Maleriraptor
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''Maleriraptor'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus of
herrerasauria Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithis ...
n
saurischia Saurischia ( , meaning "reptile-hipped" from the Greek ' () meaning 'lizard' and ' () meaning 'hip joint') is one of the two basic divisions of dinosaurs (the other being Ornithischia), classified by their hip structure. Saurischia and Ornithi ...
n dinosaurs from the
Late Triassic The Late Triassic is the third and final epoch (geology), epoch of the Triassic geologic time scale, Period in the geologic time scale, spanning the time between annum, Ma and Ma (million years ago). It is preceded by the Middle Triassic Epoch a ...
(
Norian The Norian is a division of the Triassic geological period, Period. It has the rank of an age (geology), age (geochronology) or stage (stratigraphy), stage (chronostratigraphy). It lasted from ~227.3 to Mya (unit), million years ago. It was prec ...
) Upper Maleri Formation of India. The genus contains a single species, ''M. kuttyi'', known from a partial skeleton.


Discovery and naming

The ''Maleriraptor''
holotype A holotype (Latin: ''holotypus'') is a single physical example (or illustration) of an organism used when the species (or lower-ranked taxon) was formally described. It is either the single such physical example (or illustration) or one of s ...
specimen, ISIR 282, was collected at some point prior to 1985 from outcrops of the Upper Maleri Formation of the Pranhita–Godavari Basin near
Annaram Annaram is a census town in Medak district of the Indian state of Telangana. Economy Its economy mainly depends on primary sectors of agriculture, Cattle breeding, cattle. It has Schneider Electric manufacturing industry warehouses, cycle asse ...
in south-central India. The specimen consists of the first , part of the second sacral rib, a vertebra representing a caudosacral element or the first in the series, an
anterior Standard anatomical terms of location are used to describe unambiguously the anatomy of humans and other animals. The terms, typically derived from Latin or Greek roots, describe something in its standard anatomical position. This position pro ...
(toward the front) caudal vertebra, the right , both ends of the right , and part of the left pubis. ISIR 282 is accessioned at the
Indian Statistical Institute The Indian Statistical Institute (ISI) is a public research university headquartered in Kolkata, India with centers in New Delhi, Bengaluru, Chennai and Tezpur. It was declared an Institute of National Importance by the Government of India und ...
. ISIR 282 was first described in a 2011 publication by Novas et al. reporting several new
dinosauriform Dinosauromorpha is a clade of avemetatarsalians (archosaurs closer to birds than to crocodilians) that includes the Dinosauria (dinosaurs) and some of their close relatives. It was originally defined to include dinosauriforms and lagerpetids, ...
specimens from the Upper Maleri Formation. They noted that exhibited a unique character combination that distinguished it from related species. However, the authors refrained from naming the specimen as it could not be compared with ''
Alwalkeria ''Alwalkeria'' (; "for Alick Walker") is an extinct genus of basal saurischian dinosaur from the Late Triassic Lower Maleri Formation of India. Discovery and naming ''Alwalkeria'' was originally named ''Walkeria maleriensis'' by Sankar Chatte ...
'', a similarly-aged saurischian from the slightly older
Lower Maleri Formation The Lower Maleri Formation is a sedimentary geological formation, rock formation found in Andhra Pradesh and Telangana, India. It is the lowermost member of the Pranhita–Godavari Basin. It is of late Carnian to early Norian age (Upper Triassi ...
. In 2025, Ezcurra and colleagues described ''Maleriraptor kuttyi'' as a new genus and species of early saurischian dinosaurs based on these fossil remains. The generic name, ''Maleriraptor'', combines a reference to the discovery of the holotype in the Upper Maleri Formation with the
Latin Latin ( or ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally spoken by the Latins (Italic tribe), Latins in Latium (now known as Lazio), the lower Tiber area aroun ...
word ''raptor'', meaning "thief". The
specific name Specific name may refer to: * in Database management systems, a system-assigned name that is unique within a particular database In taxonomy, either of these two meanings, each with its own set of rules: * Specific name (botany), the two-part (bino ...
, ''kuttyi'', honors Tharavat S. Kutty, the discoverer and initial describer of the holotype. Ezcurra et al. noted that a lack of overlapping material between ''Maleriraptor'' and ''Alwalkeria'' still precluded comparisons between the two and that the uncertain affinities of the latter did little to clarify their distinction. Regardless, they opted to name ''Maleriraptor kuttyi'', anticipating species-level differences between the Upper and Lower Maleri formations and hoping that future discoveries of more complete ''Alwalkeria'' specimens could allow for an improved understanding of the relationships of these taxa.


Classification

In their initial description of ISIR 282, Novas et al. (2011) reported it as belonging to an indeterminate dinosauriform; their
phylogenetic analysis In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical data ...
including the specimen recovered it in a
polytomy An internal node of a phylogenetic tree is described as a polytomy or multifurcation if (i) it is in a rooted tree and is linked to three or more child subtrees or (ii) it is in an unrooted tree and is attached to four or more branches. A tree ...
with ''
Silesaurus ''Silesaurus'' is a genus of silesaurid dinosauriform from the Late Triassic, of what is now Poland. Discovery and naming The Krasiejów claypit near Opole, Poland, was first discovered as a fossil locality in the 1980s after quarrying for ...
'',
Ornithischia Ornithischia () is an extinct clade of mainly herbivorous dinosaurs characterized by a pelvic structure superficially similar to that of birds. The name ''Ornithischia'', or "bird-hipped", reflects this similarity and is derived from the Greek ...
,
Theropoda Theropoda (; from ancient Greek iktionary:θηρίον, , (''therion'') "wild beast"; wiktionary:πούς, , wiktionary:ποδός, (''pous, podos'') "foot" is one of the three major groups (Clade, clades) of Dinosaur, dinosaurs, alon ...
,
Sauropodomorpha Sauropodomorpha ( ; from Greek, meaning "lizard-footed forms") is an extinct clade of long-necked, herbivorous, saurischian dinosaurs that includes the sauropods and their ancestral relatives. Sauropods generally grew to very large sizes, had lo ...
, and three herrerasaurs. They argued that the open and straight shaft of the pubis indicated a position more derived than
silesaurids Silesauridae is an extinct family of Triassic dinosauriforms. It is most commonly considered to be a clade of non-dinosaur dinosauriforms, and the sister group of dinosaurs. Some studies have instead suggested that most or all silesaurids compr ...
and that several anatomical characters excluded it from sauropodomorphs, but regarded its phylogenetic placement as "highly problematic". In 2021, Novas et al. reported ISIR 282 as belonging to an indeterminate herrerasaur without detailed discussion. To more accurately determine the relationships of ''Maleriraptor'', Ezcurra et al. (2025) scored it an updated version of the Ezcurra et al. (2023) phylogenetic dataset. This matrix placed ''Maleriraptor'' as an early-diverging member of the saurischian clade Herrerasauria. Scoring it in the Garcia et al. (2024) matrix yielded similar results, with ''Maleriraptor'' as a non-
herrerasaurid Herrerasauridae is a family of carnivorous dinosaurs, possibly basal to either theropods or even all of saurischians, or even their own branching from Dracohors, separate from Dinosauria altogether. They are among the oldest known dinosaurs, fi ...
herresaur. The results using the Ezcurra et al. (2023) dataset are displayed in the
cladogram A cladogram (from Greek language, Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an Phylogenetic tree, evolutionary tree because it does not s ...
below:


References

{{Taxonbar, from1=Q134384590, from2=Q134384592 Saurischia Dinosaur genera Norian dinosaurs Dinosaurs of India Fossil taxa described in 2025