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The Javan rusa or Sunda sambar (''Rusa timorensis'') is a
deer Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the re ...
native to Indonesia and East Timor. Introduced populations exist in a wide variety of locations in the Southern Hemisphere.


Taxonomy

Seven subspecies of the Javan rusa are recognised: *''R. t. timorensis'' (Timor rusa deer) –
Timor Timor is an island at the southern end of Maritime Southeast Asia, in the north of the Timor Sea. The island is divided between the sovereign states of East Timor on the eastern part and Indonesia on the western part. The Indonesian part, ...
. *''R. t. djonga'' – Muna and Butung Islands. *''R. t. floresiensis'' (Flores rusa deer) –
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Including the Komodo Islands off its west coast (but excluding the Solor Archipelago to the east of Flores), the land area is 15,530.58 km2, and t ...
and other islands. *''R. t. macassaricus'' (Celebes rusa deer) – Celebes. *''R. t. moluccensis'' (Moluccan rusa deer) –
Molucca Islands The Maluku Islands (; Indonesian: ''Kepulauan Maluku'') or the Moluccas () are an archipelago in the east of Indonesia. Tectonically they are located on the Halmahera Plate within the Molucca Sea Collision Zone. Geographically they are located ea ...
. *''R. t. renschi'' –
Bali Bali () is a province of Indonesia and the westernmost of the Lesser Sunda Islands. East of Java and west of Lombok, the province includes the island of Bali and a few smaller neighbouring islands, notably Nusa Penida, Nusa Lembongan, and ...
. *''R. t. russa'' (Javan rusa deer) –
Java Java (; id, Jawa, ; jv, ꦗꦮ; su, ) is one of the Greater Sunda Islands in Indonesia. It is bordered by the Indian Ocean to the south and the Java Sea to the north. With a population of 151.6 million people, Java is the world's mo ...
.


Characteristics

The Javan rusa is dark blackish brown and has a grey forehead. Its back is almost black, the underparts and inner thighs are yellowish brown. The abdomen is lighter brown, and the tail tuft is dark blackish brown. The hair is coarse and longer on the chest than on the remaining body. Its ears are wide and a little shorter than the head. The
antler Antlers are extensions of an animal's skull found in members of the Cervidae (deer) family. Antlers are a single structure composed of bone, cartilage, fibrous tissue, skin, nerves, and blood vessels. They are generally found only on ...
s are medium long and rather wide, the upper branch points forward.
Fawn Deer or true deer are hoofed ruminant mammals forming the family Cervidae. The two main groups of deer are the Cervinae, including the muntjac, the elk (wapiti), the red deer, and the fallow deer; and the Capreolinae, including the reindee ...
s are born without spots. Males are bigger than females; head-to-body length varies from , with a tail. Males weigh , female about .


Distribution and habitat

The Javan rusa natively occurs on the islands of Java, Bali, and Timor in Indonesia. It has been introduced to Irian Jaya,
Borneo Borneo (; id, Kalimantan) is the third-largest island in the world and the largest in Asia. At the geographic centre of Maritime Southeast Asia, in relation to major Indonesian islands, it is located north of Java, west of Sulawesi, and e ...
, the Lesser Sunda Islands, Maluku,
Sulawesi Sulawesi (), also known as Celebes (), is an island in Indonesia. One of the four Greater Sunda Islands, and the world's eleventh-largest island, it is situated east of Borneo, west of the Maluku Islands, and south of Mindanao and the Sulu ...
, Pohnpei, Mauritius, Reunion, Fiji, Tonga, Samoa, Vanuatu, the Solomon Islands, the Christmas Islands, the Cocos Islands, Nauru, Australia, New Caledonia, New Zealand, Papua New Guinea, New Britain, and New Ireland. The Javan rusa was introduced by the Dutch to
New Guinea New Guinea (; Hiri Motu: ''Niu Gini''; id, Papua, or , historically ) is the world's second-largest island with an area of . Located in Oceania in the southwestern Pacific Ocean, the island is separated from Australia by the wide Torres ...
in the early 1900s. Since its introduction to the West Papuan lowlands, the species has become widely dispersed and is common in much of its new range. However its population in its native range has declined markedly by approximately 10,000 individuals in the past two decades and likely faces further decline. As a result it has been listed as vulnerable in its native range and was declared as a protected species under Indonesian law in 2018. During the 1980s and 1990s, protected areas such as national parks were used to control poaching and the effects of land conversion that destroyed grazing areas. However large numbers of Javan rusa died in Baluran National Park in Indonesia due to the loss of grazing area as a result of the invasive thorny acacia.


Ecology

Javan rusa are nocturnal, although they do graze during the day. They are rarely seen in the open and are very difficult to approach due to their keen senses and cautious instincts. The rusa deer is often found in small groups or pairs, although males are often seen alone. When alarmed, a rusa stag lets out an extremely loud honk. This is an alarm call and alerts any other deer in the vicinity. As with other deer species, Javan rusa mainly feed on grass, leaves, and fallen fruit. Most of their fluid requirements are met by the food they consume, so they hardly drink water.


Predators

The main predators of the Javan rusa include the Javan leopard, the dhole, crocodiles, pythons, and the
Komodo dragon The Komodo dragon (''Varanus komodoensis''), also known as the Komodo monitor, is a member of the monitor lizard family Varanidae that is endemic to the Indonesian islands of Komodo, Rinca, Flores, and Gili Motang. It is the largest extant ...
on the islands of Rinca,
Komodo Komodo may refer to: Computers * Komodo Edit, a free text editor for dynamic programming languages * Komodo IDE an integrated development environment (IDE) for dynamic programming languages * Komodo (chess), a chess engine People * Komo ...
, and
Flores Flores is one of the Lesser Sunda Islands, a group of islands in the eastern half of Indonesia. Including the Komodo Islands off its west coast (but excluding the Solor Archipelago to the east of Flores), the land area is 15,530.58 km2, and t ...
.


Reproduction

The Javan rusa mates around July and August, when stags contest by calling in a loud, shrill bark and dueling with the antlers. The doe gives birth to one or two calves after a gestation period of 8 months, at the start of spring. Calves are weaned at 6–8 months, and sexual maturity is attained at 3–5 years, depending on habitat conditions. Javan rusas live 15–20 years both in the wild and in captivity.


References


External links

* * * {{Taxonbar, from=Q663412 Javan rusa Mammals of Indonesia Fauna of Java Mammals of Borneo Vulnerable fauna of Asia Javan rusa Mammals of New Guinea