen dubium''.
Gallery
Mandschurosaurus Amurensis..2H1A0401WI.jpg, Reconstructed ''M. amurensis'' skeleton in outdated tripod-pose, VSEGEI Museum, St. Petersburg
Saint Petersburg ( rus, links=no, Санкт-Петербург, a=Ru-Sankt Peterburg Leningrad Petrograd Piter.ogg, r=Sankt-Peterburg, p=ˈsankt pʲɪtʲɪrˈburk), formerly known as Petrograd (1914–1924) and later Leningrad (1924–1991), i ...
See also
* Timeline of hadrosaur research
A timeline is a display of a list of events in chronological order. It is typically a graphic design showing a long bar labelled with dates paralleling it, and usually contemporaneous events.
Timelines can use any suitable scale representi ...
References
{{Taxonbar, from=Q1076898
Late Cretaceous dinosaurs of Asia
Hadrosaurs
Fossil taxa described in 1930
Ornithischian genera