The large-tailed nightjar (''Caprimulgus macrurus'') is a species of
nightjar
Nightjars are medium-sized nocturnal or crepuscular birds in the family Caprimulgidae and order Caprimulgiformes, characterised by long wings, short legs, and very short bills. They are sometimes called bugeaters, their primary source of food ...
known for its distinctive long, broad tail and cryptic plumage that allows it to blend into its surroundings. Found along the southern
Himalayan foothills, eastern
South Asia
South Asia is the southern Subregion#Asia, subregion of Asia that is defined in both geographical and Ethnicity, ethnic-Culture, cultural terms. South Asia, with a population of 2.04 billion, contains a quarter (25%) of the world's populatio ...
,
Southeast Asia
Southeast Asia is the geographical United Nations geoscheme for Asia#South-eastern Asia, southeastern region of Asia, consisting of the regions that are situated south of China, east of the Indian subcontinent, and northwest of the Mainland Au ...
and northern
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
, its natural
habitat
In ecology, habitat refers to the array of resources, biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species' habitat can be seen as the physical manifestation of its ...
s are
subtropical or tropical moist lowland forest
Tropical and subtropical moist broadleaf forests (TSMF), also known as tropical moist forest, is a subtropical and tropical forest habitat type defined by the World Wide Fund for Nature (WWF).
Description
TSMF is generally found in large ...
, subtropical or tropical
mangrove forest
Mangrove forests, also called mangrove swamps, mangrove thickets or mangals, are productive wetlands that occur in coastal intertidal zones. Mangrove forests grow mainly at tropical and subtropical latitudes because mangrove trees cannot withsta ...
, and subtropical or tropical moist
montane forest
Montane ecosystems are found on the slopes of mountains. The alpine climate in these regions strongly affects the ecosystem because temperatures lapse rate, fall as elevation increases, causing the ecosystem to stratify. This stratification is ...
.
Description
The large-tailed nightjar measures approximately 31.5 to 33 cm in length.
The cryptic
plumage
Plumage () is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, there can b ...
and colouration of large-tailed nightjars provide effective
camouflage
Camouflage is the use of any combination of materials, coloration, or illumination for concealment, either by making animals or objects hard to see, or by disguising them as something else. Examples include the leopard's spotted coat, the b ...
within its natural habitat.
Males exhibit distinctive markings, characterized by a pale crown with a dark median stripe. A prominent row of black
scapular feathers
The following is a list of terms used in bird topography:
Plumage features
* Glossary_of_bird_terms#B, Back
* Abdomen#Vertabrates, Belly
* Breast
* Cheek
* Chin
* Crest (feathers), Crest
* Crown (anatomy), Crown
* Crown patch
* Ear-coverts
* ...
, bordered by broad buff or whitish-buff fringes, enhances their striking appearance.
Additionally, males possess a white throat patch and along, broad brown tail marked with uneven dark bars.
Large white patches on its
primaries
Primary elections or primaries are elections held to determine which candidates will run in an upcoming general election. In a partisan primary, a political party selects a candidate. Depending on the state and/or party, there may be an "open pri ...
and outer tail feathers can be observed in flight.
In contrast, females display a paler and greyer plumage compared to males. They have smaller buff-coloured wing patches and less vibrant tail patches, resulting in a drabber appearance.
Taxonomy
The large-tailed nightjar (''Caprimulgus macrurus'') is a species of nightjar belonging to the family
Caprimulgidae. There are 6
subspecies
In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...
presently recognized (''C.m. albonotatus, C.m. bimaculatus, C.m. macrurus, C.m. salvadorii, C.m. johnsoni, C.m. schlegelii'').
Originally a
species complex
In biology, a species complex is a group of closely related organisms that are so similar in appearance and other features that the boundaries between them are often unclear. The taxa in the complex may be able to hybridize readily with each oth ...
of 12 different subspecies, differences in territorial
vocalizations have been interpreted as evidence for
reproductive isolation
The mechanisms of reproductive isolation are a collection of evolutionary mechanisms, ethology, behaviors and physiology, physiological processes critical for speciation. They prevent members of different species from producing offspring, or ensu ...
and, consequently, the delegation of some members of the complex as new species of nightjar under the
biological species concept
A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of classification and ...
.
Habitat and distribution
The large-tailed nightjar is found across a diverse range of habitats within
subtropical
The subtropical zones or subtropics are geographical zone, geographical and Köppen climate classification, climate zones immediately to the Northern Hemisphere, north and Southern Hemisphere, south of the tropics. Geographically part of the Ge ...
and
tropical zones. This species is a resident of the countries of
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
,
Bangladesh
Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...
,
Bhutan
Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...
,
Brunei
Brunei, officially Brunei Darussalam, is a country in Southeast Asia, situated on the northern coast of the island of Borneo. Apart from its coastline on the South China Sea, it is completely surrounded by the Malaysian state of Sarawak, with ...
,
Cambodia
Cambodia, officially the Kingdom of Cambodia, is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. It is bordered by Thailand to the northwest, Laos to the north, and Vietnam to the east, and has a coastline ...
,
China
China, officially the People's Republic of China (PRC), is a country in East Asia. With population of China, a population exceeding 1.4 billion, it is the list of countries by population (United Nations), second-most populous country after ...
,
India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
,
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
,
Laos
Laos, officially the Lao People's Democratic Republic (LPDR), is the only landlocked country in Southeast Asia. It is bordered by Myanmar and China to the northwest, Vietnam to the east, Cambodia to the southeast, and Thailand to the west and ...
,
Malaysia
Malaysia is a country in Southeast Asia. Featuring the Tanjung Piai, southernmost point of continental Eurasia, it is a federation, federal constitutional monarchy consisting of States and federal territories of Malaysia, 13 states and thre ...
,
Myanmar
Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...
,
Nepal
Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...
,
Pakistan
Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...
,
Papua New Guinea
Papua New Guinea, officially the Independent State of Papua New Guinea, is an island country in Oceania that comprises the eastern half of the island of New Guinea and offshore islands in Melanesia, a region of the southwestern Pacific Ocean n ...
,
Philippines
The Philippines, officially the Republic of the Philippines, is an Archipelagic state, archipelagic country in Southeast Asia. Located in the western Pacific Ocean, it consists of List of islands of the Philippines, 7,641 islands, with a tot ...
,
Singapore
Singapore, officially the Republic of Singapore, is an island country and city-state in Southeast Asia. The country's territory comprises one main island, 63 satellite islands and islets, and one outlying islet. It is about one degree ...
,
Thailand
Thailand, officially the Kingdom of Thailand and historically known as Siam (the official name until 1939), is a country in Southeast Asia on the Mainland Southeast Asia, Indochinese Peninsula. With a population of almost 66 million, it spa ...
,
Timor-Leste
Timor-Leste, also known as East Timor, officially the Democratic Republic of Timor-Leste, is a country in Southeast Asia. It comprises the eastern half of the island of Timor, the coastal exclave of Oecusse in the island's northwest, and ...
and
Vietnam
Vietnam, officially the Socialist Republic of Vietnam (SRV), is a country at the eastern edge of mainland Southeast Asia, with an area of about and a population of over 100 million, making it the world's List of countries and depende ...
.
Its distribution spans a wide geographical area, extending from
northern India
North India is a geographical region, loosely defined as a cultural region comprising the northern part of India (or historically, the Indian subcontinent) wherein Indo-Aryans (speaking Indo-Aryan languages) form the prominent majority populati ...
through
eastern China
East China () is a geographical region in the People’s Republic of China, mainly consisting of seven province-level administrative divisions, namely the provinces (from north to south) Shandong, Jiangsu, Anhui, Zhejiang, Jiangxi, Fujian, a ...
and
Indonesia
Indonesia, officially the Republic of Indonesia, is a country in Southeast Asia and Oceania, between the Indian Ocean, Indian and Pacific Ocean, Pacific oceans. Comprising over List of islands of Indonesia, 17,000 islands, including Sumatra, ...
to northern
Australia
Australia, officially the Commonwealth of Australia, is a country comprising mainland Australia, the mainland of the Australia (continent), Australian continent, the island of Tasmania and list of islands of Australia, numerous smaller isl ...
.
This species commonly inhabits
secondary forest
A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
s,
forest edges,
plantation
Plantations are farms specializing in cash crops, usually mainly planting a single crop, with perhaps ancillary areas for vegetables for eating and so on. Plantations, centered on a plantation house, grow crops including cotton, cannabis, tob ...
s, and wooded parks.
It has also been observed in agricultural landscapes, where its presence may be of significant economic interest to farmers, as large-tailed nightjars are believed to play a role in the
biological control
Biological control or biocontrol is a method of controlling pests, whether pest animals such as insects and mites, weeds, or pathogens affecting animals or plants by using other organisms. It relies on predation, parasitism, herbivory, or o ...
of insect pests.
Behaviour
The large-tailed nightjar's remarkable camouflage and
nocturnal
Nocturnality is a ethology, behavior in some non-human animals characterized by being active during the night and sleeping during the day. The common adjective is "nocturnal", versus diurnality, diurnal meaning the opposite.
Nocturnal creatur ...
lifestyle pose significant challenges for studying their behaviour and
ecology
Ecology () is the natural science of the relationships among living organisms and their Natural environment, environment. Ecology considers organisms at the individual, population, community (ecology), community, ecosystem, and biosphere lev ...
. Similar to other nightjar species, they depend primarily on
acoustic communication rather than visual cues for
inter-specific and
intra-specific communication.
Vocalizations
Songs can be heard year-round, but vocal activity may vary depending on breeding activity,
population density
Population density (in agriculture: Standing stock (disambiguation), standing stock or plant density) is a measurement of population per unit land area. It is mostly applied to humans, but sometimes to other living organisms too. It is a key geog ...
, and
moon phase. The large-tailed nightjar has a distinctive, monotonous song that it uses to assert its territory.
Territorial songs are typically given by males, but there are also reports of females giving these songs. Singing is most notable during
courtship
Courtship is the period wherein some couples get to know each other prior to a possible marriage or committed romantic, ''de facto'' relationship. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marri ...
and early breeding, primarily taking place from perches or occasionally in pursuit-flights.
Long-tailed nightjars also perform grunting calls, which are very different from territorial songs. They are best described as a series of deep, low-pitched "''wroww''" calls or a series of fast-repeated "''wrrrog''" or "''grrrog''" notes.
Diet
Large-tailed nightjars are
aerial insectivores. Their diets are primarily composed of
beetle
Beetles are insects that form the Taxonomic rank, order Coleoptera (), in the superorder Holometabola. Their front pair of wings are hardened into wing-cases, elytra, distinguishing them from most other insects. The Coleoptera, with about 40 ...
s,
moth
Moths are a group of insects that includes all members of the order Lepidoptera that are not Butterfly, butterflies. They were previously classified as suborder Heterocera, but the group is Paraphyly, paraphyletic with respect to butterflies (s ...
s, and other
flying insects consumed while the bird is in flight. However, there have been a few observations of the large-tailed nightjar walking and pecking at the ground, apparently preying on
terrestrial
Terrestrial refers to things related to land or the planet Earth, as opposed to extraterrestrial.
Terrestrial may also refer to:
* Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on o ...
ant
Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...
s. It has been suggested that, while ground
foraging
Foraging is searching for wild food resources. It affects an animal's fitness because it plays an important role in an animal's ability to survive and reproduce. Foraging theory is a branch of behavioral ecology that studies the foraging behavi ...
is not common, it may occur more frequently than previously thought, as a response to low abundance of flying insects.
Reproduction
Little is known about the large-tailed nightjar's nesting and roosting behaviour. The
breeding season
Seasonal breeders are animal species that successfully mate only during certain times of the year. These times of year allow for the optimization of survival of young due to factors such as ambient temperature, food and water availability, and ch ...
most likely extends from mid-March to late September. The female does not build a nest and instead lays 1-2 eggs in a shallow depression of
leaf litter
Plant litter (also leaf litter, tree litter, soil litter, litterfall, or duff) is dead plant material (such as leaves, bark, needles, twigs, and cladodes) that has fallen to the ground. This detritus or dead organic material and its constituen ...
on the ground.
The eggs are pinkish-buff to pale yellow in colour and chicks have mealy brown feathers that ensure good camouflage in their nest site. Nightjar chicks emerge with
downy feathers and may be moved from the nest site 1-2 days after hatching. Chicks move every night and siblings will sometimes separate, likely to avoid detection by
predators
Predation is a biological interaction in which one organism, the predator, kills and eats another organism, its prey. It is one of a family of common feeding behaviours that includes parasitism and micropredation (which usually do not kill ...
.
References
External links
Large-tailed Nightjar Sounds & CallsLarge-tailed Nightjar Xeno-Canto ID''Caprimulgus macrurus'' IUCN Red List
Gallery
File:Large tailed Night I IMG 8079.jpg, at Bharatpur, Rajasthan
Rajasthan (; Literal translation, lit. 'Land of Kings') is a States and union territories of India, state in northwestern India. It covers or 10.4 per cent of India's total geographical area. It is the List of states and union territories of ...
, India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
.
File:Large-tailed Nightjar female C. m. bimaculatus I IMG 9628.jpg, Female C. m. bimaculatus in Narendrapur near Kolkata
Kolkata, also known as Calcutta ( its official name until 2001), is the capital and largest city of the Indian state of West Bengal. It lies on the eastern bank of the Hooghly River, west of the border with Bangladesh. It is the primary ...
, West Bengal
West Bengal (; Bengali language, Bengali: , , abbr. WB) is a States and union territories of India, state in the East India, eastern portion of India. It is situated along the Bay of Bengal, along with a population of over 91 million inhabi ...
, India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
.
File:Large tailed Nightjar I IMG 8461.jpg, at Bharatpur, Rajasthan
Rajasthan (; Literal translation, lit. 'Land of Kings') is a States and union territories of India, state in northwestern India. It covers or 10.4 per cent of India's total geographical area. It is the List of states and union territories of ...
, India
India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...
.
File:Large tailed nightjar.jpg, Large-tailed nightjar in Bharatpur.
File:Large Tailed Nightjar.jpg, Large-tailed nightjar at Chintamani Kar Bird Sanctuary, West Bengal, India
large-tailed nightjar
The large-tailed nightjar (''Caprimulgus macrurus'') is a species of nightjar known for its distinctive long, broad tail and cryptic plumage that allows it to blend into its surroundings. Found along the southern Himalayan foothills, eastern So ...
Birds of Southeast Asia
large-tailed nightjar
The large-tailed nightjar (''Caprimulgus macrurus'') is a species of nightjar known for its distinctive long, broad tail and cryptic plumage that allows it to blend into its surroundings. Found along the southern Himalayan foothills, eastern So ...
Taxonomy articles created by Polbot
{{caprimulgiformes-stub