List Of Subgroups Of The Order Coleoptera
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This article classifies the subgroups of the order Coleoptera (
beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s) down to the level of
families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictability, structure, and safety as ...
, following the system in "Family-group names in Coleoptera (Insecta)", Bouchard, et al. (2011), with corrections and additions from 2020, with common names from bugguide.net. *Order
Coleoptera Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
**Suborder †
Protocoleoptera The Protocoleoptera are a Paraphyly, paraphyletic group of extinct beetles, containing the earliest and most primitive lineages of beetles. They represented the dominant group of beetles during the Permian, but were largely replaced by modern bee ...
***Superfamily † Tshekardocoleoidea Rohdendorf, 1944 ****Family † Tshekardocoleidae Rohdendorf, 1944 ****Family † Labradorocoleidae Ponomarenko, 1969 ****Family † Oborocoleidae Kukalová, 1969 ***Superfamily † Permocupedoidea Martynov, 1933 ****Family † Permocupedidae Martynov, 1933 ****Family † Taldycupedidae Rohdendorf, 1961 ***Superfamily † Permosynoidea Tillyard, 1924 ****Family † Ademosynidae Ponomarenko, 1968 ****Family † Permosynidae Tillyard, 1924 **Suborder
Archostemata The Archostemata are the smallest suborder of beetles, consisting of 50 living species in five families and over 200 described fossil species. They are an ancient lineage with a number of primitive characteristics. Antenna (biology), Antennae may ...
***Superfamily Cupedoidea Laporte, 1836 ****Family Crowsoniellidae Iablokoff-Khnzorian, 1983 ****Family Cupedidae Laporte, 1836 ****Family Micromalthidae Barber, 1913 ****Family Ommatidae Sharp and Muir, 1912 ****Family Jurodidae Ponomarenko, 1985 ****Family † Triadocupedidae Ponomarenko, 1966 ****Family † Magnocoleidae Hong, 1998 ****Family † Obrieniidae Zherikhin and Gratshev, 1994 ***Superfamily † Asiocoleoidea Rohdendorf, 1961 ****Family † Asiocoleidae Rohdendorf, 1961 ****Family † Tricoleidae Ponomarenko, 1969 ***Superfamily † Rhombocoleoidea Rohdendorf, 1961 ****Family † Rhombocoleidae Rohdendorf, 1961 ***Superfamily † Schizocoleoidea Rohdendorf, 1961 (formerly Schizophoroidea Ponomarenko, 1968) ****Family † Phoroschizidae Bouchard and Bousquet, 2020 (formerly Schizophoridae Ponomarenko, 1968) ****Family † Catiniidae Ponomarenko, 1968 ****Family † Schizocoleidae Rohdendorf, 1961 **Suborder
Myxophaga Myxophaga is the second-smallest suborder of the Coleoptera after Archostemata, consisting of roughly 65 species of small to minute beetles in four families. The members of this suborder are aquatic and semiaquatic, and feed on algae. Descript ...
***Superfamily Lepiceroidea Hinton, 1936 (1882) ****Family Lepiceridae Hinton, 1936 (1882) ***Superfamily Sphaeriusoidea Erichson, 1845 ****Family Torridincolidae Steffan, 1964 ****Family Hydroscaphidae LeConte, 1874 ****Family Sphaeriusidae Erichson, 1845 **Suborder
Adephaga The Adephaga (from Greek ἀδηφάγος, ''adephagos'', "gluttonous") are a suborder of beetles, and with more than 40,000 recorded species in 10 families, the second-largest of the four beetle suborders. Members of this suborder are collecti ...
****Family † Tritarsusidae Hong, 2002 (formerly Tritarsidae Hong, 2002) ****Family
Gyrinidae The whirligig beetles are water beetles, comprising the family Gyrinidae, that usually swim on the surface of the water if undisturbed, though they swim underwater when threatened. They get their common name from their habit of swimming rapidly ...
Latreille, 1810 (whirligig beetles) ****Family Trachypachidae Thomson, 1857 (false ground beetles) ****Family
Rhysodidae Rhysodinae is a subfamily (sometimes called wrinkled bark beetles) in the family Carabidae. There are 19 genera and at least 380 described species in Rhysodinae. The group of genera making up Rhysodinae had been treated as the family Rhysodidae i ...
Laporte, 1840 ****Family
Carabidae Ground beetles are a large, cosmopolitan family of beetles, the Carabidae, with more than 40,000 species worldwide, around 2,000 of which are found in North America and 2,700 in Europe. As of 2015, it is one of the 10 most species-rich animal ...
Latreille, 1802 (ground beetles) ****Family
Haliplidae The Haliplidae are a family (biology), family of water beetles that swim using an alternating motion of the legs. They are therefore clumsy in water (compared e.g. with the Dytiscidae or Hydrophilidae), and prefer to get around by crawling. The f ...
Aubé, 1836 (crawling water beetles) ****Family † Triaplidae Ponomarenko, 1977 ****Family † Colymbotethidae Ponomarenko, 1994 ****Family † Parahygrobiidae Ponomarenko, 1977 ****Family †
Coptoclavidae Coptoclavidae is an extinct family of aquatic beetles in the suborder Adephaga. The Coptoclavidae lived from the Late Triassic to the Early Cretaceous. Coptoclavidae is a member of the adephagan clade Adephaga, Dytiscoidea, which contains living ...
Ponomarenko, 1961 ****Family † Liadytidae Ponomarenko, 1977 ****Family Meruidae Spangler and Steiner, 2005 ****Family Noteridae Thomson, 1860 (burrowing water beetles) ****Family Amphizoidae LeConte, 1853 ****Family Aspidytidae Ribera, Beutel, Balke and Vogler, 2002 ****Family Hygrobiidae Régimbart, 1879 (1837) ****Family
Dytiscidae The Dytiscidae, from the Ancient Greek word δυτικός (''dystikos''), meaning "able to dive", are the predaceous diving beetles, a family of water beetles. They occur in virtually any freshwater habitat around the world, but a few species l ...
Leach, 1815 (predaceous diving beetles) **Suborder
Polyphaga Polyphaga is the largest and most diverse suborder of beetles. It comprises 144 families in 16 superfamilies, and displays an enormous variety of specialization and adaptation, with over 350,000 described species, or approximately 90% of the b ...
***Infraorder
Staphyliniformia Staphyliniformia is a large infraorder of beetles. It contains over 70,000 described species from all regions of the world. Most species occur in moist habitats - various kinds of rotting plant debris, fungi, dung, carrion, many live in fresh wat ...
****Superfamily
Histeroidea Histeroidea is a Taxonomic rank, superfamily of beetles in the infraorder Staphyliniformia. Characteristics Characteristic to Histeroidea are an accessory posterior ridge (locking device) behind the hind margin and presence of Comstock-Needham s ...
Gyllenhal, 1808 *****Family
Histeridae Histeridae is a family (biology), family of beetles commonly known as clown beetles or hister beetles. There are more than 410 genera and 4,800 described species in Histeridae worldwide, with more than 500 species in North America. They can be id ...
Gyllenhal, 1808 *****Family
Sphaeritidae ''Sphaerites'' is a genus of beetles, the only genus in the family Sphaeritidae, sometimes called the false clown beetles. There are five known species, which are widespread in temperate areas of the Northern Hemisphere, found in forested or upla ...
Shuckard, 1839 *****Family
Synteliidae ''Syntelia'' is a genus of beetles. It is the only genus in the family Synteliidae. There are seven known species, which are native to high-elevation regions in southern North America from central Mexico to Guatemala, and in eastern Asia, from I ...
Lewis, 1882 ****Superfamily
Hydrophiloidea Hydrophiloidea, known as water scavenger beetles, is a Taxonomic rank, superfamily of beetles. Until recently it included only a single family, the Hydrophilidae (water scavenger beetles), but several of the subfamilies have been removed and rais ...
Latreille, 1802 *****Family Epimetopidae Zaitzev, 1908 *****Family Georissidae Laporte, 1840 *****Family Helophoridae Leach, 1815 *****Family Hydrochidae Thomson, 1859 *****Family
Hydrophilidae Hydrophilidae, also known colloquially as water scavenger beetles, is a family of beetles. Aquatic hydrophilids are notable for their long maxillary palps, which are longer than their antennae. Several of the former subfamilies of Hydrophilidae ...
Latreille, 1802 (water scavenger beetles) *****Family Spercheidae Erichson, 1837 ****Superfamily Staphylinoidea Latreille, 1802 *****Family
Hydraenidae Hydraenidae is a family of very small aquatic animal, aquatic beetles, sometimes called "Minute moss beetles", with a worldwide distribution. They are around 0.8 to 3.3 mm in length. The adults store air on the underside of the body as wel ...
Mulsant, 1844 *****Family Ptiliidae Erichson, 1845 *****Family Agyrtidae Thomson, 1859 *****Family
Leiodidae Leiodidae is a family of beetles with around 3800 described species found worldwide. Members of this family are commonly called round fungus beetles due to the globular shape of many species, although some are more elongated in shape. They are g ...
Fleming, 1821 *****Family
Silphidae Silphidae is a family of beetles that are known commonly as large carrion beetles, carrion beetles or burying beetles. There are two family (biology), subfamilies: Silphinae and Nicrophorinae. Members of Nicrophorinae are sometimes known as bury ...
Latreille, 1806 *****Family
Staphylinidae The rove beetles are a family (biology), family (Staphylinidae) of beetles, primarily distinguished by their short elytra (wing covers) that typically leave more than half of their abdominal segments exposed. With over 66,000 species in thousand ...
Latreille, 1802 (rove beetles) ***Infraorder
Scarabaeiformia Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Some of its constituent families are undergo ...
****Superfamily
Scarabaeoidea Scarabaeoidea is a superfamily of beetles, the only subgroup of the infraorder Scarabaeiformia. Around 35,000 species are placed in this superfamily and some 200 new species are described each year. Some of its constituent families are undergo ...
Latreille, 1802 *****Family Pleocomidae LeConte, 1861 (rain beetles) *****Family Bolboceratidae Mulsant, 1842 *****Family
Geotrupidae Geotrupidae (from Greek γῆ ''(gē)'', earth, and τρῡπητής ''(trȳpētēs)'', borer) is a family of beetles in the order Coleoptera. They are commonly called earth-boring dung beetles or dor beetles. Most excavate burrows in which t ...
Latreille, 1802 (earth-boring scarab beetles) *****Family Belohinidae Paulian, 1959 *****Family
Passalidae Passalidae is a family of beetles known variously as "bessbugs", "bess beetles", "betsy beetles", "betsy bugs", or "horned passalus beetles". Nearly all of the 500-odd species are tropical; species found in North America are notable for their siz ...
Leach, 1815 (bess beetles) *****Family
Trogidae Trogidae, sometimes called hide beetles, is a family of beetles with a distinctive warty or bumpy appearance. Found worldwide, the family includes about 300 species contained in four or five genera. Trogids range in length from 2 to 20 mm. Thei ...
MacLeay, 1819 (hide beetles) *****Family
Glaresidae ''Glaresis'' is a genus of beetles, sometimes called "Enigmatic scarab beetles", in its own family, Glaresidae. It is closely related to, and was formerly included in, the family Scarabaeidae. Although its members occur in arid and sandy areas wo ...
Prudhomme de Borre, 1886 *****Family Diphyllostomatidae Holloway, 1972 *****Family
Lucanidae Stag beetles comprise the family Lucanidae. It has about 1,200 species of beetles in four subfamilies.Smith, A.B.T. (2006). A review of the family-group names for the superfamily Scarabaeoidea (Coleoptera) with corrections to nomenclature and a c ...
Latreille, 1804 (stag beetles) *****Family
Ochodaeidae Ochodaeidae, also known as the sand-loving scarab beetles, is a small family of scarabaeiform beetles occurring in many parts of the world. These beetles are small, ranging from . Their bodies are elongate and convex, with black and brown colo ...
Streubel, 1846 (sand-loving scarab beetles) *****Family
Hybosoridae Hybosoridae, sometimes known as the scavenger scarab beetles, is a family of Scarabaeiformia, scarabaeiform beetles. The >600 species in 78 extant genera occur widely in the tropics, but little is known of their biology. Hybosorids are small, 5 ...
Erichson, 1847 *****Family Glaphyridae MacLeay, 1819 (bumble bee scarab beetles) *****Family
Scarabaeidae The family Scarabaeidae, as currently defined, consists of over 35,000 species of beetles worldwide; they are often called scarabs or scarab beetles. The classification of this family has undergone significant change. Several groups formerly tre ...
Latreille, 1802 (scarab beetles) *****Family † Coprinisphaeridae Genise, 2004 (
ichnotaxon An ichnotaxon (plural ichnotaxa) is "a taxon based on the fossilized work of an organism", i.e. the non-human equivalent of an artifact. ''Ichnotaxon'' comes from the Ancient Greek (''íchnos'') meaning "track" and English , itself derived from ...
) *****Family † Pallichnidae Genise, 2004 (ichnotaxon) ***Infraorder
Elateriformia Elateriformia is an infraorder of polyphagan beetles. The two largest families in this group are buprestids, of which there are around 15,000 described species, and click beetles, of which there are around 10,000 described species. The infraor ...
****Superfamily
Scirtoidea Scirtoidea is a superfamily of beetles. It is traditionally considered to consist of four families: Clambidae, Decliniidae, Eucinetidae and Scirtidae. However, genetic studies have suggested that Clambidae and Eucinetidae belong to a separate su ...
Fleming, 1821 *****Family
Decliniidae Decliniidae is a family of beetles belonging to Scirtoidea. It contains the single genus ''Declinia'' with two species, ''D. relicta'' and ''D. versicolor'', found in the Russian Far East The Russian Far East ( rus, Дальний Вос ...
Nikitsky, Lawrence, Kirejtshuk and Gratshev, 1994 *****Family
Eucinetidae Eucinetidae is a family (biology), family of beetles, notable for their large coxal plates that cover much of the first ventrite of the abdomen, sometimes called plate-thigh beetles. The family is small for beetles, with about 50 species in 11 ...
Lacordaire, 1857 *****Family
Clambidae Clambidae is a family of beetles. They are known commonly as the minute beetlesMajka, C. G., & Langor, D. (2009)Clambidae (Coleoptera) of Atlantic Canada.''Journal of the Acadian Entomological Society'' 5(7), 32-40. or the fringe-winged beetles.< ...
Fischer von Waldheim, 1821 *****Family
Scirtidae Scirtidae is a family of beetles (Coleoptera). These beetles are commonly referred to as marsh beetles, as the larvae are typically associated with stagnant water, but can be found in flowing water. Adults prefer decomposing plant material near ...
Fleming, 1821 *****Family † Elodophthalmidae Kirejtshuk and Azar, 2008 *****Family † Mesocinetidae Kirejtshuk and Ponomarenko, 2010 ****Superfamily
Dascilloidea Dascilloidea is a superfamily of polyphagan beetles, comprising two families Family (from ) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). It forms the basis fo ...
Guérin-Méneville, 1843 (1834) *****Family Dascillidae Guérin-Méneville, 1843 (1834) *****Family Rhipiceridae Latreille, 1834 ****Superfamily Buprestoidea Leach, 1815 *****Family Schizopodidae LeConte, 1859 *****Family
Buprestidae Buprestidae is a family (biology), family of beetles known as jewel beetles or metallic wood-boring beetles because of their glossy Iridescence, iridescent colors. Larvae of this family are known as flatheaded borers. The family is among the lar ...
Leach, 1815 ****Superfamily
Byrrhoidea Byrrhoidea is a superfamily of beetles belonging to Elateriformia that includes several families which are either aquatic or associated with a semi-aquatic habitat. Other than the superfamily Hydrophiloidea, most of the remaining Polyphagan beetle ...
Latreille, 1804 *****Family Byrrhidae Latreille, 1804 *****Family Protelmidae Jeannel, 1950 *****Family Elmidae Curtis, 1830 *****Family Dryopidae Billberg, 1820 (1817) *****Family Lutrochidae Kasap and Crowson, 1975 *****Family
Limnichidae Limnichidae, commonly called minute marsh-loving beetles, is a family of beetles belonging to Byrrhoidea. There are at least 30 genera and 350 described species in Limnichidae. They are found worldwide, with the greatest diversity in tropical regi ...
Erichson, 1846 *****Family Heteroceridae MacLeay, 1825 *****Family Psephenidae Lacordaire, 1854 *****Family Cneoglossidae Champion, 1897 *****Family Ptilodactylidae Laporte, 1836 *****Family Podabrocephalidae Pic, 1930 *****Family Chelonariidae Blanchard, 1845 *****Family Eulichadidae Crowson, 1973 *****Family Callirhipidae Emden, 1924 ****Superfamily
Elateroidea The Elateroidea are a large superfamily of beetles. It contains the familiar click beetles, fireflies, and soldier beetles and their relatives. It consists of about 25,000 species. Description Elateroidea is a morphologically diverse group ...
Leach, 1815 *****Family Artematopodidae Lacordaire, 1857 *****Family Brachypsectridae Horn, 1881 *****Family
Cantharidae The soldier beetles (Cantharidae) are relatively soft-bodied, straight-sided beetles. They are Cosmopolitan distribution, cosmopolitan in distribution. One of the first described species has a color pattern reminiscent of the Red coat (Bri ...
Imhoff, 1856 (1815) (soldier beetles) *****Family
Cerophytidae The Cerophytidae are a family of beetles belonging to Elateroidea. Larvae are associated with rotting wood, on which they are presumed to feed.Costa, Cleide, Vanin, Sergio A., Lawrence, John F. and Ide, Sergio. "4.4. Cerophytidae Latreille, 1834 ...
Latreille, 1834 *****Family
Elateridae Elateridae or click beetles (or "typical click beetles" to distinguish them from the related families Cerophytidae and Eucnemidae, which are also capable of clicking) are a family of beetles. Other names include elaters, snapping beetles, sp ...
Leach, 1815 (click beetles; includes former Drilidae, Omalisidae, and Plastoceridae) *****Family
Eucnemidae Eucnemidae, or false click beetles, are a family of elateroid beetles based on the type genus '' Eucnemis''; they include about 1700 species, distributed worldwide. Description Closely related to the family Elateridae, specimens of Eucnemida ...
Eschscholtz, 1829 *****Family
Lampyridae The Lampyridae are a family of elateroid beetles with more than 2,000 described species, many of which are light-emitting. They are soft-bodied beetles commonly called fireflies, lightning bugs, or glowworms for their conspicuous production ...
Rafinesque, 1815 (fireflies) *****Family Lycidae Laporte, 1836 *****Family Omethidae LeConte, 1861 (includes former Telegeusidae) *****Family Phengodidae LeConte, 1861 *****Family Rhagophthalmidae Olivier, 1907 *****Family Sinopyrophoridae Bi, 2018 *****Family Throscidae Laporte, 1840 nomen protectum *****Family † Berendtimiridae Winkler, 1987 *****Family † Praelateriidae Dolin, 1973 ****Superfamily
Rhinorhipoidea ''Rhinorhipus'' is a genus of beetles that contains a single species, ''Rhinorhipus tamborinensis'' from southern Queensland, Australia. It is the sole member of the family Rhinorhipidae and superfamily Rhinorhipoidea. It is an isolated lineage n ...
Lawrence, 1988 *****Family Rhinorhipidae Lawrence, 1988 ***Infraorder Bostrichiformia ****Superfamily Derodontoidea LeConte, 1861 *****Family
Derodontidae Derodontidae is a family of beetles, in its own superfamily, Derodontoidea, sometimes known as the tooth-necked fungus beetles. Beetles of this family are small, between 2 and 6 mm in length, typically with spiny margins on their pronotum ...
LeConte, 1861 *****Family Nosodendridae Erichson, 1846 *****Family Jacobsoniidae Heller, 1926 ****Superfamily
Bostrichoidea Bostrichoidea is a Taxonomic rank, superfamily of beetles. It is the type superfamily of the infraorder Bostrichiformia. Description The Bostrichoidea are united by the following features: modified cryptonephridism, the structure of the aedea ...
Latreille, 1802 *****Family
Dermestidae Dermestidae are a family of Coleoptera that are commonly referred to as skin beetles or carpet beetles. Other common names include larder beetles, hide or leather beetles, and khapra beetles. There are over 1,800 species described. Dermestids ha ...
Latreille, 1804 (carpet beetles) *****Family
Endecatomidae ''Endecatomus'' is a genus of beetles, the sole member of the family Endecatomidae. There are at least four described species in ''Endecatomus''. Endecatomidae was formerly treated as a subfamily of Ciidae or Bostrichidae. They are found in the H ...
LeConte, 1861 *****Family
Bostrichidae The Bostrichidae are a family of beetles with more than 700 described species. They are commonly called auger beetles, false powderpost beetles, or horned powderpost beetles. Anatomy The head of most auger beetles cannot be seen from above, as ...
Latreille, 1802 *****Family
Ptinidae Ptinidae is a family of beetles in the superfamily Bostrichoidea. There are at least 220 genera and 2,200 described species in Ptinidae worldwide. The family includes spider beetles and deathwatch beetle, as well as the Cigarette beetle, cigare ...
Latreille, 1802 ***Infraorder Cucujiformia ****Superfamily Lymexyloidea Fleming, 1821 *****Family Lymexylidae Fleming, 1821 ****Superfamily
Cleroidea Cleroidea is a small Taxonomic rank, superfamily of beetles containing over 10,000 species. Most of the members of the group are somewhat slender, often with fairly soft, flexible elytra, and typically hairy or scaly. Description Cleroidea is ...
Latreille, 1802 *****Family
Phloiophilidae ''Phloiophilus edwardsii'' is the sole known species of the beetle family Phloiophilidae in the superfamily Cleroidea. It is native to Europe. The larvae are mycophagous, and have been observed feeding on Basidiomycota, basidiomycetes of the genu ...
Kiesenwetter, 1863 *****Family
Trogossitidae Trogossitidae, also known as bark-gnawing beetles, are a small family in the superfamily Cleroidea. Many taxa formerly within this family have been removed (as of 2019) to other families, such as Lophocateridae, Peltidae, Protopeltidae, Rentoniid ...
Latreille, 1802 *****Family Chaetosomatidae Crowson, 1952 *****Family Metaxinidae Kolibáč, 2004 *****Family
Thanerocleridae Thanerocleridae is a family of beetles belonging to the superfamily Cleroidea. It was formerly considered a subfamily of Cleridae, but was recently elevated to the rank of family. The family has 36 living species in 10 genera, which are found glo ...
Chapin, 1924 *****Family
Cleridae Cleridae are a family of beetles of the superfamily Cleroidea. They are commonly known as checkered beetles. The family Cleridae has a worldwide distribution, and a variety of habitats and feeding preferences. Cleridae have many Ecological nich ...
Latreille, 1802 (checkered beetles) *****Family
Acanthocnemidae ''Acanthocnemus nigricans'' is a species of cleroid beetle, the only species in the genus ''Acanthocnemus'' and the family Acanthocnemidae. Originally native to Australia, the species has spread in modern times to numerous parts of the world, in ...
Crowson, 1964 *****Family Phycosecidae Crowson, 1952 *****Family
Prionoceridae Prionoceridae is a small family of beetles, in the suborder Polyphaga. They form a group within the cleroid beetles and were formerly treated as a subfamily (Prionocerinae) within the family Melyridae. Very little is known of their life history ...
Lacordaire, 1857 *****Family
Mauroniscidae Mauroniscidae is a family of Cleroidea, cleroid beetles, formerly included in the family Melyridae. There are presently five or six genera and roughly 30 described species in Mauroniscidae, all of which are native to the Americas. Almost nothing ...
Majer, 1995 *****Family
Melyridae Melyridae (common name: soft-winged flower beetles) are a family of beetles of the superfamily Cleroidea. Description Most are elongate-oval, soft-bodied beetles 10 mm long or less. Many are brightly patterned in black and brown, yellow, ...
Leach, 1815 (soft-winged flower beetles) ****Superfamily
Cucujoidea Cucujoidea is a superfamily of beetles. This group formerly included all of the families now included in the superfamily Coccinelloidea. They include some fungus beetles and a diversity of lineages of "bark beetles" unrelated to the "true" bark ...
Latreille, 1802 *****Family † Parandrexidae Kirejtshuk, 1994 *****Family † Sinisilvanidae Hong, 2002 *****Family Boganiidae Sen Gupta and Crowson, 1966 *****Family Byturidae Gistel, 1848 *****Family Helotidae Chapuis, 1876 *****Family
Protocucujidae Protocucujidae is a family of beetle Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most oth ...
Crowson, 1954 *****Family Sphindidae Jacquelin du Val, 1860 *****Family Biphyllidae LeConte, 1861 *****Family Erotylidae Latreille, 1802 *****Family Monotomidae Laporte, 1840 *****Family Hobartiidae Sen Gupta and Crowson, 1966 *****Family Cryptophagidae Kirby, 1826 *****Family Agapythidae Sen Gupta and Crowson, 1969 *****Family Priasilphidae Crowson, 1973 *****Family Phloeostichidae Reitter, 1911 *****Family
Silvanidae Silvanidae, "silvan flat bark beetles", is a family of beetles in the superfamily Cucujoidea,Thomas, M. C., and R.A. B. Leschen. 2010. Silvanidae Kirby, 1837. p. 346-350. In: Leschen, R.A.B., R.G. Beutel, and J.F. Lawrence. Coleoptera, Beetles. V ...
Kirby, 1837 *****Family Cucujidae Latreille, 1802 *****Family Myraboliidae Lawrence and Britton, 1991 *****Family Cavognathidae Sen Gupta and Crowson, 1966 *****Family Lamingtoniidae Sen Gupta and Crowson, 1969 *****Family
Passandridae Passandridae, the "parasitic flat bark beetles," are a family of beetles notable for being one of the very few beetle families with larvae that are, as far as known, exclusively ectoparasitic on the immature stages of other beetles and Hymenopter ...
Blanchard, 1845 *****Family Phalacridae Leach, 1815 *****Family Propalticidae Crowson, 1952 *****Family Laemophloeidae Ganglbauer, 1899 *****Family Tasmosalpingidae Lawrence and Britton, 1991 *****Family Cyclaxyridae Gimmel, Leschen and Ślipiński, 2009 *****Family Kateretidae Kirby, 1837 *****Family
Nitidulidae The sap beetles, also known as Nitidulidae, are a family of beetles. They are small (2–6 mm) ovoid, usually dull-coloured beetles, with knobbed antennae. Some have red or yellow spots or bands. They feed mainly on decaying vegetable ma ...
Latreille, 1802 *****Family Smicripidae Horn, 1880 *****Family
Bothrideridae Bothrideridae is a family of beetles in the superfamily Coccinelloidea. They are known commonly as the cocoon-forming beetles or dry bark beetles. They occur worldwide with most native to the Old World tropics. In older literature, the family was ...
Erichson, 1845 *****Family Cerylonidae Billberg, 1820 *****Family Alexiidae Imhoff, 1856 *****Family Discolomatidae Horn, 1878 *****Family Endomychidae Leach, 1815 *****Family
Coccinellidae Coccinellidae () is a widespread family (biology), family of small beetles. They are commonly known as ladybugs in North America and ladybirds in the United Kingdom; "lady" refers to Mary, mother of Jesus, mother Mary. Entomologists use the ...
Latreille, 1807 (ladybirds or lady beetles) *****Family
Corylophidae Corylophidae is a family of minute hooded beetles, sometimes called minute fungus beetles, in the superfamily Coccinelloidea. There are about 18 genera and at least 120 described species in Corylophidae. They feed on microfungi such as molds, ...
LeConte, 1852 *****Family Akalyptoischiidae Lord, Hartley, Lawrence, McHugh, Whiting and Miller, 2010 *****Family
Latridiidae Latridiidae (sometimes spelled "Lathridiidae") is a family of tiny, little-known beetles commonly called minute brown scavenger beetles or fungus beetles. The number of described species currently stands at around 1050 in 29 genera but the number ...
Erichson, 1842 ****Superfamily
Tenebrionoidea The Tenebrionoidea are a very large and diverse superfamily of beetles. It generally corresponds to the Heteromera of earlier authors. Taxonomy Tenebrionoidea contains the following families: * Aderidae Winkler 1927 (ant-like leaf beetles) ...
Latreille, 1802 *****Family
Mycetophagidae The Mycetophagidae or hairy fungus beetles are a family of beetles in the superfamily Tenebrionoidea. The different species are between 1.0 and 6.5 mm in length. The larvae and adults live in decaying leaf litter, fungi, and under bark. Most ...
Leach, 1815 *****Family Archeocrypticidae Kaszab, 1964 *****Family
Pterogeniidae Pterogeniidae is a family of beetles belonging to the superfamily Tenebrionoidea. They are found in South Asia, Southeast Asia, China, New Guinea and the Solomon Islands. Adults and larvae have been found associated with the fruiting bodies of po ...
Crowson, 1953 *****Family
Ciidae The minute tree-fungus beetles, family Ciidae, are a sizeable group of beetles which inhabit Polyporales bracket fungi or coarse woody debris. Most numerous in warmer regions, they are nonetheless widespread and a considerable number of species ...
Leach, 1819 *****Family Tetratomidae Billberg, 1820 *****Family
Melandryidae Melandryidae is a family of beetles in the superfamily Tenebrionoidea. Members of the family are found worldwide, with around 420 species in 60 genera. Larvae and adults are generally associated with rotting wood and wood-decomposing fungi. Gen ...
Leach, 1815 *****Family Mordellidae Latreille, 1802 (tumbling flower beetles) *****Family
Ripiphoridae Ripiphoridae (formerly spelled Rhipiphoridae) is a cosmopolitan family of some 450 described species of beetles sometimes called "wedge-shaped beetles". Ripiphoridae are unusual among beetle families in that many species are hypermetamorphic p ...
Laporte, 1840 *****Family
Zopheridae Zopheridae is a family of beetles belonging to Tenebrionoidea. It has grown considerably in recent years as the members of two other families are now included (Monommatidae and Colydiidae), as subfamilies or (in the former case) even as tribe ( ...
Solier, 1834 *****Family Ulodidae Pascoe, 1869 *****Family Promecheilidae Lacordaire, 1859 *****Family Chalcodryidae Watt, 1974 *****Family Trachelostenidae Lacordaire, 1859 *****Family Darkling beetle, Tenebrionidae Latreille, 1802 (darkling beetles) *****Family Prostomidae Thomson, 1859 *****Family Synchroidae Lacordaire, 1859 *****Family Stenotrachelidae Thomson, 1859 *****Family Oedemeridae Latreille, 1810 *****Family Meloidae Gyllenhal, 1810 (blister beetles) *****Family Mycteridae Perty, 1840 (palm beetles and flower beetles) *****Family Boridae Thomson, 1859 *****Family Trictenotomidae Blanchard, 1845 *****Family Pythidae Solier, 1834 *****Family Pyrochroidae Latreille, 1806 *****Family Salpingidae Leach, 1815 *****Family Anthicidae Latreille, 1819 *****Family Aderidae Csiki, 1909 *****Family Scraptiidae Gistel, 1848 ****Clade Phytophaga *****Superfamily Chrysomeloidea Latreille, 1802 ******Family Oxypeltidae Lacordaire, 1868 ******Family Vesperidae Mulsant, 1839 ******Family Disteniidae Thomson, 1861 ******Family Longhorn beetle, Cerambycidae Latreille, 1802 (longhorn beetles) ******Family Megalopodidae Latreille, 1802 ******Family Orsodacnidae Thomson, 1859 ******Family Leaf beetle, Chrysomelidae Latreille, 1802 (leaf beetles) *****Superfamily Curculionoidea Latreille, 1802 ******Family Nemonychidae Bedel, 1882 ******Family Anthribidae Billberg, 1820 ******Family †Ulyanidae Zherikhin, 1993 ******Family Belidae Schönherr, 1826 ******Family Caridae Thompson, 1992 ******Family Attelabidae Billberg, 1820 ******Family Brentidae Billberg, 1820 ******Family Brachyceridae Billberg, 1820 ******Family Curculionidae Latreille, 1802 (snout beetles, weevils, and bark beetles)


References

* Lawrence, J.F., Newton, A.F. Jr. (1995) Families and subfamilies of Coleoptera (with selected genera, notes, references, and data on family-group names), pp. 779–1006. In: Pakaluk, J., Slipinski, S.A. (eds.), ''Biology, phylogeny, and classification of Coleoptera: Papers celebrating the 80th birthday of Roy A. Crowson''. Muzeum i Instytut Zoologii PAN, Warszawa. * Ross H. Arnett, Jr. and Michael C. Thomas, ''American Beetles'' (CRC Press, 2001–2002) {{Coleoptera Beetle taxonomy, *