''Lansium parasiticum'', commonly known as langsat (),
lanzones (), or longkong in English; duku in
Indonesian
Indonesian is anything of, from, or related to Indonesia, an archipelagic country in Southeast Asia. It may refer to:
* Indonesians, citizens of Indonesia
** Native Indonesians, diverse groups of local inhabitants of the archipelago
** Indonesia ...
or dokong in
Terengganu Malay, is a species of tree in the
Mahogany
Mahogany is a straight- grained, reddish-brown timber of three tropical hardwood species of the genus '' Swietenia'', indigenous to the AmericasBridgewater, Samuel (2012). ''A Natural History of Belize: Inside the Maya Forest''. Austin: Un ...
family with commercially cultivated edible fruits. The species is native to Southeast Asia.
[
]
Description
The tree is average sized, reaching in height and in diameter. 30 years old trees grown from seed and planted at 8 × 8 meter spacing can have a height of 10 meters and diameter of 25 cm. The trunk grows in an irregular manner, with its buttress root
Buttress roots also known as plank roots are large, wide roots on all sides of a shallowly rooted tree. Typically, they are found in nutrient-poor tropical forest soils that may not be very deep. They prevent the tree from falling over (hence t ...
s showing above ground. The tree's bark is a greyish colour, with light and dark spots. Its resin is thick and milk coloured.
The pinnately compound
Pinnation (also called pennation) is the arrangement of feather-like or multi-divided features arising from both sides of a common axis. Pinnation occurs in biological morphology, in crystals, such as some forms of ice or metal crystals, and in ...
leaves are odd numbered, with thin hair, and 6 to 9 buds at intervals. The buds are long and elliptical, approximately by in size. The upper edge shines, and the leaves themselves have pointed bases and tips. The stems of the buds measure .[
The flowers are located in ]inflorescence
An inflorescence is a group or cluster of flowers arranged on a stem that is composed of a main branch or a complicated arrangement of branches. Morphologically, it is the modified part of the shoot of seed plants where flowers are formed ...
s that grow and hang from large branches or the trunk; the bunches may number up to 5 in one place. They are often branched at their base, measure in size, and have short fur. The flowers are small, with short stems, and have two genders
Gender is the range of characteristics pertaining to femininity and masculinity and differentiating between them. Depending on the context, this may include sex-based social structures (i.e. gender roles) and gender identity. Most cultures u ...
. The sheathe is shaped like a five lobed cup and is coloured a greenish-yellow. The corona is egg-shaped and hard, measuring by . There is one stamen
The stamen (plural ''stamina'' or ''stamens'') is the pollen-producing reproductive organ of a flower. Collectively the stamens form the androecium., p. 10
Morphology and terminology
A stamen typically consists of a stalk called the filam ...
, measuring in length. The top of the stamen is round. The pistil
Gynoecium (; ) is most commonly used as a collective term for the parts of a flower that produce ovules and ultimately develop into the fruit and seeds. The gynoecium is the innermost whorl of a flower; it consists of (one or more) '' pis ...
is short and thick.[
The fruit can be elliptical, ovoid or round, measuring by in size. Fruits look much like small potatoes and are borne in clusters similar to grapes. The larger fruits are on the variety known as duku. It is covered by thin, yellow hair giving a slightly fuzzy aspect. The skin thickness varies with the varieties, from to approximately . With the skin removed, the fruit resembles peeled ]garlic
Garlic (''Allium sativum'') is a species of bulbous flowering plant in the genus ''Allium''. Its close relatives include the onion, shallot, leek, chive, Welsh onion and Chinese onion. It is native to South Asia, Central Asia and northe ...
in appearance, with half a dozen white, translucent lobes, some of which contain a flat, bitter tasting seed. The seeds are covered with a thick, clear-white aril that tastes sweet and sour.[ The taste has been likened to a combination of grape and grapefruit and is considered excellent by most. The sweet juicy flesh contains ]sucrose
Sucrose, a disaccharide, is a sugar composed of glucose and fructose subunits. It is produced naturally in plants and is the main constituent of white sugar. It has the molecular formula .
For human consumption, sucrose is extracted and refi ...
, fructose
Fructose, or fruit sugar, is a ketonic simple sugar found in many plants, where it is often bonded to glucose to form the disaccharide sucrose. It is one of the three dietary monosaccharides, along with glucose and galactose, that are absorb ...
, and glucose
Glucose is a simple sugar with the molecular formula . Glucose is overall the most abundant monosaccharide, a subcategory of carbohydrates. Glucose is mainly made by plants and most algae during photosynthesis from water and carbon dioxide, usi ...
. For consumption, cultivars with small or undeveloped seeds and thick aril are preferred.
Cultivars
There are numerous cultivar
A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s of ''L. parasiticum''. Overall, there are two main groups of cultivars, those named ''duku'' and those named ''langsat''. There are also mixed duku-langsat varieties.[
Those called duku generally have a large ]crown
A crown is a traditional form of head adornment, or hat, worn by monarchs as a symbol of their power and dignity. A crown is often, by extension, a symbol of the monarch's government or items endorsed by it. The word itself is used, partic ...
, thick with bright green leaves, with short bunches of few fruit. The individual fruit are large, generally round, and have somewhat thick skin that does not release sap when cooked. The seeds are small, with thick flesh, a sweet scent, and a sweet or sour taste.
Meanwhile, the variant commonly known as langsat generally has thinner trees, with a less dense crown consisting of dark green leaves and stiff branches. The bunches are longer, and each bunch holds between 15 and 25 large, egg-shaped fruit. The skin is thin and releases a white sap when cooked. The flesh is watery and tastes sweet and sour.[
]
''Lansium domesticum'' var. ''aquaeum'' is distinguished by its hairy leaves, as well as the tightly packed dark yellow fruit on its bunches. The fruit tends to be small, with thin skin and little sap; the skin is difficult to remove. To be eaten, the fruit is bitten and the flesh sucked through the hole created,[ or rubbed until the skin breaks and the seeds are retrieved. In Indonesia the fruit has several names, including ''kokosan'', ''pisitan'', ''pijetan'', and ''bijitan''.][ The seeds are relatively large, with thin, sour flesh.
]
Reproduction
The seeds of ''L. parasiticum'' are polyembryonic, the multiple embryos resulting from apomixis
In botany, apomixis is asexual reproduction without fertilization. Its etymology is Greek for "away from" + "mixing". This definition notably does not mention meiosis. Thus "normal asexual reproduction" of plants, such as propagation from cutt ...
.
''Lansium parasiticum'' is traditionally reproduced by spreading seedlings, either cultivated or collected from below the tree.[ It has been said that new seedlings require 20 to 25 years to bear fruit, with the possibility of the quality being inferior.] However other sources quote 12 years to first production from seed and no variations. Production often varies from year to year, and depends to some extent on having a dry period to induce flowering. One example of ten trees in Costa Rica about twenty-five years old produced during five years the following weights of salable fruits: 2008: 50 kilos, 2009: 2000 kilos, 2010: 1000 kilos, 2011: 100 kilos, 2012: 1500 kilos. Experiments in the Philippines with grafting where two trees are planted close to each other and then grafted when one to two meters tall to leave twin root systems on a single main trunk have resulted in earlier and less erratic fruit production.
Another common method is by air layering
Layering has evolved as a common means of vegetative propagation of numerous species in natural environments. Layering is also utilized by horticulturists to propagate desirable plants.
Natural layering typically occurs when a branch touches ...
. Although the process requires up to several months,[ the new rooted tree produced is itself ready to bear fruit within two years. Trees cultivated with this method have a high death rate,][ and the growths are less resilient.][
The third common way to reproduce ''L. parasiticum'' is with ]grafting
Grafting or graftage is a horticultural technique whereby tissues of plants are joined so as to continue their growth together. The upper part of the combined plant is called the scion () while the lower part is called the rootstock. The succ ...
. This results in the new trees having the same genetic characteristics as their parent, and being ready to bear fruit within 5 to 6 years. The offspring are relatively stronger than transplanted shoots.[
]
Ecology
''Lansium parasiticum'' grows well in mixed agroforests. The plant, especially the ''duku'' variant, prefers damp, shaded areas. It can be grown in the same agroforest as durian
The durian (, ) is the edible fruit of several tree species belonging to the genus ''Durio''. There are 30 recognised ''Durio'' species, at least nine of which produce edible fruit. '' Durio zibethinus'', native to Borneo and Sumatra, is the o ...
, petai, and jengkol, as well as wood-producing trees.
''Lansium parasiticum'' is grown from low grounds up to heights of above sea level, in areas with an average rainfall of a year. The plant can grow and blossom in latosol Latosols, also known as tropical red earth, are soils found under tropical rainforests which have a relatively high content of iron and aluminium oxides. They are typically classified as oxisols (USDA soil taxonomy) or ferralsols (World Reference B ...
, yellow podzol, and alluvium
Alluvium (from Latin ''alluvius'', from ''alluere'' 'to wash against') is loose clay, silt, sand, or gravel that has been deposited by running water in a stream bed, on a floodplain, in an alluvial fan or beach, or in similar settings. Alluv ...
.[ The plant prefers slightly acidic soil with good drainage and rich in mulch. The ''langsat'' variant is hardier, and can weather dry seasons with a little shade and water.][ The plant cannot handle floods.]
''Lansium parasiticum'' generally bears fruit once a year. This period can vary between areas, but blooming is generally after the beginning of the rainy season and fruit production some four months later.
Distribution
''Lansium parasiticum'' is native to the Malesian phytochorion
A phytochorion, in phytogeography, is a geographic area with a relatively uniform composition of plant species. Adjacent phytochoria do not usually have a sharp boundary, but rather a soft one, a transitional area in which many species from both r ...
of Southeast Asia, from Peninsular Thailand
Southern Thailand, Southern Siam or Tambralinga is a southernmost cultural region of Thailand, separated from Central Thailand region by the Kra Isthmus.
Geography
Southern Thailand is on the Malay Peninsula, with an area of around , bound ...
and Peninsular Malaysia
Peninsular Malaysia ( ms, Semenanjung Malaysia; Jawi: سمننجڠ مليسيا), or the States of Malaya ( ms, Negeri-negeri Tanah Melayu; Jawi: نڬري-نڬري تانه ملايو), also known as West Malaysia or the Malaysian Peninsula, ...
, to Java
Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
, Sumatra, Borneo
Borneo (; id, Kalimantan) is the List of islands by area, third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java Isl ...
, the Philippines
The Philippines (; fil, Pilipinas, links=no), officially the Republic of the Philippines ( fil, Republika ng Pilipinas, links=no),
* bik, Republika kan Filipinas
* ceb, Republika sa Pilipinas
* cbk, República de Filipinas
* hil, Republ ...
(Luzon
Luzon (; ) is the largest and most populous island in the Philippines. Located in the northern portion of the Philippines archipelago, it is the economic and political center of the nation, being home to the country's capital city, Manila, ...
, Camiguin
Camiguin, officially the Province of Camiguin ( ceb, Probinsya sa Camiguin; tl, Lalawigan ng Camiguin; Kamigin: ''Probinsya ta Kamigin''), is an island province in the Philippines located in the Bohol Sea, about off the northern coast of Min ...
, Basilan
Basilan, officially the Province of Basilan ( cbk, Provincia de Basilan; yka, Wilayah Basilanin; tsg, Wilaya' sin Basilan; fil, Lalawigan ng Basilan), is an island province of the Philippines located primarily in the Bangsamoro Autonomous Re ...
, and