Bioluminescence Bioluminescence is the emission of light during a chemiluminescence reaction by living organisms. Bioluminescence occurs in multifarious organisms ranging from marine vertebrates and invertebrates, as well as in some Fungus, fungi, microorgani ...

is the production of light by living organisms. The environment organizes this list of bioluminescent organisms, covering terrestrial, marine, and microorganisms.

Terrestrial animals

*Certain

arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...

s **Coleoptera (beetles) ***Lampyridae (

Fireflies The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...

) *** Phengodidae (Glowworm beetles) **** railroad worms *** Rhagophthalmidae *** Sinopyrophoridae ***Certain Elateridae (

Click beetle Elateridae or click beetles (or "typical click beetles" to distinguish them from the related families Cerophytidae and Eucnemidae, which are also capable of clicking) are a family (biology), family of beetles. Other names include elaters, sna ...

s) **** Pyrophorini ****'' Balgus'' ****'' Campyloxenus'' **Diptera (true flies) ***Certain keroplatid fungus gnats **Myriapoda ***Certain

centipede Centipedes (from Neo-Latin , "hundred", and Latin , "foot") are predatory arthropods belonging to the class Chilopoda (Ancient Greek , ''kheilos'', "lip", and Neo-Latin suffix , "foot", describing the forcipules) of the subphylum Myriapoda, ...

s ****'' Geophilus carpophagus'' ****''

Orphnaeus brevilabiatus ''Orphnaeus brevilabiatus'' is a species of centipede in the family Oryidae. Description Adult specimens are typically 60–65 mm long, yellowish-orange in colour and ca. 1 mm wide at the head. Females can have from 73 to 85 pairs of l ...

'' ***Certain

millipede Millipedes (originating from the Latin , "thousand", and , "foot") are a group of arthropods that are characterised by having two pairs of jointed legs on most body segments; they are known scientifically as the class Diplopoda, the name derive ...

s ****Genus '' Motyxia'' *Some

land snail A land snail is any of the numerous species of snail that live on land, as opposed to the sea snails and freshwater snails. ''Land snail'' is the common name for terrestrial molluscs, terrestrial gastropod mollusks that have gastropod shell, shel ...

s **'' Quantula striata'' **'' Quantula weinkauffiana'' **Genus '' Phuphania'' *

Annelid The annelids (), also known as the segmented worms, are animals that comprise the phylum Annelida (; ). The phylum contains over 22,000 extant species, including ragworms, earthworms, and leeches. The species exist in and have adapted to vario ...

s **'' Octochaetus multiporus'' **'' Microscolex phosphoreus'' **'' Pontodrilus litoralis''

Marine animals

Fish

Anglerfish The anglerfish are ray-finned fish in the order Lophiiformes (). Both the order's common name, common and scientific name comes from the characteristic mode of predation, in which a modified dorsal Fish fin#Ray-fins, fin ray acts as a Aggressiv ...

* Gulper eel *

Lanternfish Lanternfish (or myctophids, from the Greek language, Greek μυκτήρ ''myktḗr'', "nose" and ''ophis'', "serpent") are small mesopelagic fish of the large family (biology), family Myctophidae. One of two families in the order Myctophiformes, ...

Stomiiformes Stomiiformes is an order (biology), order of deep-sea ray-finned fishes of very diverse Morphology (biology), morphology. It includes, for example, Barbeled dragonfish, dragonfishes, lightfishes (Gonostomatidae and Phosichthyidae), loosejaws, ma ...

Marine hatchetfish Marine hatchetfishes or deep-sea hatchetfishes are small deep-sea mesopelagic ray-finned fish of the stomiiform subfamily Sternoptychinae. They should not be confused with the freshwater hatchetfishes, which are not particularly closely relate ...

Viperfish A viperfish is any species of marine fish in the genus ''Chauliodus''. Viperfish are mostly found in the mesopelagic zone and are characterized by long, needle-like teeth and hinged lower jaws. A typical viperfish grows to lengths of . Viperfis ...

** Black dragonfish * Midshipman fish * Pineconefish * Lanterneye fish *Some

Squaliformes The Squaliformes are an order of sharks that includes about 126 species in seven families. Members of the order have two dorsal fins, which usually possess spines, they usually have a sharp head, no anal fin or nictitating membrane, and fi ...

** Lantern sharks ** Kitefin sharks ** Velvet dogfish

Invertebrates

*A deep-sea species of carnivorous sponge ( Cladorhizidae) *Many

Cnidarian Cnidaria ( ) is a phylum under kingdom Animalia containing over 11,000 species of aquatic invertebrates found both in fresh water, freshwater and marine environments (predominantly the latter), including jellyfish, hydroid (zoology), hydroids, ...

s **

Sea pen Sea pens are marine cnidarians belonging to the superfamily Pennatuloidea, which are colony-forming benthic filter feeders within the order Scleralcyonacea. There are 14 families within the order and 35 extant genera, and it is estimated a ...

s *** '' Renilla reniformis'' **

Coral Corals are colonial marine invertebrates within the subphylum Anthozoa of the phylum Cnidaria. They typically form compact Colony (biology), colonies of many identical individual polyp (zoology), polyps. Coral species include the important Coral ...

**Certain

Jellyfish Jellyfish, also known as sea jellies or simply jellies, are the #Life cycle, medusa-phase of certain gelatinous members of the subphylum Medusozoa, which is a major part of the phylum Cnidaria. Jellyfish are mainly free-swimming marine animal ...

***''

Aequorea victoria ''Aequorea victoria'', also sometimes called the crystal jelly, is a bioluminescent hydrozoan jellyfish, or hydromedusa, that is found off the west coast of North America. The species is best known as the source of aequorin (a photoprotein), and ...

'' ***'' Atolla jellyfish'' ***'' Helmet jellyfish'' *Certain

Ctenophore Ctenophora (; : ctenophore ) is a phylum of marine invertebrates, commonly known as comb jellies, that inhabit sea waters worldwide. They are notable for the groups of cilia they use for swimming (commonly referred to as "combs"), and they ar ...

s (comb jellies) *Some

Tunicate Tunicates are marine invertebrates belonging to the subphylum Tunicata ( ). This grouping is part of the Chordata, a phylum which includes all animals with dorsal nerve cords and notochords (including vertebrates). The subphylum was at one time ...

s: **

Larvacean Larvaceans, copelates or appendicularians, class Appendicularia, are solitary, free-swimming tunicates found throughout the world's oceans. While larvaceans are filter feeders like most other tunicates, they keep their tadpole-like shape as adu ...

s **

Salp A salp (: salps, also known colloquially as “sea grape”) or salpa (: salpae or salpas) is a barrel-shaped, Plankton, planktonic tunicate in the family Salpidae. The salp moves by contracting its gelatinous body in order to pump water thro ...

s **

Ascidiacea Ascidiacea, commonly known as the ascidians or sea squirts, is a paraphyletic class in the subphylum Tunicata of sac-like marine invertebrate filter feeders. Ascidians are characterized by a tough outer test or "tunic" made of the polysacchari ...

Doliolida The Doliolida are an order of small marine chordates of the subphylum Tunicata. They are in the class Thaliacea, which also includes the salps and pyrosomes. The doliolid body is small, typically 1–2 mm long, and barrel-shaped; it fea ...

** Pyrosomes *Certain

echinoderm An echinoderm () is any animal of the phylum Echinodermata (), which includes starfish, brittle stars, sea urchins, sand dollars and sea cucumbers, as well as the sessile sea lilies or "stone lilies". While bilaterally symmetrical as ...

s (e.g.

Ophiurida The Ophiurida are an order of echinoderms within the class Ophiuroidea. It includes the vast majority of living brittle stars. Characteristics Ophiurida have bursae for respiration and excretion, and dorsal and ventral arm shields are present an ...

) ** '' Amphiura filiformis'' ** '' Ophiopsila aranea'' ** '' Ophiopsila californica'' ** '' Amphipholis squamata'' *Many

Crustacean Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

s: ** Seed shrimp (

Myodocopa Traditionally, the Myodocopa and Podocopa have been classified as subclasses within the class Ostracoda, although there is some question about how closely related the two groups actually are. The Myodocopa are defined by possession of a poorly c ...

) **

Copepod Copepods (; meaning 'oar-feet') are a group of small crustaceans found in nearly every freshwater and saltwater habitat (ecology), habitat. Some species are planktonic (living in the water column), some are benthos, benthic (living on the sedimen ...

s ** Lophogastrids ( Gnathophausia) **

Amphipods Amphipoda () is an order (biology), order of malacostracan crustaceans with no carapace and generally with laterally compressed bodies. Amphipods () range in size from and are mostly detritivores or scavengers. There are more than 10,700 amphip ...

Krill Krill ''(Euphausiids)'' (: krill) are small and exclusively marine crustaceans of the order (biology), order Euphausiacea, found in all of the world's oceans. The name "krill" comes from the Norwegian language, Norwegian word ', meaning "small ...

Decapods The Decapoda or decapods, from Ancient Greek δεκάς (''dekás''), meaning "ten", and πούς (''poús''), meaning "foot", is a large order (biology), order of crustaceans within the class Malacostraca, and includes crabs, lobsters, crayfis ...

(

shrimp A shrimp (: shrimp (American English, US) or shrimps (British English, UK)) is a crustacean with an elongated body and a primarily Aquatic locomotion, swimming mode of locomotion – typically Decapods belonging to the Caridea or Dendrobranchi ...

and

prawn Prawn is a common name for small aquatic crustaceans with an exoskeleton An exoskeleton () . is a skeleton that is on the exterior of an animal in the form of hardened integument, which both supports the body's shape and protects the intern ...

) ***Genus '' Heterocarpus'' *Two species of Chaetognaths (arrow worms) **'' Caecosagitta macrocephala'' **'' Eukrohnia fowleri'' *

a **Genus '' Tomopteris'' **Genus '' Swima'' **Certain polynoids *

Mollusca Mollusca is a phylum of protostome, protostomic invertebrate animals, whose members are known as molluscs or mollusks (). Around 76,000 extant taxon, extant species of molluscs are recognized, making it the second-largest animal phylum ...

**Certain

Bivalves Bivalvia () or bivalves, in previous centuries referred to as the Lamellibranchiata and Pelecypoda, is a class of aquatic molluscs (marine and freshwater) that have laterally compressed soft bodies enclosed by a calcified exoskeleton consis ...

(clams) ***'' Plocamopherus maderae'' ***'' Pholas dactylus'' **Certain

Nudibranch Nudibranchs () are a group of soft-bodied marine gastropod molluscs, belonging to the order Nudibranchia, that shed their shells after their larval stage. They are noted for their often extraordinary colours and striking forms, and they have b ...

s (sea slugs) **Certain

sea snail Sea snails are slow-moving marine (ocean), marine gastropod Mollusca, molluscs, usually with visible external shells, such as whelk or abalone. They share the Taxonomic classification, taxonomic class Gastropoda with slugs, which are distinguishe ...

s ***'' Hinea brasiliana'' **Many

Cephalopods A cephalopod is any member of the molluscan Taxonomic rank, class Cephalopoda (Greek language, Greek plural , ; "head-feet") such as a squid, octopus, cuttlefish, or nautilus. These exclusively marine animals are characterized by bilateral symm ...

***Certain

Octopus An octopus (: octopuses or octopodes) is a soft-bodied, eight-limbed mollusc of the order Octopoda (, ). The order consists of some 300 species and is grouped within the class Cephalopoda with squids, cuttlefish, and nautiloids. Like oth ...

es **** Bolitaenidae **** Stauroteuthis *** Vampire squid ***

Sepiolida Bobtail squid (order Sepiolida) are a group of cephalopods closely related to cuttlefish. Bobtail squid tend to have a rounder mantle than cuttlefish and have no cuttlebone. They have eight suckered arms and two tentacles and are generally quit ...

***Many Teuthida (squid) **** Cranchiidae ***** Colossal Squid **** Mastigoteuthidae **** Histioteuthidae **** Enoploteuthoidea ***** Firefly squid

Freshwater animals

*'' Latia'', a genus of four species of freshwater snail

Fungi

Bacteria

*''

Photorhabdus luminescens ''Photorhabdus luminescens'' (previously called ''Xenorhabdus luminescens'') is a Gammaproteobacterium of the family Morganellaceae, and is a lethal pathogen of insects. It lives in the gut of an entomopathogenic nematode of the family Hetero ...

'' *Certain species of the family Vibrionaceae (e.g. '' Vibrio fischeri'', '' Vibrio harveyi'', '' Photobacterium phosphoreum'') *Certain species of the family Shewanellaceae, (e.g. '' Shewanella hanedai'' and '' Shewanella woodyi'')

Other microorganisms

Protists A protist ( ) or protoctist is any Eukaryote, eukaryotic organism that is not an animal, Embryophyte, land plant, or fungus. Protists do not form a Clade, natural group, or clade, but are a Paraphyly, paraphyletic grouping of all descendants o ...

**Certain

Dinoflagellate The Dinoflagellates (), also called Dinophytes, are a monophyletic group of single-celled eukaryotes constituting the phylum Dinoflagellata and are usually considered protists. Dinoflagellates are mostly marine plankton, but they are also commo ...

s (e.g. '' Noctiluca scintillans'', '' Pyrodinium bahamense'', '' Pyrocystis fusiformis'' and '' Lingulodinium polyedra''

References

{{Use dmy dates, date=May 2017

Bioluminescent Bioluminescence is the emission of light during a chemiluminescence reaction by living organisms. Bioluminescence occurs in multifarious organisms ranging from marine vertebrates and invertebrates, as well as in some fungi, microorganisms inc ...

List of