The laggar falcon (''Falco jugger''), also known as the lugger falcon or jugger (from

Hindi Modern Standard Hindi (, ), commonly referred to as Hindi, is the Standard language, standardised variety of the Hindustani language written in the Devanagari script. It is an official language of India, official language of the Government ...

जग्गर — jaggar, "falcon"), is a mid-sized

bird of prey Birds of prey or predatory birds, also known as (although not the same as) raptors, are hypercarnivorous bird species that actively predation, hunt and feed on other vertebrates (mainly mammals, reptiles and smaller birds). In addition to speed ...

which occurs in the Indian subcontinent from extreme southeastern

Iran Iran, officially the Islamic Republic of Iran (IRI) and also known as Persia, is a country in West Asia. It borders Iraq to the west, Turkey, Azerbaijan, and Armenia to the northwest, the Caspian Sea to the north, Turkmenistan to the nort ...

, southeastern

Afghanistan Afghanistan, officially the Islamic Emirate of Afghanistan, is a landlocked country located at the crossroads of Central Asia and South Asia. It is bordered by Pakistan to the Durand Line, east and south, Iran to the Afghanistan–Iran borde ...

Pakistan Pakistan, officially the Islamic Republic of Pakistan, is a country in South Asia. It is the List of countries and dependencies by population, fifth-most populous country, with a population of over 241.5 million, having the Islam by country# ...

, through

India India, officially the Republic of India, is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area; the List of countries by population (United Nations), most populous country since ...

Nepal Nepal, officially the Federal Democratic Republic of Nepal, is a landlocked country in South Asia. It is mainly situated in the Himalayas, but also includes parts of the Indo-Gangetic Plain. It borders the Tibet Autonomous Region of China Ch ...

Bhutan Bhutan, officially the Kingdom of Bhutan, is a landlocked country in South Asia, in the Eastern Himalayas between China to the north and northwest and India to the south and southeast. With a population of over 727,145 and a territory of , ...

Bangladesh Bangladesh, officially the People's Republic of Bangladesh, is a country in South Asia. It is the List of countries and dependencies by population, eighth-most populous country in the world and among the List of countries and dependencies by ...

and northwestern

Myanmar Myanmar, officially the Republic of the Union of Myanmar; and also referred to as Burma (the official English name until 1989), is a country in northwest Southeast Asia. It is the largest country by area in Mainland Southeast Asia and has ...

. It resembles the

lanner falcon The lanner falcon (''Falco biarmicus'') is a medium-sized bird of prey that breeds in Africa, southeast Europe and just into Asia. It prefers open habitat and is mainly resident, but some birds disperse more widely after the breeding season. A l ...

but is darker overall, and has blackish "trousers" (

tibiotarsus The tibiotarsus is the large bone between the femur and the tarsometatarsus in the leg of a bird. It is the fusion of the proximal part of the tarsus with the tibia. A similar structure also occurred in the Mesozoic Heterodontosauridae. These ...

feathers). Fledglings have an almost entirely dark underside, and first-year subadult birds still retain much dark on the belly. This

species A species () is often defined as the largest group of organisms in which any two individuals of the appropriate sexes or mating types can produce fertile offspring, typically by sexual reproduction. It is the basic unit of Taxonomy (biology), ...

belongs to a close-knit complex of

falcon Falcons () are birds of prey in the genus ''Falco'', which includes about 40 species. Some small species of falcons with long, narrow wings are called hobbies, and some that hover while hunting are called kestrels. Falcons are widely distrib ...

s known as

hierofalcon The hierofalcons are four closely related species of falcon which make up the subgenus ''Hierofalco'': * Lanner falcon, ''Falco biarmicus'' * Laggar falcon, ''Falco jugger'' * Saker falcon, ''Falco cherrug'' * Gyrfalcon, ''Falco rusticolus'' ...

s. In this group, there is ample evidence for rampant hybridization and incomplete lineage sorting which confounds analyses of

DNA sequence A nucleic acid sequence is a succession of bases within the nucleotides forming alleles within a DNA (using GACT) or RNA (GACU) molecule. This succession is denoted by a series of a set of five different letters that indicate the order of the nu ...

data to a massive extent; molecular studies with small

sample size Sample size determination or estimation is the act of choosing the number of observations or replicates to include in a statistical sample. The sample size is an important feature of any empirical study in which the goal is to make inferences abo ...

s can simply not be expected to yield reliable conclusions in the entire hierofalcon group. The

radiation In physics, radiation is the emission or transmission of energy in the form of waves or particles through space or a material medium. This includes: * ''electromagnetic radiation'' consisting of photons, such as radio waves, microwaves, infr ...

of the entire living diversity of hierofalcons seems to have taken place in the Eemian interglacial at the start of the

Late Pleistocene The Late Pleistocene is an unofficial Age (geology), age in the international geologic timescale in chronostratigraphy, also known as the Upper Pleistocene from a Stratigraphy, stratigraphic perspective. It is intended to be the fourth division ...

, a mere 130,000-115,000 years ago; the laggar falcon represents a lineage that arrived at its present range out of eastern

Africa Africa is the world's second-largest and second-most populous continent after Asia. At about 30.3 million km2 (11.7 million square miles) including adjacent islands, it covers 20% of Earth's land area and 6% of its total surfac ...

by way of the

Arabian Peninsula The Arabian Peninsula (, , or , , ) or Arabia, is a peninsula in West Asia, situated north-east of Africa on the Arabian plate. At , comparable in size to India, the Arabian Peninsula is the largest peninsula in the world. Geographically, the ...

which during that time had a more humid climate than today.Helbig ''et al.'' (1994), Wink ''et al.'' (1998), Wink ''et al.'' (2004), Nittinger ''et al.'' (2005) Laggar Falcon juvenile plumage

Laggar falcons used to be the most common falcons in the region, but numbers have declined markedly in recent times and today it is probably nowhere a common species anymore. The main threats are the intensification of pesticide use in the region and use as a decoy to trap large falcons.

Gallery

File:AB011 Laggar Falcon.JPG, Adult bird,

Thar desert The Thar Desert (), also known as the Great Indian Desert, is an arid region in the north-western part of the Indian subcontinent that covers an area of in India and Pakistan. It is the world's 18th-largest desert, and the world's 9th-large ...

Rajasthan Rajasthan (; Literal translation, lit. 'Land of Kings') is a States and union territories of India, state in northwestern India. It covers or 10.4 per cent of India's total geographical area. It is the List of states and union territories of ...

(

). File:AB013 Laggar Falcon.JPG, Same as previous. File:AB012 Laggar Falcon.JPG, Same as previous; feeding on

Indian spiny-tailed lizard ''Saara hardwickii'', commonly known as Hardwicke's spiny-tailed lizard or the Indian spiny-tailed lizard is a species of lizard in the Family (biology), family Agamidae. The species is found in patches across the Thar Desert, Kutch, and surrou ...

(''Saara hardwickii''). File:Falcon prey (Spiny-tailed Lizard).JPG, The prey's spiny tail was not eaten. File:Laggar Falcon in Flight.jpg, Laggar falcon begins a swoop at

Tal Chhapar Sanctuary Tal Chhapar Sanctuary is a sanctuary located in the Churu district of Northwestern Rajasthan, in the Shekhawati region of India. It is known for blackbucks and is also home to a variety of birds. The sanctuary is 210 km from Jaipur on the ...

(

) File:Falco jugger MHNT 227 India.jpg, Egg of ''Falco jugger'' MHNT File:Laggar Falcon feeding on a spiny tailed lizard.jpg, Laggar falcon with a just caught Indian spiny-tailed lizard (''Saara hardwickii'') Laggar Falcon adult male

References

* Helbig, A.J.; Seibold, I.; Bednarek, W.; Brüning, H.; Gaucher, P.; Ristow, D.; Scharlau, W.; Schmidl, D. & Wink, Michael (1994): Phylogenetic relationships among falcon species (genus Falco) according to DNA sequence variation of the cytochrome b gene. ''In:'' Meyburg, B.-U. & Chancellor, R.D. (eds.): ''Raptor conservation today'': 593–599
PDF fulltext
* Nittinger, F.; Haring, E.; Pinsker, W.; Wink, Michael & Gamauf, A. (2005): Out of Africa? Phylogenetic relationships between ''Falco biarmicus'' and other hierofalcons (Aves Falconidae). ''Journal of Zoological Systematics and Evolutionary Research'' 43(4): 321–331. PDF fulltext
*Wink, Michael; Seibold, I.; Lotfikhah, F. & Bednarek, W. (1998): Molecular systematics of holarctic raptors (Order Falconiformes). ''In:'' Chancellor, R.D., Meyburg, B.-U. & Ferrero, J.J. (eds.): ''Holarctic Birds of Prey'': 29–48. Adenex & WWGBP
PDF fulltext
* Wink, Michael; Sauer-Gürth, Hedi; Ellis, David & Kenward, Robert (2004): Phylogenetic relationships in the Hierofalco complex (Saker-, Gyr-, Lanner-, Laggar Falcon). ''In:'' Chancellor, R.D. & Meyburg, B.-U. (eds.): ''Raptors Worldwide'': 499–504. WWGBP, Berlin
PDF fulltext

External links

* ttp://www.xeno-canto.org/species/Falco-jugger Sound recordings from Xeno-canto.org {{Taxonbar, from=Q845955 laggar falcon Birds of South Asia National symbols of India National symbols of Myanmar National symbols of Bangladesh National symbols of Bhutan National symbols of Nepal National symbols of Pakistan Birds of Myanmar laggar falcon laggar falcon