''Juniperus macrocarpa'' (large-fruited juniper, syn. ''J. oxycedrus'' subsp. ''macrocarpa'' (Sibth. & Sm.) Ball) is a species of

juniper Junipers are coniferous trees and shrubs in the genus ''Juniperus'' ( ) of the cypress family Cupressaceae. Depending on the taxonomy, between 50 and 67 species of junipers are widely distributed throughout the Northern Hemisphere as far south ...

, native across the northern Mediterranean Region from southwestern

Spain Spain, or the Kingdom of Spain, is a country in Southern Europe, Southern and Western Europe with territories in North Africa. Featuring the Punta de Tarifa, southernmost point of continental Europe, it is the largest country in Southern Eur ...

east to western

Turkey Turkey, officially the Republic of Türkiye, is a country mainly located in Anatolia in West Asia, with a relatively small part called East Thrace in Southeast Europe. It borders the Black Sea to the north; Georgia (country), Georgia, Armen ...

and

Cyprus Cyprus (), officially the Republic of Cyprus, is an island country in the eastern Mediterranean Sea. Situated in West Asia, its cultural identity and geopolitical orientation are overwhelmingly Southeast European. Cyprus is the List of isl ...

, growing on

coast A coast (coastline, shoreline, seashore) is the land next to the sea or the line that forms the boundary between the land and the ocean or a lake. Coasts are influenced by the topography of the surrounding landscape and by aquatic erosion, su ...

sand dune A dune is a landform composed of wind- or water-driven sand. It typically takes the form of a mound, ridge, or hill. An area with dunes is called a dune system or a dune complex. A large dune complex is called a dune field, while broad, flat ...

s from sea level up to in altitude.Adams, R. P. (2004). ''Junipers of the World''. Trafford. Farjon, A. (2005). ''Monograph of Cupressaceae and Sciadopitys''. Royal Botanic Gardens, Kew. A single, isolated tree is found further west, in a cliff in southern

Portugal Portugal, officially the Portuguese Republic, is a country on the Iberian Peninsula in Southwestern Europe. Featuring Cabo da Roca, the westernmost point in continental Europe, Portugal borders Spain to its north and east, with which it share ...

. It is a spreading

shrub A shrub or bush is a small to medium-sized perennial woody plant. Unlike herbaceous plants, shrubs have persistent woody stems above the ground. Shrubs can be either deciduous or evergreen. They are distinguished from trees by their multiple ...

tall, rarely a small

tree In botany, a tree is a perennial plant with an elongated stem, or trunk, usually supporting branches and leaves. In some usages, the definition of a tree may be narrower, e.g., including only woody plants with secondary growth, only ...

up to tall. The

leaves A leaf (: leaves) is a principal appendage of the stem of a vascular plant, usually borne laterally above ground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, stem, ...

are broad lanceolate, produced in whorls of three, green, long and 2–3 mm broad, with a double white

stoma In botany, a stoma (: stomata, from Greek language, Greek ''στόμα'', "mouth"), also called a stomate (: stomates), is a pore found in the Epidermis (botany), epidermis of leaves, stems, and other organs, that controls the rate of gas exc ...

tal band split by a green midrib on the inner surface. It is

dioecious Dioecy ( ; ; adj. dioecious, ) is a characteristic of certain species that have distinct unisexual individuals, each producing either male or female gametes, either directly (in animals) or indirectly (in seed plants). Dioecious reproduction is ...

, with separate male and female plants. The seed cones are berry-like, green ripening in 18 months to orange-red with a variable pink waxy coating; they are spherical, 12–18 mm diameter, and have six fused scales in two whorls, three of the scales with a single

seed In botany, a seed is a plant structure containing an embryo and stored nutrients in a protective coat called a ''testa''. More generally, the term "seed" means anything that can be Sowing, sown, which may include seed and husk or tuber. Seeds ...

. The seeds are dispersed when

bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...

s eat the cones, digesting the fleshy scales and passing the hard seeds in their droppings. The

pollen Pollen is a powdery substance produced by most types of flowers of seed plants for the purpose of sexual reproduction. It consists of pollen grains (highly reduced Gametophyte#Heterospory, microgametophytes), which produce male gametes (sperm ...

cones are yellow, 2–3 mm long, and fall soon after shedding their pollen in late winter.Arboretum de Villardebelle
photos of cones and shoots
/ref> Despite its distinct morphology with large cones and broad leaves more like those of '' Juniperus drupacea'', it has often been treated as a

subspecies In Taxonomy (biology), biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics (Morphology (biology), morpholog ...

of ''

Juniperus oxycedrus ''Juniperus oxycedrus'', vernacularly called Cade, cade juniper, prickly juniper, prickly cedar, or sharp cedar, is a species of juniper, native across the Mediterranean region, growing on a variety of rocky sites from sea level. The specific ep ...

'', though recent genetic studiesAdams, R. P. (2000). Systematics of ''Juniperus'' section ''Juniperus'' based on leaf essential oils and RAPD DNA fingerprinting. ''Biochem. Syst. Ecol''. 28: 515-528
available online (pdf file)
Adams, R. P., Morris, J. A., Pandey, R. N., & Schwarzbach, A. E. (2005). Cryptic speciation between ''Juniperus deltoides'' and ''J. oxycedrus'' (Cupressaceae) in the Mediterranean. ''Biochem. Syst. Ecol''. 33: 771-787
available online (pdf file)
/ref> have shown its

DNA Deoxyribonucleic acid (; DNA) is a polymer composed of two polynucleotide chains that coil around each other to form a double helix. The polymer carries genetic instructions for the development, functioning, growth and reproduction of al ...

is distinct from that of ''J. oxycedrus''.

References

{{Taxonbar, from=Q138307 macrocarpa Trees of Mediterranean climate Trees of Europe Trees of Turkey Flora of Cyprus Dioecious plants Flora of the Mediterranean basin