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''Laurelia novae-zelandiae'', also called pukatea, is a large
evergreen In botany, an evergreen is a plant which has foliage that remains green and functional through more than one growing season. This also pertains to plants that retain their foliage only in warm climates, and contrasts with deciduous plants, whic ...
tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondary growth, plants that are ...
,
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
to the forests of
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
. Pukatea has 'toothed' leaves and produces small flowers. It is a species in the
Atherospermataceae The Atherospermataceae, commonly known as the southern sassafrases, are a family of broadleaf evergreen trees and shrubs. The family includes 14 species in seven genera. The atherosperms are today mostly distributed in the Southern Hemisphere, wi ...
(formerly
Monimiaceae The Monimiaceae is a family of flowering plants in the magnoliid order Laurales.Peter F. Stevens (2001 onwards). "Monimiaceae" At: Angiosperm Phylogeny Website. At: Botanical Databases At: Missouri Botanical Garden Website. (see ''External links' ...
) family, typical representative of
laurel forest Laurel forest, also called laurisilva or laurissilva, is a type of subtropical forest found in areas with high humidity and relatively stable, mild temperatures. The forest is characterized by broadleaf tree species with evergreen, glossy and elo ...
ecoregion An ecoregion (ecological region) or ecozone (ecological zone) is an ecologically and geographically defined area that is smaller than a bioregion, which in turn is smaller than a biogeographic realm. Ecoregions cover relatively large areas of ...
.


Distribution

Pukatea is generally found in lowland forest and grows throughout the
North Island The North Island, also officially named Te Ika-a-Māui, is one of the two main islands of New Zealand, separated from the larger but much less populous South Island by the Cook Strait. The island's area is , making it the world's 14th-larges ...
of New Zealand, and the northern third of the
South Island The South Island, also officially named , is the larger of the two major islands of New Zealand in surface area, the other being the smaller but more populous North Island. It is bordered to the north by Cook Strait, to the west by the Tasman ...
, usually where moisture is plentiful, such as in damp, lowland forests, gullies, and on the edges of streams. Pukatea grows well in poorly drained soil, but is equally at home on hillsides. It requires a temperate to warm subtropical climate, but also a frost-free environment with only very slight winter frosts not below 24.8 °F (-4 °C), and with high summer heat. Growth is best on well-drained, slightly acidic
soil Soil, also commonly referred to as earth or dirt, is a mixture of organic matter, minerals, gases, liquids, and organisms that together support life. Some scientific definitions distinguish ''dirt'' from ''soil'' by restricting the former ...
s rich in organic matter.


Description

Pukatea grows slowly to a height of 130 feet (40 m), usually 115 feet (35 m), and is the only New Zealand native tree developing large plank buttresses to support the tree's growth in swamp or shallow-soil areas. ''L. novae-zelandiae'' has specialized respiratory root structures called
pneumatophore Aerial roots are roots above the ground. They are almost always adventitious. They are found in diverse plant species, including epiphytes such as orchids (''Orchidaceae''), tropical coastal swamp trees such as mangroves, banyan figs (''Fic ...
s in certain waterlogged ground or mud. These fragrant trees are characteristic of the lower strata of the tropical rainforest. The tree has thin bark and a pale brownish-grey trunk that becomes attractively buttressed at the base. Its dark green, glossy, elliptical leaves are 2-3 inches (5-7 cm) long and have coarsely serrated edges and paler undersides. The odorous opposite
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
have oil cells in the
parenchyma Parenchyma () is the bulk of functional substance in an animal organ or structure such as a tumour. In zoology it is the name for the tissue that fills the interior of flatworms. Etymology The term ''parenchyma'' is New Latin from the word � ...
, and brochidodromous venations. Juvenile leaves and stems can be difficult to distinguish from another native tree
hutu The Hutu (), also known as the Abahutu, are a Bantu ethnic or social group which is native to the African Great Lakes region. They mainly live in Rwanda, Burundi and the eastern Democratic Republic of the Congo, where they form one of the p ...
, to which it is only distantly related. Pukatea is a mostly
dioecious Dioecy (; ; adj. dioecious , ) is a characteristic of a species, meaning that it has distinct individual organisms (unisexual) that produce male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproducti ...
species, male and female flowers are on separate individuals. Some specimens had a ratio as high as 100 male flowers to every female or hermaphrodite one. These results suggest that the species is not truly dioecious. The female and hermaphrodite flowers are very similar. The flowers are tiny, inconspicuous, and in small
raceme A raceme ( or ) or racemoid is an unbranched, indeterminate type of inflorescence bearing flowers having short floral stalks along the shoots that bear the flowers. The oldest flowers grow close to the base and new flowers are produced as the s ...
s. The star-shaped flowers are whitish with yellow glistening glands and scarlet anther flaps. The glands at the base of the stamens in ''L. novae-zelandiae'' secrete nectar that accumulates at the base of the flower. This attracts a large number of bees, blowflies, small flightless animals, and
Bombyliidae The Bombyliidae are a family of flies, commonly known as bee flies. Adults generally feed on nectar and pollen, some being important pollinators. Larvae are mostly parasitoids of other insects. Overview The Bombyliidae are a large family of fl ...
to the flowers. The nectar is visible as a colourless liquid which has a glistening appearance to the outer faces of the glands. Nectar runs down from the glands and accumulates on the floor of the flower as a result of continuing secretion. The fruit are little pear-shaped capsules which contain numerous
achene An achene (; ), also sometimes called akene and occasionally achenium or achenocarp, is a type of simple dry fruit produced by many species of flowering plants. Achenes are monocarpellate (formed from one carpel) and indehiscent (they do not o ...
s attached to fine feathery strands which aid their dispersal by the wind. Often, only a few seeds are viable.


Uses

Historically, the light, but tough timber of pukatea has been used for boat building. The timber was used by
Māori Māori or Maori can refer to: Relating to the Māori people * Māori people of New Zealand, or members of that group * Māori language, the language of the Māori people of New Zealand * Māori culture * Cook Islanders, the Māori people of the Co ...
to create figureheads for canoes. It is rather soft, but very strong. It yields a pale hardwood that is difficult to split and will dent upon impact rather than break. Its wood is pale-yellowish, with
growth ring Dendrochronology (or tree-ring dating) is the scientific method of dating tree rings (also called growth rings) to the exact year they were formed. As well as dating them, this can give data for dendroclimatology, the study of climate and atmo ...
s, and is homogeneous and fine-textured. An extract from the bark containing the
alkaloid Alkaloids are a class of basic, naturally occurring organic compounds that contain at least one nitrogen atom. This group also includes some related compounds with neutral and even weakly acidic properties. Some synthetic compounds of simila ...
pukateine is used in traditional Māori herbal medicine as an
analgesic An analgesic drug, also called simply an analgesic (American English), analgaesic (British English), pain reliever, or painkiller, is any member of the group of drugs used to achieve relief from pain (that is, analgesia or pain management). It ...
.Nigel Perry. 'Plant extracts', Te Ara - the Encyclopedia of New Zealand, updated 2-Mar-09 - use as an analgesic
/ref> The pulp of the cambium was boiled in water and the resulting liquid used for treating
tuberculosis Tuberculosis (TB) is an infectious disease usually caused by '' Mycobacterium tuberculosis'' (MTB) bacteria. Tuberculosis generally affects the lungs, but it can also affect other parts of the body. Most infections show no symptoms, ...
.


References


Further reading


"STUDIES ON THE MONIMIACEAE II.The Floral morphology of ''Laurelia novae-zelandiae'' A. Cunn. (SUBFAMILY ATHEROSPERMOIDEAE)" by F.B. Sampson, in ''New Zealand Journal of Botany'', September 1969 Item 20: pages 214-240 - pdf format.
{{Taxonbar, from=Q1005204 Atherospermataceae Trees of New Zealand Flora of the North Island Flora of the South Island Plants described in 1838 Plants used in traditional Māori medicine