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''Lansium parasiticum'', commonly known as langsat (), lanzones (), or longkong in English; duku in Indonesian or dokong in
Terengganu Malay Terengganu Malay ( ms, Bahasa Melayu Terengganu; Terengganu Malay: ) is a Malayic language spoken in the Malaysian state of Terengganu all the way southward to coastal Pahang and northeast Johor. It is the native language of Terengganu Malays a ...
, is a species of tree in the
Mahogany Mahogany is a straight- grained, reddish-brown timber of three tropical hardwood species of the genus '' Swietenia'', indigenous to the AmericasBridgewater, Samuel (2012). ''A Natural History of Belize: Inside the Maya Forest''. Austin: U ...
family with commercially cultivated edible fruits. The species is native to Southeast Asia.


Description

The tree is average sized, reaching in height and in diameter. 30 years old trees grown from seed and planted at 8 × 8 meter spacing can have a height of 10 meters and diameter of 25 cm. The trunk grows in an irregular manner, with its
buttress root Buttress roots also known as plank roots are large, wide roots on all sides of a shallowly rooted tree. Typically, they are found in nutrient-poor tropical forest soils that may not be very deep. They prevent the tree from falling over (hen ...
s showing above ground. The tree's bark is a greyish colour, with light and dark spots. Its resin is thick and milk coloured. The pinnately compound leaves are odd numbered, with thin hair, and 6 to 9 buds at intervals. The buds are long and elliptical, approximately by in size. The upper edge shines, and the leaves themselves have pointed bases and tips. The stems of the buds measure . The flowers are located in
inflorescence An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are forme ...
s that grow and hang from large branches or the trunk; the bunches may number up to 5 in one place. They are often branched at their base, measure in size, and have short fur. The flowers are small, with short stems, and have two
genders Gender is the range of characteristics pertaining to femininity and masculinity and differentiating between them. Depending on the context, this may include sex-based social structures (i.e. gender roles) and gender identity. Most cultures us ...
. The sheathe is shaped like a five lobed cup and is coloured a greenish-yellow. The corona is egg-shaped and hard, measuring by . There is one
stamen The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10 Morphology and terminology A stamen typically consists of a stalk called the fila ...
, measuring in length. The top of the stamen is round. The
pistil Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pistils ...
is short and thick. The fruit can be elliptical, ovoid or round, measuring by in size. Fruits look much like small potatoes and are borne in clusters similar to grapes. The larger fruits are on the variety known as duku. It is covered by thin, yellow hair giving a slightly fuzzy aspect. The skin thickness varies with the varieties, from to approximately . With the skin removed, the fruit resembles peeled
garlic Garlic (''Allium sativum'') is a species of bulbous flowering plant in the genus '' Allium''. Its close relatives include the onion, shallot, leek, chive, Welsh onion and Chinese onion. It is native to South Asia, Central Asia and northeas ...
in appearance, with half a dozen white, translucent lobes, some of which contain a flat, bitter tasting seed. The seeds are covered with a thick, clear-white
aril An aril (pronounced ), also called an arillus, is a specialized outgrowth from a seed that partly or completely covers the seed. An arillode or false aril is sometimes distinguished: whereas an aril grows from the attachment point of the see ...
that tastes sweet and sour. The taste has been likened to a combination of grape and grapefruit and is considered excellent by most. The sweet juicy flesh contains
sucrose Sucrose, a disaccharide, is a sugar composed of glucose and fructose subunits. It is produced naturally in plants and is the main constituent of white sugar. It has the molecular formula . For human consumption, sucrose is extracted and refine ...
,
fructose Fructose, or fruit sugar, is a ketonic simple sugar found in many plants, where it is often bonded to glucose to form the disaccharide sucrose. It is one of the three dietary monosaccharides, along with glucose and galactose, that are absorb ...
, and
glucose Glucose is a simple sugar with the molecular formula . Glucose is overall the most abundant monosaccharide, a subcategory of carbohydrates. Glucose is mainly made by plants and most algae during photosynthesis from water and carbon dioxide, u ...
. For consumption, cultivars with small or undeveloped seeds and thick aril are preferred.


Cultivars

There are numerous
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s of ''L. parasiticum''. Overall, there are two main groups of cultivars, those named ''duku'' and those named ''langsat''. There are also mixed duku-langsat varieties. Those called duku generally have a large crown, thick with bright green leaves, with short bunches of few fruit. The individual fruit are large, generally round, and have somewhat thick skin that does not release sap when cooked. The seeds are small, with thick flesh, a sweet scent, and a sweet or sour taste. Meanwhile, the variant commonly known as langsat generally has thinner trees, with a less dense crown consisting of dark green leaves and stiff branches. The bunches are longer, and each bunch holds between 15 and 25 large, egg-shaped fruit. The skin is thin and releases a white sap when cooked. The flesh is watery and tastes sweet and sour. ''Lansium domesticum'' var. ''aquaeum'' is distinguished by its hairy leaves, as well as the tightly packed dark yellow fruit on its bunches. The fruit tends to be small, with thin skin and little sap; the skin is difficult to remove. To be eaten, the fruit is bitten and the flesh sucked through the hole created, or rubbed until the skin breaks and the seeds are retrieved. In Indonesia the fruit has several names, including ''kokosan'', ''pisitan'', ''pijetan'', and ''bijitan''. The seeds are relatively large, with thin, sour flesh.


Reproduction

The seeds of ''L. parasiticum'' are
polyembryonic Polyembryony is the phenomenon of two or more embryos developing from a single fertilized egg. Due to the embryos resulting from the same egg, the embryos are identical to one another, but are genetically diverse from the parents. The genetic differ ...
, the multiple embryos resulting from
apomixis In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cuttin ...
. ''Lansium parasiticum'' is traditionally reproduced by spreading seedlings, either cultivated or collected from below the tree. It has been said that new seedlings require 20 to 25 years to bear fruit, with the possibility of the quality being inferior. However other sources quote 12 years to first production from seed and no variations. Production often varies from year to year, and depends to some extent on having a dry period to induce flowering. One example of ten trees in Costa Rica about twenty-five years old produced during five years the following weights of salable fruits: 2008: 50 kilos, 2009: 2000 kilos, 2010: 1000 kilos, 2011: 100 kilos, 2012: 1500 kilos. Experiments in the Philippines with grafting where two trees are planted close to each other and then grafted when one to two meters tall to leave twin root systems on a single main trunk have resulted in earlier and less erratic fruit production. Another common method is by
air layering Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants. Natural layering typically occurs when a branch touches ...
. Although the process requires up to several months, the new rooted tree produced is itself ready to bear fruit within two years. Trees cultivated with this method have a high death rate, and the growths are less resilient. The third common way to reproduce ''L. parasiticum'' is with
grafting Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...
. This results in the new trees having the same genetic characteristics as their parent, and being ready to bear fruit within 5 to 6 years. The offspring are relatively stronger than transplanted shoots.


Ecology

''Lansium parasiticum'' grows well in mixed agroforests. The plant, especially the ''duku'' variant, prefers damp, shaded areas. It can be grown in the same agroforest as
durian The durian (, ) is the edible fruit of several tree species belonging to the genus ''Durio''. There are 30 recognised ''Durio'' species, at least nine of which produce edible fruit. '' Durio zibethinus'', native to Borneo and Sumatra, is the on ...
,
petai ''Parkia speciosa'', the bitter bean, twisted cluster bean or stink bean, is a plant of the genus ''Parkia'' in the family Fabaceae. It bears long, flat edible beans with bright green seeds the size and shape of plump almonds which have a ra ...
, and
jengkol ''Archidendron pauciflorum'', commonly known as djenkol, jengkol or jering is a species of flowering tree in the pea family, Fabaceae. It is native to Southeast Asia, where the seeds are a popular dish.Lim, T. K. "Archidendron jiringa." Edible Me ...
, as well as wood-producing trees. ''Lansium parasiticum'' is grown from low grounds up to heights of above sea level, in areas with an average rainfall of a year. The plant can grow and blossom in
latosol Latosols, also known as tropical red earth, are soils found under tropical rainforests which have a relatively high content of iron and aluminium oxides. They are typically classified as oxisols ( USDA soil taxonomy) or ferralsols ( World Referenc ...
, yellow
podzol In soil science, podzols are the typical soils of coniferous or boreal forests and also the typical soils of eucalypt forests and heathlands in southern Australia. In Western Europe, podzols develop on heathland, which is often a construct of ...
, and
alluvium Alluvium (from Latin ''alluvius'', from ''alluere'' 'to wash against') is loose clay, silt, sand, or gravel that has been deposited by running water in a stream bed, on a floodplain, in an alluvial fan or beach, or in similar settings. ...
. The plant prefers slightly acidic soil with good drainage and rich in mulch. The ''langsat'' variant is hardier, and can weather dry seasons with a little shade and water. The plant cannot handle floods. ''Lansium parasiticum'' generally bears fruit once a year. This period can vary between areas, but blooming is generally after the beginning of the rainy season and fruit production some four months later.


Distribution

''Lansium parasiticum'' is native to the Malesian
phytochorion A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both r ...
of Southeast Asia, from
Peninsular Thailand Southern Thailand, Southern Siam or Tambralinga is a southernmost cultural region of Thailand, separated from Central Thailand region by the Kra Isthmus. Geography Southern Thailand is on the Malay Peninsula, with an area of around , bounded t ...
and
Peninsular Malaysia Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
, to
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
,
Sumatra Sumatra is one of the Sunda Islands of western Indonesia. It is the largest island that is fully within Indonesian territory, as well as the sixth-largest island in the world at 473,481 km2 (182,812 mi.2), not including adjacent i ...
,
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and e ...
, the
Philippines The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
(
Luzon Luzon (; ) is the largest and most populous island in the Philippines. Located in the northern portion of the Philippines archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, as ...
,
Camiguin Camiguin, officially the Province of Camiguin ( ceb, Probinsya sa Camiguin; tl, Lalawigan ng Camiguin; Kamigin: ''Probinsya ta Kamigin''), is an island province in the Philippines located in the Bohol Sea, about off the northern coast of ...
,
Basilan Basilan, officially the Province of Basilan ( cbk, Provincia de Basilan; yka, Wilayah Basilanin; tsg, Wilaya' sin Basilan; fil, Lalawigan ng Basilan), is an island province of the Philippines located primarily in the Bangsamoro Autonomous Reg ...
, and
Mindanao Mindanao ( ) ( Jawi: مينداناو) is the second-largest island in the Philippines, after Luzon, and seventh-most populous island in the world. Located in the southern region of the archipelago, the island is part of an island group of ...
),
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu ...
, the
Moluccas The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ...
, and
Western New Guinea Western New Guinea, also known as Papua, Indonesian New Guinea, or Indonesian Papua, is the western half of the Melanesian island of New Guinea which is administered by Indonesia. Since the island is alternatively named as Papua, the region ...
. It has also been introduced to
Laos Laos (, ''Lāo'' )), officially the Lao People's Democratic Republic ( Lao: ສາທາລະນະລັດ ປະຊາທິປະໄຕ ປະຊາຊົນລາວ, French: République démocratique populaire lao), is a socialist s ...
,
Cambodia Cambodia (; also Kampuchea ; km, កម្ពុជា, UNGEGN: ), officially the Kingdom of Cambodia, is a country located in the southern portion of the Indochinese Peninsula in Southeast Asia, spanning an area of , bordered by Thailand ...
,
Myanmar Myanmar, ; UK pronunciations: US pronunciations incl. . Note: Wikipedia's IPA conventions require indicating /r/ even in British English although only some British English speakers pronounce r at the end of syllables. As John Wells explai ...
,
Micronesia Micronesia (, ) is a subregion of Oceania, consisting of about 2,000 small islands in the western Pacific Ocean. It has a close shared cultural history with three other island regions: the Philippines to the west, Polynesia to the east, ...
,
Hawaii Hawaii ( ; haw, Hawaii or ) is a state in the Western United States, located in the Pacific Ocean about from the U.S. mainland. It is the only U.S. state outside North America, the only state that is an archipelago, and the only stat ...
,
Sri Lanka Sri Lanka (, ; si, ශ්‍රී ලංකා, Śrī Laṅkā, translit-std=ISO (); ta, இலங்கை, Ilaṅkai, translit-std=ISO ()), formerly known as Ceylon and officially the Democratic Socialist Republic of Sri Lanka, is an ...
,
India India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
, the
Seychelles Seychelles (, ; ), officially the Republic of Seychelles (french: link=no, République des Seychelles; Creole: ''La Repiblik Sesel''), is an archipelagic state consisting of 115 islands in the Indian Ocean. Its capital and largest city, ...
,
Trinidad and Tobago Trinidad and Tobago (, ), officially the Republic of Trinidad and Tobago, is the southernmost island country in the Caribbean. Consisting of the main islands Trinidad and Tobago, and numerous much smaller islands, it is situated south of ...
, and
Suriname Suriname (; srn, Sranankondre or ), officially the Republic of Suriname ( nl, Republiek Suriname , srn, Ripolik fu Sranan), is a country on the northeastern Atlantic coast of South America. It is bordered by the Atlantic Ocean to the nor ...
, among others. The tree is cultivated commercially in Thailand ( th, ลางสาด, langsat), Cambodia, Vietnam, India, Malaysia and the Philippines ( tl, lansones; ceb, buwahan, lansones;
Philippine Spanish The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no), * bik, Republika kan Filipinas * ceb, Republika sa Pilipinas * cbk, República de Filipinas * hil, Republ ...
: ''lanzones''). It grows well in the wetter areas (120 inches/3 meters or more annual rainfall) of Costa Rica, where it is still very rare, having been introduced decades ago by the United Fruit Company. A major hindrance to its acceptance seems to be that it is very slow in bearing, said to take 12 years or more from seed. However, air layering from mature trees, as well as grafting, are said to work well and produce much faster. The name Duku is reserved for the larger-sized varieties of Langsat, near the size of golf balls, claimed sweeter and with less sap in the peel. A variety called Dokong exported to mainland Malaysia from Thailand (this variety is called 'Longkang' th, ลองกอง in Thailand) grows tighter in the clusters, giving it a faceted shape, and is preferred by many over the standard Langsat.


Uses

''Lansium parasiticum'' is cultivated mainly for its fruit, which can be eaten raw. The fruit can also be bottled in syrup. The wood is hard, thick, heavy, and resilient, allowing it to be used in the construction of rural houses. Some parts of the plant are used in making traditional medicine. The bitter seeds can be pounded and mixed with water to make a
deworming Deworming (sometimes known as worming, drenching or dehelmintization) is the giving of an anthelmintic drug (a wormer, dewormer, or drench) to a human or animals to rid them of helminths parasites, such as roundworm, flukes and tapeworm. Pur ...
and ulcer medication. The bark is used to treat
dysentery Dysentery (UK pronunciation: , US: ), historically known as the bloody flux, is a type of gastroenteritis that results in bloody diarrhea. Other symptoms may include fever, abdominal pain, and a feeling of incomplete defecation. Complications ...
and
malaria Malaria is a mosquito-borne infectious disease that affects humans and other animals. Malaria causes symptoms that typically include fever, tiredness, vomiting, and headaches. In severe cases, it can cause jaundice, seizures, coma, or death. ...
; the powdered bark can also be used to treat scorpion stings. The fruit's skin is used to treat diarrhea, and in the Philippines the dried skin is burned as a mosquito repellent. The skin, especially of the ''langsat'' variety, can be dried and burned as incense. The greatest producers of ''Lansium parasiticum'' are Indonesia, Thailand, the Philippines and Malaysia. The production is mostly for internal consumption, although some is exported to
Singapore Singapore (), officially the Republic of Singapore, is a sovereign island country and city-state in maritime Southeast Asia. It lies about one degree of latitude () north of the equator, off the southern tip of the Malay Peninsula, bor ...
,
Hong Kong Hong Kong ( (US) or (UK); , ), officially the Hong Kong Special Administrative Region of the People's Republic of China (abbr. Hong Kong SAR or HKSAR), is a List of cities in China, city and Special administrative regions of China, special ...
and
Kuwait Kuwait (; ar, الكويت ', or ), officially the State of Kuwait ( ar, دولة الكويت '), is a country in Western Asia. It is situated in the northern edge of Eastern Arabia at the tip of the Persian Gulf, bordering Iraq to Iraq–Ku ...
.


See also

*''
Dysoxylum parasiticum ''Dysoxylum parasiticum'', known as yellow mahogany, is a species of rainforest trees in the family Meliaceae. The specific epithet ' is from the Latin meaning "parasitic", referring to the idea (now known to be incorrect) that the flowers are pa ...
'' *
Lychee Lychee (US: ; UK: ; ''Litchi chinensis''; ) is a Monotypic taxon, monotypic taxon and the sole member in the genus ''Litchi'' in the Sapindus, soapberry family, ''Sapindaceae''. It is a tropical tree native to Southeast and Southwest China (t ...
*
Longan ''Dimocarpus longan'', commonly known as the longan () and dragon's eye, is a tropical tree species that produces edible fruit. It is one of the better-known tropical members of the soapberry family Sapindaceae, to which the lychee and rambu ...


References


External links

* *
''Lansium domesticum''
at Tree Functional Attributes and Ecological Database (World Agro Forestry) {{Taxonbar, from=Q860731 Meliaceae Tropical fruit Fruits originating in Asia Trees of Indo-China Trees of Malesia Fruit trees