The leopard (''Panthera pardus'') is one of the five extant cat

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

in the

genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...

Panthera ''Panthera'' is a genus within the family (biology), family Felidae, and one of two extant genera in the subfamily Pantherinae. It contains the largest living members of the cat family. There are five living species: the jaguar, leopard, lion, ...

''. It has a pale yellowish to dark golden fur with dark spots grouped in rosettes. Its body is slender and muscular reaching a length of with a long tail and a shoulder height of . Males typically weigh , and females . The leopard was first described in 1758, and several

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

were proposed in the 19th and 20th centuries. Today, eight subspecies are recognised in its wide range in

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

and

Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...

. It initially evolved in Africa during the

Early Pleistocene The Early Pleistocene is an unofficial epoch (geology), sub-epoch in the international geologic timescale in chronostratigraphy, representing the earliest division of the Pleistocene Epoch within the ongoing Quaternary Period. It is currently esti ...

, before migrating into Eurasia around the Early–

Middle Pleistocene The Chibanian, more widely known as the Middle Pleistocene (its previous informal name), is an Age (geology), age in the international geologic timescale or a Stage (stratigraphy), stage in chronostratigraphy, being a division of the Pleistocen ...

transition. Leopards were formerly present across Europe, but became extinct in the region at around the end of the

Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...

-early

Holocene The Holocene () is the current geologic time scale, geological epoch, beginning approximately 11,700 years ago. It follows the Last Glacial Period, which concluded with the Holocene glacial retreat. The Holocene and the preceding Pleistocene to ...

. The leopard is adapted to a variety of habitats ranging from

rainforest Rainforests are forests characterized by a closed and continuous tree Canopy (biology), canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforests can be generally classified as tropi ...

steppe In physical geography, a steppe () is an ecoregion characterized by grassland plains without closed forests except near rivers and lakes. Steppe biomes may include: * the montane grasslands and shrublands biome * the tropical and subtropica ...

, including

arid Aridity is the condition of geographical regions which make up approximately 43% of total global available land area, characterized by low annual precipitation, increased temperatures, and limited water availability.Perez-Aguilar, L. Y., Plata ...

and montane areas. It is an

opportunistic 300px, ''Opportunity Seized, Opportunity Missed'', engraving by Theodoor Galle, 1605 Opportunism is the practice of taking advantage of circumstances — with little regard for principles or with what the consequences are for others. Opport ...

predator, hunting mostly

ungulate Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with Hoof, hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined ...

s and

primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...

s. It relies on its spotted pattern for

camouflage Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...

as it stalks and ambushes its prey, which it sometimes drags up a tree. It is a

solitary animal Sociality is the degree to which individuals in an animal population tend to associate in social groups (gregariousness) and form cooperative societies. Sociality is a survival response to evolutionary pressures. For example, when a mother was ...

outside the mating season and when raising cubs. Females usually give birth to a litter of 2–4 cubs once in 15–24 months. Both male and female leopards typically reach

sexual maturity Sexual maturity is the capability of an organism to reproduce. In humans, it is related to both puberty and adulthood. ''Puberty'' is the biological process of sexual maturation, while ''adulthood'', the condition of being socially recognized ...

at the age 2–2.5 years. Listed as Vulnerable on the

IUCN Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is an inventory of the global conservation status and extinction risk of biological ...

, leopard populations are currently threatened by

habitat loss Habitat destruction (also termed habitat loss or habitat reduction) occurs when a natural habitat is no longer able to support its native species. The organisms once living there have either moved elsewhere, or are dead, leading to a decrease ...

and fragmentation, and are declining in large parts of the global range. Leopards have had cultural roles in

Ancient Greece Ancient Greece () was a northeastern Mediterranean civilization, existing from the Greek Dark Ages of the 12th–9th centuries BC to the end of classical antiquity (), that comprised a loose collection of culturally and linguistically r ...

West Africa West Africa, also known as Western Africa, is the westernmost region of Africa. The United Nations geoscheme for Africa#Western Africa, United Nations defines Western Africa as the 16 countries of Benin, Burkina Faso, Cape Verde, The Gambia, Gha ...

and modern Western culture. Leopard skins are popular in fashion.

Etymology

The English name "leopard" comes from

Old French Old French (, , ; ) was the language spoken in most of the northern half of France approximately between the late 8th -4; we might wonder whether there's a point at which it's appropriate to talk of the beginnings of French, that is, when it wa ...

or Middle French , that derives from Latin and ancient Greek (). could be a Compound (linguistics), compound of (), meaning , and (), meaning . The word originally referred to a cheetah (''Acinonyx jubatus''). "Panther" is another common name, derived from Latin and ancient Greek (); The Generic name (biology), generic name ''Panthera'' originates in Latin , a hunting net for catching wild beasts to be used by the

Roman Roman or Romans most often refers to: *Rome, the capital city of Italy *Ancient Rome, Roman civilization from 8th century BC to 5th century AD *Roman people, the people of Roman civilization *Epistle to the Romans, shortened to Romans, a letter w ...

s in combats. is the masculine singular form.

Taxonomy

''Felis pardus'' was the

scientific name In Taxonomy (biology), taxonomy, binomial nomenclature ("two-term naming system"), also called binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, both of which use Latin gramm ...

proposed by

Carl Linnaeus Carl Linnaeus (23 May 1707 – 10 January 1778), also known after ennoblement in 1761 as Carl von Linné,#Blunt, Blunt (2004), p. 171. was a Swedish biologist and physician who formalised binomial nomenclature, the modern system of naming o ...

in 1758. The Generic name (biology), generic name ''

'' was first used by

Lorenz Oken Lorenz Oken (1 August 1779 – 11 August 1851) was a Germans, German natural history, naturalist, botany, botanist, biologist, and ornithology, ornithologist. Biography Oken was born Lorenz Okenfuss () in Bohlsbach (now part of Offenburg), Ortena ...

in 1816, who included all the known spotted cats into this group. Oken's classification was not widely accepted, and ''

Felis ''Felis'' is a genus of small and medium-sized cat species native to most of Africa and south of 60° latitude in Europe and Asia to Indochina. The genus includes the domestic cat. The smallest of the seven ''Felis'' species is the black-footed ...

'' or ''

Leopardus ''Leopardus'' is a genus comprising eight species of Felinae, small cats native to the Americas. This genus is considered the oldest branch of a genetic lineage of small cats in the Americas whose common ancestor crossed the Bering land bridge fr ...

'' was used as the generic name until the early 20th century. The leopard was designated as the

type species In International_Code_of_Zoological_Nomenclature, zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the spe ...

of ''Panthera'' by

Joel Asaph Allen Joel Asaph Allen (July 19, 1838 – August 29, 1921) was an American zoology, zoologist, mammalogy, mammalogist, and ornithology, ornithologist. He became the first president of the American Ornithologists' Union, the first curator of birds and ma ...

in 1902. In 1917,

Reginald Innes Pocock Reginald Innes Pocock, (4 March 1863 – 9 August 1947) was a British zoologist. Pocock was born in Clifton, Bristol, the fourth son of Rev. Nicholas Pocock and Edith Prichard. He began showing interest in natural history at St. Edward's ...

also subordinated the

tiger The tiger (''Panthera tigris'') is a large Felidae, cat and a member of the genus ''Panthera'' native to Asia. It has a powerful, muscular body with a large head and paws, a long tail and orange fur with black, mostly vertical stripes. It is ...

(''P. tigris''),

lion The lion (''Panthera leo'') is a large Felidae, cat of the genus ''Panthera'', native to Sub-Saharan Africa and India. It has a muscular, broad-chested body (biology), body; a short, rounded head; round ears; and a dark, hairy tuft at the ...

(''P. leo''), and

jaguar The jaguar (''Panthera onca'') is a large felidae, cat species and the only extant taxon, living member of the genus ''Panthera'' that is native to the Americas. With a body length of up to and a weight of up to , it is the biggest cat spe ...

(''P. onca'') to ''Panthera''.

Living subspecies

Following Linnaeus' first description, 27 leopard

were proposed by naturalists between 1794 and 1956. Since 1996, only eight subspecies have been considered valid on the basis of

mitochondrial A mitochondrion () is an organelle found in the cells of most eukaryotes, such as animals, plants and fungi. Mitochondria have a double membrane structure and use aerobic respiration to generate adenosine triphosphate (ATP), which is used ...

analysis. Later analysis revealed a ninth valid subspecies, the

Arabian leopard The Arabian leopard (''Panthera pardus nimr'') is the smallest leopard subspecies. It was described in 1830 and is native to the Arabian Peninsula, where it was widely distributed in rugged hilly and montane terrain until the late 1970s. Today, t ...

. In 2017, the Cat Classification Task Force of the Cat Specialist Group recognized the following eight subspecies as valid taxa: Results of an analysis of molecular variance and pairwise

fixation index The fixation index (FST) is a measure of population differentiation due to genetic structure. It is frequently estimated from Polymorphism (biology), genetic polymorphism data, such as single-nucleotide polymorphisms (SNP) or Microsatellite (genet ...

of 182 African leopard museum specimens showed that some African leopards exhibit higher genetic differences than Asian leopard subspecies.

Evolution

Results of

phylogenetic In biology, phylogenetics () is the study of the evolutionary history of life using observable characteristics of organisms (or genes), which is known as phylogenetic inference. It infers the relationship among organisms based on empirical dat ...

studies based on

nuclear DNA Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. ...

and

mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...

analysis showed that the last

common ancestor Common descent is a concept in evolutionary biology applicable when one species is the ancestor of two or more species later in time. According to modern evolutionary biology, all living beings could be descendants of a unique ancestor commonl ...

of the ''Panthera'' and ''

Neofelis ''Neofelis'' is a genus comprising two extant cat species in Southeast Asia: the clouded leopard (''Neofelis nebulosa'') of mainland Asia, and the Sunda clouded leopard (''Neofelis diardi'') of Sumatra and Borneo. The scientific name ''Neofel ...

'' genera is thought to have lived about . ''Neofelis'' diverged about from the ''Panthera'' lineage. The tiger diverged about , followed by the snow leopard about and the leopard about . The leopard is a

sister taxon In phylogenetics, a sister group or sister taxon, also called an adelphotaxon, comprises the closest relative(s) of another given unit in an evolutionary tree. Definition The expression is most easily illustrated by a cladogram: Taxon A and ...

to a

clade In biology, a clade (), also known as a Monophyly, monophyletic group or natural group, is a group of organisms that is composed of a common ancestor and all of its descendants. Clades are the fundamental unit of cladistics, a modern approach t ...

within ''Panthera'', consisting of the lion and the jaguar. Results of a phylogenetic analysis of chemical

secretion Secretion is the movement of material from one point to another, such as a secreted chemical substance from a cell or gland. In contrast, excretion is the removal of certain substances or waste products from a cell or organism. The classical mec ...

s amongst cats indicated that the leopard is closely related to the lion. The geographic origin of the ''Panthera'' is most likely northern Central Asia. The leopard-lion clade was distributed in the Asian and African

Palearctic The Palearctic or Palaearctic is a biogeographic realm of the Earth, the largest of eight. Confined almost entirely to the Eastern Hemisphere, it stretches across Europe and Asia, north of the foothills of the Himalayas, and North Africa. Th ...

since at least the early

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58mitochondrial genome Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the DNA contained in ...

s of the leopard, lion and

snow leopard The snow leopard (''Panthera uncia'') is a species of large cat in the genus ''Panthera'' of the family Felidae. The species is native to the mountain ranges of Central and South Asia. It is listed as Vulnerable on the IUCN Red List because ...

are more similar to each other than their

nuclear genome Nuclear DNA (nDNA), or nuclear deoxyribonucleic acid, is the DNA contained within each cell nucleus of a eukaryotic organism. It encodes for the majority of the genome in eukaryotes, with mitochondrial DNA and plastid DNA coding for the rest. It ...

s, indicating that their ancestors hybridized with the snow leopard at some point in their

evolution Evolution is the change in the heritable Phenotypic trait, characteristics of biological populations over successive generations. It occurs when evolutionary processes such as natural selection and genetic drift act on genetic variation, re ...

. The oldest unambiguous

fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserve ...

s of the leopard are from Eastern Africa, dating to around 2 million years ago. Leopard-like fossil bones and teeth possibly dating to the

Pliocene The Pliocene ( ; also Pleiocene) is the epoch (geology), epoch in the geologic time scale that extends from 5.33 to 2.58Perrier in

France France, officially the French Republic, is a country located primarily in Western Europe. Overseas France, Its overseas regions and territories include French Guiana in South America, Saint Pierre and Miquelon in the Atlantic Ocean#North Atlan ...

, northeast of

London London is the Capital city, capital and List of urban areas in the United Kingdom, largest city of both England and the United Kingdom, with a population of in . London metropolitan area, Its wider metropolitan area is the largest in Wester ...

, and in

Valdarno The Valdarno is the valley of the river Arno, from Florence to the sea. The name applies to the entire river basin, though usage of the term generally excludes Casentino and the valleys formed by major tributaries. Some towns in the area: * R ...

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

. Until 1940, similar fossils dating back to the

Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...

were excavated mostly in

loess A loess (, ; from ) is a clastic rock, clastic, predominantly silt-sized sediment that is formed by the accumulation of wind-blown dust. Ten percent of Earth's land area is covered by loesses or similar deposition (geology), deposits. A loess ...

and caves at 40 sites in Europe, including Furninha Cave near

Lisbon Lisbon ( ; ) is the capital and largest city of Portugal, with an estimated population of 567,131, as of 2023, within its administrative limits and 3,028,000 within the Lisbon Metropolitan Area, metropolis, as of 2025. Lisbon is mainlan ...

, Genista Caves in

Gibraltar Gibraltar ( , ) is a British Overseas Territories, British Overseas Territory and British overseas cities, city located at the southern tip of the Iberian Peninsula, on the Bay of Gibraltar, near the exit of the Mediterranean Sea into the A ...

, and Santander Province in northern

Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...

to several sites across France,

Switzerland Switzerland, officially the Swiss Confederation, is a landlocked country located in west-central Europe. It is bordered by Italy to the south, France to the west, Germany to the north, and Austria and Liechtenstein to the east. Switzerland ...

, Italy,

Austria Austria, formally the Republic of Austria, is a landlocked country in Central Europe, lying in the Eastern Alps. It is a federation of nine Federal states of Austria, states, of which the capital Vienna is the List of largest cities in Aust ...

Germany Germany, officially the Federal Republic of Germany, is a country in Central Europe. It lies between the Baltic Sea and the North Sea to the north and the Alps to the south. Its sixteen States of Germany, constituent states have a total popu ...

, in the north up to

Derby Derby ( ) is a City status in the United Kingdom, city and Unitary authorities of England, unitary authority area on the River Derwent, Derbyshire, River Derwent in Derbyshire, England. Derbyshire is named after Derby, which was its original co ...

England England is a Countries of the United Kingdom, country that is part of the United Kingdom. It is located on the island of Great Britain, of which it covers about 62%, and List of islands of England, more than 100 smaller adjacent islands. It ...

, in the east to

Přerov Přerov (; ) is a city in the Olomouc Region of the Czech Republic. It has about 41,000 inhabitants. It lies on the Bečva River. In the past it was a major crossroad in the heart of Moravia in the Czech Republic. The historic city centre is we ...

in the

Czech Republic The Czech Republic, also known as Czechia, and historically known as Bohemia, is a landlocked country in Central Europe. The country is bordered by Austria to the south, Germany to the west, Poland to the northeast, and Slovakia to the south ...

and the Baranya in southern

Hungary Hungary is a landlocked country in Central Europe. Spanning much of the Pannonian Basin, Carpathian Basin, it is bordered by Slovakia to the north, Ukraine to the northeast, Romania to the east and southeast, Serbia to the south, Croatia and ...

. Leopards arrived in Eurasia during the late Early to Middle Pleistocene around 1.2 to 0.6 million years ago. Four European Pleistocene leopard subspecies were proposed. ''P. p. begoueni'' from the beginning of the

was replaced about by ''P. p. sickenbergi'', which in turn was replaced by ''P. p. antiqua'' around 0.3 million years ago. ''P. p. spelaea'' is the most recent subspecies that appeared at the beginning of the

and survived until about 11,000 years ago and possibly into the early

in the

Iberian Peninsula The Iberian Peninsula ( ), also known as Iberia, is a peninsula in south-western Europe. Mostly separated from the rest of the European landmass by the Pyrenees, it includes the territories of peninsular Spain and Continental Portugal, comprisin ...

. Leopards depicted in cave paintings in

Chauvet Cave The Chauvet-Pont-d'Arc Cave ( ) in the Ardèche department of southeastern France is a cave that contains some of the best-preserved figurative cave paintings in the world, as well as other evidence of Upper Paleolithic life.Clottes (2003b), p. ...

provide indirect evidence of leopard presence in Europe. Leopard fossils dating to the Late Pleistocene were found in Biśnik Cave in south-central

Poland Poland, officially the Republic of Poland, is a country in Central Europe. It extends from the Baltic Sea in the north to the Sudetes and Carpathian Mountains in the south, bordered by Lithuania and Russia to the northeast, Belarus and Ukrai ...

. Fossil remains were also excavated in the Iberian and Italian Peninsula, and in the

Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...

. Leopard fossils dating to the Pleistocene were also excavated in the

Japanese archipelago The is an archipelago of list of islands of Japan, 14,125 islands that form the country of Japan. It extends over from the Sea of Okhotsk in the northeast to the East China Sea, East China and Philippine Sea, Philippine seas in the southwest al ...

. Leopard fossils were also found in

Taiwan Taiwan, officially the Republic of China (ROC), is a country in East Asia. The main geography of Taiwan, island of Taiwan, also known as ''Formosa'', lies between the East China Sea, East and South China Seas in the northwestern Pacific Ocea ...

Hybrids

In 1953, a male leopard and a female lion were crossbred in Hanshin Park in Nishinomiya, Japan. Their offspring known as a

leopon A ''Panthera'' hybrid is a crossbreed between individuals of any of the five species of the genus ''Panthera'': the tiger, lion, jaguar, leopard, and snow leopard. Most hybrids would not be perpetuated in the wild as the territories of the par ...

was born in 1959 and 1961, all cubs were spotted and bigger than a juvenile leopard. Attempts to mate a leopon with a tigress proved unsuccessful.

Characteristics

The leopard's fur is generally soft and thick, notably softer on the belly than on the back. Its skin colour varies between individuals from pale yellowish to dark golden with dark spots grouped in rosettes. Its underbelly is white and its ringed tail is shorter than its body. Its pupils are round. Leopards living in arid regions are pale cream, yellowish to ochraceous and

rufous Rufous () is a color that may be described as reddish-brown or brownish- red, as of rust or oxidised iron. The first recorded use of ''rufous'' as a color name in English was in 1782. However, the color is also recorded earlier in 1527 as a d ...

in colour; those living in forests and mountains are much darker and deep golden. Spots fade toward the white underbelly and the insides and lower parts of the legs. Rosettes are circular in East African leopard populations, and tend to be squarish in Southern African and larger in Asian leopard populations. The fur tends to be grayish in colder climates, and dark golden in

habitats. Rosette patterns are unique in each individual. This pattern is thought to be an adaptation to dense vegetation with patchy shadows, where it serves as

. Its white-tipped tail is about long, white underneath and with spots that form incomplete bands toward the end of the tail. The guard hairs protecting the basal hairs are short, in face and head, and increase in length toward the flanks and the belly to about . Juveniles have woolly fur that appear to be dark-coloured due to the densely arranged spots. Its fur tends to grow longer in colder climates. The leopard's rosettes differ from those of the

, which are darker and with smaller spots inside. The leopard has a

diploid Ploidy () is the number of complete sets of chromosomes in a cell, and hence the number of possible alleles for autosomal and pseudoautosomal genes. Here ''sets of chromosomes'' refers to the number of maternal and paternal chromosome copies, ...

chromosome A chromosome is a package of DNA containing part or all of the genetic material of an organism. In most chromosomes, the very long thin DNA fibers are coated with nucleosome-forming packaging proteins; in eukaryotic cells, the most import ...

number of 38. Melanistic leopards are also known as

black panther A black panther is the Melanism, melanistic colour variant of the leopard (''Panthera pardus'') and the jaguar (''Panthera onca''). Black panthers of both species have excess black pigments, but their typical Rosette (zoology), rosettes are al ...

Melanism Melanism is the congenital excess of melanin in an organism resulting in dark pigment. Pseudomelanism, also called abundism, is another variant of pigmentation, identifiable by dark spots or enlarged stripes, which cover a large part of the bod ...

in leopards is caused by a

recessive allele In genetics, dominance is the phenomenon of one variant (allele) of a gene on a chromosome masking or overriding the effect of a different variant of the same gene on the other copy of the chromosome. The first variant is termed dominant and ...

and is inherited as a recessive trait. In India, nine pale and white leopards were reported between 1905 and 1967. Leopards exhibiting

erythrism Erythrism or erythrochroism refers to an unusual reddish pigmentation of an animal's hair Hair is a protein filament that grows from follicles found in the dermis. Hair is one of the defining characteristics of mammals. The human body, ap ...

were recorded between 1990 and 2015 in South Africa's Madikwe Game Reserve and in

Mpumalanga Mpumalanga () is one of the nine provinces of South Africa. The name means "East", or literally "The Place Where the Sun Rises" in the Nguni languages. Mpumalanga lies in eastern South Africa, bordering Eswatini and Mozambique. It shares bor ...

. The cause of this morph known as a "strawberry leopard" or "pink panther" is not well understood.

Size

The leopard is a slender and muscular cat, with relatively short limbs and a broad head. It is

sexually dimorphic Sexual dimorphism is the condition where sexes of the same species exhibit different Morphology (biology), morphological characteristics, including characteristics not directly involved in reproduction. The condition occurs in most dioecy, di ...

with males larger and heavier than females. Males stand at the shoulder, while females are tall. The head-and-body length ranges between with a long tail. Sizes vary geographically. Males typically weigh , and females . Occasionally, large males can grow up to . Leopards from the

Cape Province The Province of the Cape of Good Hope (), commonly referred to as the Cape Province () and colloquially as The Cape (), was a province in the Union of South Africa and subsequently the Republic of South Africa. It encompassed the old Cape Co ...

in South Africa are generally smaller, reaching only in males. The heaviest wild leopard in Southern Africa weighed around , and it measured . In 2016, an Indian leopard killed in

Himachal Pradesh Himachal Pradesh (; Sanskrit: ''himācāl prādes;'' "Snow-laden Mountain Province") is a States and union territories of India, state in the northern part of India. Situated in the Western Himalayas, it is one of the thirteen Indian Himalayan ...

measured with an estimated weight of ; it was perhaps the largest known wild leopard in India. The largest recorded skull of a leopard was found in India in 1920 and measured in basal length, in breadth, and weighed . The skull of an African leopard measured in basal length, and in breadth, and weighed .

Distribution and habitat

The leopard has the largest distribution of all wild cats, occurring widely in Africa and Asia, although populations are fragmented and declining. It inhabits foremost

savanna A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) biome and ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach th ...

and

, and areas where

grassland A grassland is an area where the vegetation is dominance (ecology), dominated by grasses (Poaceae). However, sedge (Cyperaceae) and rush (Juncaceae) can also be found along with variable proportions of legumes such as clover, and other Herbaceo ...

woodland A woodland () is, in the broad sense, land covered with woody plants (trees and shrubs), or in a narrow sense, synonymous with wood (or in the U.S., the '' plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunli ...

s and

riparian forest A riparian forest or riparian woodland is a forested or wooded area of land adjacent to a body of water such as a river, stream, pond, lake, marshland, estuary, canal, Sink (geography), sink, or reservoir. Due to the broad nature of the definitio ...

s remain largely undisturbed. It also persists in urban environments, if it is not persecuted, has sufficient prey and patches of vegetation for shelter during the day. The leopard's range in

is estimated to have drastically declined by 95%, and in the

Sahara desert The Sahara (, ) is a desert spanning across North Africa. With an area of , it is the largest hot desert in the world and the list of deserts by area, third-largest desert overall, smaller only than the deserts of Antarctica and the northern Ar ...

by 97%. In

sub-Saharan Africa Sub-Saharan Africa is the area and regions of the continent of Africa that lie south of the Sahara. These include Central Africa, East Africa, Southern Africa, and West Africa. Geopolitically, in addition to the list of sovereign states and ...

, it is still numerous and surviving in marginal habitats where other large cats have disappeared. In southeastern

Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...

, an individual found killed in 2017 was the first sighting of the leopard in this area in 65 years. In

West Asia West Asia (also called Western Asia or Southwest Asia) is the westernmost region of Asia. As defined by most academics, UN bodies and other institutions, the subregion consists of Anatolia, the Arabian Peninsula, Iran, Mesopotamia, the Armenian ...

, the leopard inhabits the areas of southern and southeastern

Anatolia Anatolia (), also known as Asia Minor, is a peninsula in West Asia that makes up the majority of the land area of Turkey. It is the westernmost protrusion of Asia and is geographically bounded by the Mediterranean Sea to the south, the Aegean ...

. Leopard populations in the

Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...

are small and fragmented. In the

Indian subcontinent The Indian subcontinent is a physiographic region of Asia below the Himalayas which projects into the Indian Ocean between the Bay of Bengal to the east and the Arabian Sea to the west. It is now divided between Bangladesh, India, and Pakista ...

, the leopard is still relatively abundant, with greater numbers than those of other ''Panthera'' species. Some leopard populations in

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

live quite close to human settlements and even in semi-developed areas. Although adaptable to human disturbances, leopards require healthy prey populations and appropriate vegetative cover for hunting for prolonged survival and thus rarely linger in heavily developed areas. Due to the leopard's stealth, people often remain unaware that it lives in nearby areas. As of 2020, the leopard population within forested habitats in India's tiger range landscapes was estimated at 12,172 to 13,535 individuals. Surveyed landscapes included elevations below in the

Shivalik Hills The Sivalik Hills, also known as Churia Hills, are a mountain range of the outer Himalayas. The literal translation of "Sivalik" is 'tresses of Shiva'. The hills are known for their numerous fossils, and are also home to the Soanian Middle Paleo ...

and Gangetic plains,

Central India Central India refers to a geographical region of India that generally includes the states of Chhattisgarh and Madhya Pradesh. The Central Zonal Council, established by the Government of India, includes these states as well as Uttar Prades ...

and

Eastern Ghats The Eastern Ghats is a mountain range that stretches along the East Coast of India, eastern coast of the Indian peninsula. Covering an area of , it traverses the states and union territories of India, states of Odisha, Telangana, Andhra Prade ...

, Western Ghats, the

Brahmaputra River The Brahmaputra is a trans-boundary river which flows through Southwestern China, Northeastern India, and Bangladesh. It is known as Brahmaputra or Luit in Assamese language, Assamese, Yarlung Tsangpo in Lhasa Tibetan, Tibetan, the Siang/Dihan ...

basin and hills in

Northeast India Northeast India, officially the North Eastern Region (NER), is the easternmost region of India representing both a geographic and political Administrative divisions of India, administrative division of the country. It comprises eight States and ...

. In Nepal's

Kanchenjunga Conservation Area Kanchenjunga Conservation Area is a protected area in the Himalayas of eastern Nepal that was established in 1997. It covers in the Taplejung District and comprises two peaks of Kanchenjunga. In the north it adjoins the Qomolangma National Nat ...

, a melanistic leopard was photographed at an elevation of by a camera trap in May 2012. In Sri Lanka, leopards were recorded in

Yala National Park Yala National Park is the most visited and second largest national park in Sri Lanka, bordering the Indian Ocean. The park consists of five blocks, three of which are now open to the public. There are also two adjoining parks, Kumana National Pa ...

and in unprotected forest patches, tea estates, grasslands, home gardens,

pine A pine is any conifer tree or shrub in the genus ''Pinus'' () of the family Pinaceae. ''Pinus'' is the sole genus in the subfamily Pinoideae. ''World Flora Online'' accepts 134 species-rank taxa (119 species and 15 nothospecies) of pines as cu ...

and

eucalyptus ''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...

plantations. In Myanmar, leopards were recorded for the first time by camera traps in the hill forests of Myanmar's

Karen State Kayin State (, ; ; , ), formerly known as Karen State, is a state of Myanmar. The capital city is Hpa-An, also spelled Pa-An. The terrain of the state is mountainous; with the Dawna Range running along the state in a NNW–SSE direction, and ...

. The Northern Tenasserim Forest Complex in southern Myanmar is considered a leopard stronghold. In Thailand, leopards are present in the Western Forest Complex, Kaeng Krachan- Kui Buri, Khlong Saeng- Khao Sok protected area complexes and in Hala Bala Wildlife Sanctuary bordering Malaysia. In

Peninsular Malaysia Peninsular Malaysia, historically known as Malaya and also known as West Malaysia or the Malaysian Peninsula, is the western part of Malaysia that comprises the southern part of the Malay Peninsula on Mainland Southeast Asia and the list of isla ...

, leopards are present in Belum-Temengor, Taman Negara and Endau-Rompin National Parks. In Laos, leopards were recorded in Nam Et-Phou Louey National Biodiversity Conservation Area and Nam Kan National Protected Area. In Cambodia, leopards inhabit deciduous

dipterocarp forest Dipterocarpaceae is a family of flowering plants with 22 genera and about 695 known species of mainly lowland tropical forest trees. Their distribution is pantropical, from northern South America to Africa, the Seychelles, India, Indochina, Indo ...

Phnom Prich Wildlife Sanctuary Phnom Prich Wildlife Sanctuary () is a large protected area in eastern Cambodia that was established in 1993. It is part of maybe the largest protected area complex in southeast Asia. Phnom Prich Wildlife Sanctuary borders Mondulkiri Protected F ...

and

Mondulkiri Protected Forest Sre Pok Wildlife Sanctuary (, formerly Mondulkiri Protected Forest) is a large wildlife sanctuary in Mondulkiri Province, eastern Cambodia established on May 9, 2016, according to Sub-decree No. 85 ANKr.BK. Formerly, classified as Mondulkiri P ...

. In southern China, leopards were recorded only in the

Qinling The Qinling () or Qin Mountains, formerly known as the Nanshan ("Southern Mountains"), are a major east–west mountain range in southern Shaanxi Province, China. The mountains mark the divide between the drainage basins of the Yangtze and Ye ...

Mountains during surveys in 11 nature reserves between 2002 and 2009. In Java, leopards inhabit dense

tropical rainforest Tropical rainforests are dense and warm rainforests with high rainfall typically found between 10° north and south of the Equator. They are a subset of the tropical forest biome that occurs roughly within the 28° latitudes (in the torrid zo ...

s and dry

deciduous forest In the fields of horticulture and botany, the term deciduous () means "falling off at maturity" and "tending to fall off", in reference to trees and shrubs that seasonally shed leaves, usually in the autumn; to the shedding of petals, after flo ...

s at elevations from sea level to . Outside protected areas, leopards were recorded in mixed

agricultural Agriculture encompasses crop and livestock production, aquaculture, and forestry for food and non-food products. Agriculture was a key factor in the rise of sedentary human civilization, whereby farming of domesticated species created f ...

land,

secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...

and production forest between 2008 and 2014. In the Russian Far East, it inhabits

temperate coniferous forest Temperate coniferous forest is a terrestrial biome defined by the World Wide Fund for Nature. Temperate coniferous forests are found predominantly in areas with warm summers and cool winters, and vary in their kinds of plant life. In some, needle ...

s where winter temperatures reach a low of .

Behaviour and ecology

The leopard is a solitary and

territorial A territory is an area of land, sea, or space, belonging or connected to a particular country, person, or animal. In international politics, a territory is usually a geographic area which has not been granted the powers of self-government, ...

animal. It is typically shy and alert when crossing roadways and encountering oncoming vehicles, but may be emboldened to attack people or other animals when threatened. Adults associate only in the mating season. Females continue to interact with their offspring even after weaning and have been observed sharing kills with their offspring when they can not obtain any prey. They produce a number of vocalizations, including growls, snarls and purrs. Cubs call their mother with meows and an ''urr-urr'' sound. The most notable vocalization is the 'sawing'

roar A roar is a type of animal vocalization that is loud, deep and resonating. Many mammals have evolved to produce roars and other roar-like vocals for purposes such as long-distance communication and intimidation. These include various species of ...

, which consists of deep, repeated strokes. This likely functions in establishing territories and attracting mates. The whitish spots on the back of its ears are thought to play a role in communication. It has been hypothesized that the white tips of their tails may function as a 'follow-me' signal in intraspecific communication. However, no significant association were found between a conspicuous colour of tail patches and behavioural variables in carnivores. Leopards are mainly active from dusk till dawn and will rest for most of the day and some hours at night in thickets, among rocks or over tree branches. Leopards have been observed walking across their range at night; wandering up to if disturbed. In some regions, they are

nocturnal Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite. Nocturnal creatur ...

. In western African forests, they have been observed to be largely diurnal and hunting during twilight, when their prey animals are active; activity patterns vary between seasons. Leopards can climb trees quite skillfully, often resting on tree branches and descending headfirst. They can run at over , leap over horizontally, and jump up to vertically.

Social spacing

Kruger National Park Kruger National Park () is a national park in South Africa covering an area of in the provinces of Limpopo and Mpumalanga in the country's northeast. It extends from north to south and from east to west. The administrative headquarters are i ...

, most leopards tend to keep apart. Males occasionally interact with their partners and cubs, and exceptionally this can extend beyond to two generations. Aggressive encounters are rare, typically limited to defending territories from intruders. In a South African reserve, a male was wounded in a male–male territorial battle over a carcass. Males occupy

home range A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. ...

s that often overlap with a few smaller female home ranges, probably as a strategy to enhance access to females. In the

Ivory Coast Ivory Coast, also known as Côte d'Ivoire and officially the Republic of Côte d'Ivoire, is a country on the southern coast of West Africa. Its capital city of Yamoussoukro is located in the centre of the country, while its largest List of ci ...

, the home range of a female was completely enclosed within a male's. Females live with their cubs in home ranges that overlap extensively, probably due to the association between mothers and their offspring. There may be a few other fluctuating home ranges belonging to young individuals. It is not clear if male home ranges overlap as much as those of females do. Individuals try to drive away intruders of the same sex. A study of leopards in the Namibian farmlands showed that the size of home ranges was not significantly affected by sex, rainfall patterns or season; the higher the prey availability in an area, the greater the leopard population density and the smaller the size of home ranges, but they tend to expand if there is human interference. Sizes of home ranges vary geographically and depending on habitat and availability of prey. In the

Serengeti The Serengeti ( ) ecosystem is a geographical region in Africa, spanning the Mara and Arusha Regions of Tanzania. The protected area within the region includes approximately of land, including the Serengeti National Park and several game r ...

, males have home ranges of and females of ; but males in northeastern Namibia of and females of . They are even larger in arid and montane areas. In Nepal's Bardia National Park, male home ranges of and female ones of are smaller than those generally observed in Africa.

Hunting and diet

The leopard is a

carnivore A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant Plants are the eukaryotes that form the Kingdom (biology), kingdom Plantae; they ar ...

that prefers medium-sized prey with a body mass ranging from . Prey species in this weight range tend to occur in dense habitat and to form small herds. Species that prefer open areas and have well-developed anti-predator strategies are less preferred. More than 100 prey species have been recorded. The most preferred species are

ungulates Ungulates ( ) are members of the diverse clade Euungulata ("true ungulates"), which primarily consists of large mammals with hooves. Once part of the clade "Ungulata" along with the clade Paenungulata, "Ungulata" has since been determined to b ...

, such as

impala The impala or rooibok (''Aepyceros melampus'', lit. 'black-footed high-horn' in Ancient Greek) is a medium-sized antelope found in eastern and southern Africa. The only extant member of the genus '' Aepyceros'', and tribe Aepycerotini, it ...

, bushbuck, common duiker and

chital The chital or cheetal (''Axis axis''; ), also called spotted deer, chital deer and axis deer, is a deer species native to the Indian subcontinent. It was first described by Johann Christian Polycarp Erxleben in 1777. A moderate-sized deer, mal ...

Primate Primates is an order (biology), order of mammals, which is further divided into the Strepsirrhini, strepsirrhines, which include lemurs, galagos, and Lorisidae, lorisids; and the Haplorhini, haplorhines, which include Tarsiiformes, tarsiers a ...

s preyed upon include white-eyelid mangabeys,

guenon The guenons (, ) are Old World monkeys of the genus ''Cercopithecus'' (). Not all members of this genus have the word "guenon" in their common names; also, because of changes in scientific classification, some monkeys in other genera may have co ...

s and

gray langur ''Semnopithecus'' is a genus of Old World monkeys native to the Indian subcontinent, with all species with the exception of two being commonly known as gray langurs. Traditionally only the species ''Semnopithecus entellus'' was recognized, but si ...

s. Leopards also kill smaller carnivores like

black-backed jackal The black-backed jackal (''Lupulella mesomelas'') is a medium-sized Caninae, canine native to East Africa, eastern and southern Africa. These regions are separated by roughly . One region includes the southernmost tip of the continent, includin ...

bat-eared fox The bat-eared fox (''Otocyon megalotis'') is a species of fox found on the African savanna. It is the only extant species of the genus ''Otocyon'' and a Basal (phylogenetics), basal species of Canidae, canid. Fossil records indicate this canid ...

, genet and cheetah. In urban environments,

domestic dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers ...

s provide an important food source. The largest prey killed by a leopard was reportedly a male eland weighing . A study in

Wolong National Nature Reserve Wolong National Nature Reserve (from Chinese ), officially known as Wolong Special Administrative Region, is a national protected area located in Wenchuan County in China. Established in 1963 with an initial size of about , the reserve was fur ...

in southern China demonstrated variation in the leopard's diet over time; over the course of seven years, the vegetative cover receded, and leopards opportunistically shifted from primarily consuming tufted deer to pursuing bamboo rats and other smaller prey. The leopard depends mainly on its acute senses of hearing and vision for hunting. It primarily hunts at night in most areas. In western African forests and Tsavo National Park, they have also been observed hunting by day. They usually hunt on the ground. In the Serengeti, they have been seen to ambush prey by descending on it from trees. It stalks its prey and tries to approach as closely as possible, typically within of the target, and, finally, pounces on it and kills it by suffocation. It kills small prey with a bite to the back of the neck, but holds larger animals by the throat and strangles them. It caches kills up to apart. It is able to take large prey due to its powerful jaw muscles, and is therefore strong enough to drag carcasses heavier than itself up into trees; an individual was seen to haul a young giraffe weighing nearly up into a tree. It eats small prey immediately, but drags larger carcasses over several hundred metres and caches it safely in trees, bushes or even caves; this behaviour allows the leopard to store its prey away from rivals, and offers it an advantage over them. The way it stores the kill depends on local topography and individual preferences, varying from trees in Kruger National Park to bushes in the plain terrain of the Kalahari. Average daily consumption rates of were estimated for males and of for females. In the southern

Kalahari Desert The Kalahari Desert is a large semiarid climate, semiarid sandy savanna in Southern Africa covering including much of Botswana as well as parts of Namibia and South Africa. It is not to be confused with the Angolan, Namibian, and South African ...

, leopards meet their water requirements by the bodily fluids of prey and

succulent plant In botany, succulent plants, also known as succulents, are plants with parts that are thickened, fleshy, and engorged, usually to retain water in arid climates or soil conditions. The word ''succulent'' comes from the Latin word ''sucus'', meanin ...

s; they drink water every two to three days and feed infrequently on moisture-rich plants such as gemsbok cucumbers,

watermelon The watermelon (''Citrullus lanatus'') is a species of flowering plant in the family Cucurbitaceae, that has a large, edible fruit. It is a Glossary of botanical terms#scandent, scrambling and trailing vine-like plant, and is plant breeding ...

and Kalahari sour grass.

Enemies and competitors

Lioness vs Leopard 9 July 2016 Latest Sightings 1

Across its range, the leopard coexists with a number of other large predators. In Africa, it is part of a large predator

guild A guild ( ) is an association of artisans and merchants who oversee the practice of their craft/trade in a particular territory. The earliest types of guild formed as organizations of tradespeople belonging to a professional association. They so ...

with lions, cheetahs, spotted and

brown hyena The brown hyena (''Parahyaena brunnea''), also called the strandwolf, is a species of hyena found in Namibia, Botswana, western and southern Zimbabwe, southern Mozambique and South Africa. It is the only extant species in the genus ''Parahyaena' ...

s, and

African wild dog The African wild dog (''Lycaon pictus''), also called painted dog and Cape hunting dog, is a wild canine native to sub-Saharan Africa. It is the largest wild canine in Africa, and the only extant member of the genus '' Lycaon'', which is disti ...

s. The leopard is dominant only over the cheetah while the others have the advantage of size, pack numbers or both. Lions pose a great mortal threat and can be responsible for 22% of leopard deaths in

Sabi Sand Game Reserve Sabi Sand Game Reserve consists of a group of private game reserves located adjacent to the Kruger National Park in the Lowveld of Mpumalanga, South Africa. Its name comes from the Sabie River on its southern boundary and the Sand River flow ...

. Spotted hyenas are less threatening but are more likely to steal kills, being the culprits of up to 50% of stolen leopard kills in the same area. To counter this, leopards store their kills in the trees and out of reach. Lions have a high success rate in fetching leopard kills from trees. Leopards do not seem to actively avoid their competitors but rather difference in prey and habitat preferences appear to limit their spatial overlap. In particular, leopards use heavy vegetation regardless of whether lions are present in an area and both cats are active at the same time of day. In Asia, the leopard's main competitors are tigers and

dhole The dhole ( ; ''Cuon alpinus'') is a canid native to South, East and Southeast Asia. It is anatomically distinguished from members of the genus ''Canis'' in several aspects: its skull is convex rather than concave in profile, it lacks a third ...

s. Both the larger tiger and pack-living dhole dominate leopards during encounters. Interactions between the three predators involve chasing, stealing kills and direct killing. Tigers appear to inhabit the deep parts of the forest while leopards and dholes are pushed closer to the fringes. The three predators coexist by hunting different sized prey. In Nagarhole National Park, the average size for a leopard kill was compared to for tigers and for dholes. At Kui Buri National Park, following a reduction in prey numbers, tigers continued to feed on favoured prey while leopards and dholes had to increase their consumption of small prey. Leopards can live successfully in tiger habitat when there is abundant food and vegetation cover. Otherwise, they appear to be less common where tigers are numerous. The recovery of the tiger population in Rajaji National Park during the 2000s led to a reduction in leopard population densities.

Reproduction and life cycle

In some areas, leopards mate all year round. In Manchuria and Siberia, they mate during January and February. On average, females begin to breed between the ages of 2½ and three, and males between the ages of two and three. The female's estrous cycle lasts about 46 days, and she is usually in heat for 6–7 days. Gestation lasts for 90 to 105 days. Cubs are usually born in a Litter (zoology), litter of 2–4 cubs. The mortality rate of cubs is estimated at 41–50% during the first year. Predators are the biggest cause for leopard cub mortality during their first year. Male leopards are known to cause infanticide (zoology), infanticide, in order to bring the female back into heat. Intervals between births average 15 to 24 months, but can be shorter, depending on the survival of the cubs. Females give birth in a cave, crevice among boulders, hollow tree or thicket. Newborn cubs weigh , and are born with closed eyes, which open four to nine days after birth. The fur of the young tends to be longer and thicker than that of adults. Their pelage is also more gray in colour with less defined spots. They begin to eat meat at around nine weeks. Around three months of age, the young begin to follow the mother on hunts. At one year of age, cubs can probably fend for themselves, but will remain with the mother for 18–24 months. After separating from their mother, sibling cubs may travel together for months. Both male and female leopards typically reach sexual maturity at 2–2⅓ years. The Generation time, generation length of the leopard is 9.3 years. The average life span of a leopard is 12–17 years. The oldest leopard was a captive female that died at the age of 24 years, 2 months and 13 days.

Conservation

The leopard is listed on CITES Appendix I, and hunting is banned in Botswana and Afghanistan; in 11 sub-Saharan countries, trade is restricted to skins and body parts of 2,560 individuals. In 2007, a leopard reintroduction programme was initiated in the Russian Caucasus, where captive bred individuals are reared and trained in large enclosures in Sochi National Park; six individuals released into Caucasus Nature Reserve and Alaniya National Park in 2018 survived as of February 2022.

Threats

The leopard is primarily threatened by habitat fragmentation and conversion of forest to agriculturally used land, which lead to a declining natural prey base, human–wildlife conflict with livestock herders and high leopard mortality rates. It is also threatened by trophy hunting and poaching. Contemporary records suggest that the leopard occurs in only 25% of its historical range. Between 2002 and 2012, at least four leopards were estimated to have been poached per week in India for the illegal wildlife trade of its skins and bones. In spring 2013, 37 leopard skins were found during a 7-week long market survey in major Moroccan cities. In 2014, 43 leopard skins were detected during two surveys in Morocco. Vendors admitted to have imported skins from sub-Saharan Africa. Surveys in the Central African Republic's Chinko area revealed that the leopard population decreased from 97 individuals in 2012 to 50 individuals in 2017. In this period, transhumant Pastoralism, pastoralists from the border area with Sudan moved in the area with their livestock. Rangers confiscated large amounts of poison in the camps of livestock herders who were accompanied by armed merchants. They engaged in poaching large herbivores, sale of bushmeat and trading leopard skins in Am Dafok. In Java, the leopard is threatened by illegal hunting and trade. Between 2011 and 2019, body parts of 51 Javan leopards were seized including six live individuals, 12 skins, 13 skulls, 20 canines and 22 claws.

Human relations

Cultural significance

Leopards have been featured in art, mythology and folklore of many countries. In Greek mythology, it was a symbol of the god Dionysus, who was depicted wearing leopard skin and using leopards as means of transportation. In one myth, the god was captured by pirates but two leopards rescued him. Numerous Roman mosaics from North African sites depict fauna now found only in tropical Africa. During the Benin Empire, the leopard was commonly represented on engravings and sculptures and was used to symbolise the power of the king or ''Oba (ruler), oba'', since the leopard was considered the king of the forest. The Ashanti people also used the leopard as a symbol of leadership, and only the king was permitted to have a ceremonial leopard stool. Some African cultures considered the leopard to be a smarter, better hunter than the lion and harder to kill. In Rudyard Kipling's "How the Leopard Got His Spots", one of his ''Just So Stories'', a leopard with no spots in the Highveld lives with his hunting partner, the Ethiopian. When they set off to the forest, the Ethiopian changed his brown skin, and the leopard painted spots on his skin. A leopard played an important role in the 1938 Hollywood film ''Bringing Up Baby''. African chiefs, European queens, Hollywood actors and burlesque dancers wore coats made of leopard skins. The leopard is a frequently used motif in Leopard (heraldry), heraldry, most commonly as ''Attitude (heraldry)#Passant, passant''. The heraldic leopard lacks spots and sports a mane, making it visually almost identical to the Lion (heraldry), heraldic lion, and the two are often used interchangeably. Naturalistic leopard-like depictions appear on the coat of arms of Benin, Coat of arms of Malawi, Malawi, Coat of arms of Somalia, Somalia, the Coat of arms of the Democratic Republic of the Congo, Democratic Republic of the Congo and Coat of arms of Gabon, Gabon, the last of which uses a black panther.

Attacks on people

The Leopard of Rudraprayag killed more than 125 people; the Panar Leopard was thought to have killed over 400 people. Both were shot by British hunter Jim Corbett. The ''spotted devil of Gummalapur'' killed about 42 people in Karnataka, India.

In captivity

The ancient Romans kept leopards in captivity to be slaughtered in Venatio, hunts as well as Damnatio ad bestias, execute criminals. In Benin, leopards were kept and paraded as mascots, totems and sacrifices to deities. Several leopards were kept in a menagerie originally established by King John of England at the Tower of London in the 13th century; around 1235, three of these animals were given to Henry III of England, Henry III by Holy Roman Emperor Frederick II, Holy Roman Emperor, Frederick II. In modern times, leopards have been Animal training, trained and Tame animal, tamed in circuses.

References

External links

IUCN/SSC Cat Specialist Group: ''Panthera pardus'' in Africa
an
''Panthera pardus'' in Asia
* {{DEFAULTSORT:Leopard Leopards, Articles containing video clips Big cats Felids of Africa Felids of Asia Mammals described in 1758 National symbols of Benin National symbols of Malawi National symbols of Somalia National symbols of the Democratic Republic of the Congo Panthera Taxa named by Carl Linnaeus Apex predators

Etymology

Taxonomy

Living subspecies

Evolution

Hybrids

Characteristics

Size

Distribution and habitat

Behaviour and ecology

Social spacing

Hunting and diet

Enemies and competitors

Reproduction and life cycle

Conservation

Threats

Human relations

Cultural significance

Attacks on people

In captivity

See also

References

Further reading

External links