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''Larix gmelinii'', the Dahurian larch or Gmelin larch, is a species of
larch Larches are deciduous conifers in the genus ''Larix'', of the family Pinaceae (subfamily Laricoideae). Growing from tall, they are native to much of the cooler temperate northern hemisphere, on lowlands in the north and high on mountains furt ...
native to eastern
Siberia Siberia ( ; rus, Сибирь, r=Sibir', p=sʲɪˈbʲirʲ, a=Ru-Сибирь.ogg) is an extensive geographical region, constituting all of North Asia, from the Ural Mountains in the west to the Pacific Ocean in the east. It has been a part of ...
and adjacent northeastern
Mongolia Mongolia; Mongolian script: , , ; literal translation, lit. "Mongol Nation" or "State of Mongolia" () is a landlocked country in East Asia, bordered by Russia Mongolia–Russia border, to the north and China China–Mongolia border, to the s ...
, northeastern China (
Heilongjiang Heilongjiang () formerly romanized as Heilungkiang, is a province in northeast China. The standard one-character abbreviation for the province is (). It was formerly romanized as "Heilungkiang". It is the northernmost and easternmost province ...
),
South Korea South Korea, officially the Republic of Korea (ROK), is a country in East Asia, constituting the southern part of the Korean Peninsula and sharing a land border with North Korea. Its western border is formed by the Yellow Sea, while its ea ...
and
North Korea North Korea, officially the Democratic People's Republic of Korea (DPRK), is a country in East Asia. It constitutes the northern half of the Korea, Korean Peninsula and shares borders with China and Russia to the north, at the Yalu River, Y ...
.


Description

''Larix gmelinii'' is a medium-sized
deciduous In the fields of horticulture and Botany, the term ''deciduous'' () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, afte ...
coniferous Conifers are a group of cone-bearing seed plants, a subset of gymnosperms. Scientifically, they make up the division Pinophyta (), also known as Coniferophyta () or Coniferae. The division contains a single extant class, Pinopsida. All extan ...
tree In botany, a tree is a perennial plant with an elongated Plant stem, stem, or trunk (botany), trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, including only woody plants with secondar ...
reaching 10–30 m tall, rarely 40 m, with a trunk up to 1 m diameter. The crown is broad conic; both the main branches and the side branches are level, the side branches only rarely drooping. The shoots are dimorphic, with growth divided into long shoots (typically 5–30 cm long) and bearing several
bud In botany, a bud is an undeveloped or embryonic shoot and normally occurs in the axil of a leaf or at the tip of a stem. Once formed, a bud may remain for some time in a dormant condition, or it may form a shoot immediately. Buds may be spec ...
s, and short shoots only 1–2 mm long with only a single bud. The
leaves A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, ste ...
are needle-like, light green, 2–3 cm long; they turn bright yellow to orange before they fall in the autumn, leaving the variably downy reddish-brown shoots bare until the next spring. The
cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines con ...
are erect, ovoid, 1–2 cm (rarely 2.5 cm) long, with 15–25 moderately reflexed seed scales; they are green (rarely purple) when immature, turning brown and opening to release the
seed A seed is an Plant embryogenesis, embryonic plant enclosed in a testa (botany), protective outer covering, along with a food reserve. The formation of the seed is a part of the process of reproduction in seed plants, the spermatophytes, includ ...
s when mature, 3–5 months after pollination. The old cones commonly remain on the tree for many years, turning dull grey-black.


Taxonomy and systematics

The scientific name honours
Johann Georg Gmelin Johann Georg Gmelin (8 August 1709 – 20 May 1755) was a German naturalist, botanist and geographer. Early life and education Gmelin was born in Tübingen, the son of a professor at the University of Tübingen. He was a gifted child and began ...
. Due to the species' variability, it has acquired numerous
synonyms A synonym is a word, morpheme, or phrase that means exactly or nearly the same as another word, morpheme, or phrase in a given language. For example, in the English language, the words ''begin'', ''start'', ''commence'', and ''initiate'' are a ...
in the botanical literature, including ''L. cajanderi, L. dahurica, L. kamtschatica, L. komarovii, L. kurilensis, L. lubarskii, L. ochotensis''. Dahurian larch intergrades with the closely related Siberian larch '' L. sibirica'' of central and western Siberia where their ranges meet along the Yenisei Valley; the resulting
hybrid Hybrid may refer to: Science * Hybrid (biology), an offspring resulting from cross-breeding ** Hybrid grape, grape varieties produced by cross-breeding two ''Vitis'' species ** Hybridity, the property of a hybrid plant which is a union of two dif ...
is named ''Larix'' × ''czekanowskii''.


Varieties

,
Plants of the World Online Plants of the World Online (POWO) is an online database published by the Royal Botanic Gardens, Kew. It was launched in March 2017 with the ultimate aim being "to enable users to access information on all the world's known seed-bearing plants by ...
accepts four varieties: *''Larix gmelinii'' var. ''gmelinii''. Most of the species' range, from the
Yenisei The Yenisey (russian: Енисе́й, ''Yeniséy''; mn, Горлог мөрөн, ''Gorlog mörön''; Buryat: Горлог мүрэн, ''Gorlog müren''; Tuvan: Улуг-Хем, ''Uluğ-Hem''; Khakas: Ким суғ, ''Kim suğ''; Ket: Ӄук ...
Valley east to
Kamchatka The Kamchatka Peninsula (russian: полуостров Камчатка, Poluostrov Kamchatka, ) is a peninsula in the Russian Far East, with an area of about . The Pacific Ocean and the Sea of Okhotsk make up the peninsula's eastern and west ...
. *''Larix gmelinii'' var. ''japonica''. The
Kuril Islands The Kuril Islands or Kurile Islands (; rus, Кури́льские острова́, r=Kuril'skiye ostrova, p=kʊˈrʲilʲskʲɪjə ɐstrɐˈva; Japanese: or ) are a volcanic archipelago currently administered as part of Sakhalin Oblast in the ...
and
Sakhalin Sakhalin ( rus, Сахали́н, r=Sakhalín, p=səxɐˈlʲin; ja, 樺太 ''Karafuto''; zh, c=, p=Kùyèdǎo, s=库页岛, t=庫頁島; Manchu: ᠰᠠᡥᠠᠯᡳᠶᠠᠨ, ''Sahaliyan''; Orok: Бугата на̄, ''Bugata nā''; Nivkh: ...
. * ''Larix gmelinii'' var. ''olgensis''. North Korea,
Heilongjiang Heilongjiang () formerly romanized as Heilungkiang, is a province in northeast China. The standard one-character abbreviation for the province is (). It was formerly romanized as "Heilungkiang". It is the northernmost and easternmost province ...
, and the
Sikhote-Alin The Sikhote-Alin (russian: Сихотэ́-Али́нь, , , ) is a mountain range in Primorsky and Khabarovsk Krais, Russia, extending about to the northeast of the Russian Pacific seaport of Vladivostok. The highest summits are Tordoki Yani ...
mountains of
Primorsky Krai Primorsky Krai (russian: Приморский край, r=Primorsky kray, p=prʲɪˈmorskʲɪj kraj), informally known as Primorye (, ), is a federal subject (a krai) of Russia, located in the Far East region of the country and is a part of the ...
,
Russia Russia (, , ), or the Russian Federation, is a transcontinental country spanning Eastern Europe and Northern Asia. It is the largest country in the world, with its internationally recognised territory covering , and encompassing one-eight ...
. Sometimes treated as a distinct species ''Larix olgensis''. * ''Larix gmelinii'' var. ''principis-rupprechtii'' (Prince Rupprecht's larch), syn. ''Larix principis-rupprechtii''. It occurs in a disjunct region in the
Wutai Shan Mount Wutai, also known by its Chinese name Wutaishan and as is a sacred Buddhist site at the headwaters of the Qingshui in Shanxi Province, China. Its central area is surrounded by a cluster of flat-topped peaks roughly corresponding to the c ...
mountains west of
Beijing } Beijing ( ; ; ), alternatively romanized as Peking ( ), is the capital of the People's Republic of China. It is the center of power and development of the country. Beijing is the world's most populous national capital city, with over 2 ...
, separated from typical ''L. gmelinii'' by about 1000 km, and differs in having larger (2–4 cm) cones with more scales.


Ecology and uses

''Larix gmelinii'' forms enormous forests in the eastern Siberian taiga, growing at 50–1,200 m altitude on both boggy and well-drained soils, including on the shallow soils above
permafrost Permafrost is ground that continuously remains below 0 °C (32 °F) for two or more years, located on land or under the ocean. Most common in the Northern Hemisphere, around 15% of the Northern Hemisphere or 11% of the global surface ...
. It is unique in two respects, being the northernmost tree in the world, reaching 73° 04' 32" N 102° 00' 00" E as creeping forms of the trees in the tundra of the Taymyr Peninsula, 72° 55' 07" N 106° 08' E as comparatively vertical growing single trees, 72° 31' N 105° 03' E at Lukunsky grove and 72° 28' N 102° 15' E at Ary-Mas as the subtundra sparse forests which are situated both in the Taymyr Peninsula, Khatanga river basin, and also the most cold-hardy tree in the world, tolerating temperatures below -70 °C in the
Oymyakon Oymyakon, ; sah, Өймөкөөн, ''Öymököön'', is a rural locality (a '' selo'') in Oymyakonsky District of the Sakha Republic, Russia, located in the Yana-Oymyakon Highlands, along the Indigirka River, northwest of Tomtor on the Kol ...
Verkhoyansk Verkhoyansk ( rus, Верхоянск, p=vʲɪrxɐˈjansk; sah, Верхоянскай, ''Verkhoyanskay'') is a town in Verkhoyansky District of the Sakha Republic, Russia, located on the Yana River in the Arctic Circle, from Batagay, the ...
region of
Yakutia Sakha, officially the Republic of Sakha (Yakutia),, is the largest republic of Russia, located in the Russian Far East, along the Arctic Ocean, with a population of roughly 1 million. Sakha comprises half of the area of its governing Far E ...
. One tree in Yakutia was recorded as being 919 years old. Dahurian larch is occasionally grown in botanical gardens in Europe and North America; it is not an easy tree to grow in areas with mild winters as it is adapted to a long period of winter rest; the warm winter weather in Britain can tempt it into leaf as early as the start of January, with the tender young leaves then being killed by the next frost. In its native region, daily minimum temperatures above freezing do not occur until late May or June, with no further frost until the brief summer is over. ''Larix gmelinii'' cone scales are used as food by the
caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder Symph ...
s of the
tortrix moth The Tortricidae are a family of moths, commonly known as tortrix moths or leafroller moths, in the order Lepidoptera. This large family has over 11,000 species described, and is the sole member of the superfamily Tortricoidea, although the genu ...
'' Cydia illutana''.


Gallery

File:Larix gmelinii female cone.jpg, Young female cone and emerging needles File:Larix gmelinii bark.jpg, Bark File:Larix gmelinii short shoots.jpg, Short shoots, just after bud break File:Larix gmelinii var. olgensis (OIga Bay Larch) (37535368894).jpg, Shoots in fall File:Larix gmelinii var. olgensis (OIga Bay Larch) (26470319579).jpg, ''Larix gmelinii'' var. ''olgensis'' in fall,
Arnold Arboretum of Harvard University The Arnold Arboretum of Harvard University is a botanical research institution and free public park, located in the Jamaica Plain and Roslindale neighborhoods of Boston, Massachusetts. Established in 1872, it is the oldest public arboretum in No ...
, accession #365-97*A. File:Larix gmelinii old cone.jpg, Old female cone


References


External links


''Larix gmelinii'' images at the Arnold Arboretum of Harvard University Plant Image Database"''Larix dahurica'' var. ''Principis Rupprechtii'' China."
''Library Featured Images,'' Arnold Arboretum of Harvard University website, 30 October 2014. Accessed 7 May 2020.
Gymnosperm Database: ''Larix gmelinii''images (Larix gmelinii var. gmelinii)
{{Authority control gmelinii Flora of Northeast Asia Trees of Siberia Trees of Russia Trees of Asia Trees of China Trees of Korea Trees of continental subarctic climate Least concern plants Deciduous conifers