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The ivory-billed woodpecker (''Campephilus principalis'') is a possibly
extinct Extinction is the termination of a kind of organism or of a group of kinds (taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed and ...
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions ...
that is native to the
bottomland hardwood forest The bottomland hardwood forest is a type of deciduous and evergreen hardwood forest found in broad lowland floodplains along large rivers and lakes in the United States and elsewhere. They are occasionally flooded, which builds up the alluvial s ...
s and
temperate coniferous forest Temperate coniferous forest is a terrestrial biome defined by the World Wide Fund for Nature. Temperate coniferous forests are found predominantly in areas with warm summers and cool winters, and vary in their kinds of plant life. In some, needle ...
s of the
Southern United States The Southern United States (sometimes Dixie, also referred to as the Southern States, the American South, the Southland, or simply the South) is a geographic and cultural region of the United States of America. It is between the Atlantic Ocean ...
and
Cuba Cuba ( , ), officially the Republic of Cuba ( es, República de Cuba, links=no ), is an island country comprising the island of Cuba, as well as Isla de la Juventud and several minor archipelagos. Cuba is located where the northern Caribb ...
.
Habitat destruction Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
and hunting have reduced populations so thoroughly that the species is listed by the
International Union for Conservation of Nature The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natu ...
(IUCN) on its
Red List The International Union for Conservation of Nature (IUCN) Red List of Threatened Species, also known as the IUCN Red List or Red Data Book, founded in 1964, is the world's most comprehensive inventory of the global conservation status of biologi ...
as critically endangered, and by the
American Birding Association The American Birding Association (ABA) is a nonprofit organization, founded in 1969, dedicated to recreational birding in Canada and the United States. It has been called "the standard-bearer for serious birding in North America." Originally con ...
as "definitely or probably extinct". The last universally accepted sighting of an American ivory-billed woodpecker occurred in
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is bord ...
in 1944, and the last universally accepted sighting of a Cuban ivory-billed woodpecker occurred in 1987, after the bird's rediscovery there the prior year. Sporadic reports of sightings and other evidence of the persistence of the species have continued since then. The bird's preferred diet consists of large beetle larvae, particularly wood-boring
Cerambycidae The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...
beetles, supplemented by vegetable matter including such varied fruits as southern magnolia, pecans,
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera '' Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and b ...
s,
hickory nuts Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in Mex ...
, wild grapes, and persimmons. To hunt wood-boring beetle larvae, the bird uses its large bill to wedge and peel bark off dead trees to expose the larvae tunnels; no other species present in its range is able to remove tightly bound tree bark, and the ivory-bill faces no real competitor in hunting these larvae. It is, or was, the largest woodpecker in the United States, and one of the largest in the world, with a total length of and a typical wingspan of . In adults the bill is ivory in color, hence the bird's common name, while in juveniles it is chalky white. The bird has been found in habitat including dense
swampland A swamp is a forested wetland.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p. Swamps are considered to be transition zones because both land and water play a role in ...
, comparatively open
old-growth forest An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological feature ...
and, in Cuba, upland pine forests. Both parents work together to dig out a tree cavity roughly from the ground to create the nest, the typical depth of which is roughly . In the 21st century, reported sightings and analyses of audio and visual recordings were published in peer-reviewed scientific journals as evidence that the species persists in
Arkansas Arkansas ( ) is a landlocked state in the South Central United States. It is bordered by Missouri to the north, Tennessee and Mississippi to the east, Louisiana to the south, and Texas and Oklahoma to the west. Its name is from the O ...
,
Louisiana Louisiana , group=pronunciation (French: ''La Louisiane'') is a state in the Deep South and South Central regions of the United States. It is the 20th-smallest by area and the 25th most populous of the 50 U.S. states. Louisiana is bord ...
, and
Florida Florida is a state located in the Southeastern region of the United States. Florida is bordered to the west by the Gulf of Mexico, to the northwest by Alabama, to the north by Georgia, to the east by the Bahamas and Atlantic Ocean, and ...
. Various land purchases and
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
restoration efforts to protect any surviving individuals have been initiated in areas where sightings and other evidence have suggested a relatively high probability the species exists. In September 2021, the U.S. Fish and Wildlife Service proposed that the species be declared extinct. A virtual public hearing was held on January 26, 2022, and the public comment period ended February 10, 2022. A final decision by the Fish and Wildlife Service was due by September 2022. On July 11, 2022, the U.S. Fish and Wildlife Service announced it is extending its review period by 6 months to determine the status of the ivory-billed woodpecker.


Taxonomy

The ivory-billed woodpecker was first described as ''Picus maximus rostra albo'' (
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
for "the largest white-bill woodpecker") in
English English usually refers to: * English language * English people English may also refer to: Peoples, culture, and language * ''English'', an adjective for something of, from, or related to England ** English national ...
naturalist
Mark Catesby Mark Catesby (24 March 1683 – 23 December 1749) was an English naturalist who studied the flora and fauna of the New World. Between 1729 and 1747 Catesby published his ''Natural History of Carolina, Florida and the Bahama Islands'', the fi ...
's 1731 publication of ''Natural History of Carolina, Florida, and the Bahamas''. Noting his report,
Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, the ...
later described it in the landmark 1758
10th edition 1 (one, unit, unity) is a number representing a single or the only entity. 1 is also a numerical digit and represents a single unit of counting or measurement. For example, a line segment of ''unit length'' is a line segment of length 1. ...
of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...
'', where it was given the binomial name of ''Picus principalis''. The genus ''
Campephilus ''Campephilus'' is a genus of large American woodpeckers in the family Picidae. Taxonomy The genus ''Campephilus'' was introduced by English zoologist George Robert Gray in 1840, with the ivory-billed woodpecker (''Campephilus principalis'') ...
'' was introduced by the English zoologist
George Robert Gray George Robert Gray FRS (8 July 1808 – 6 May 1872) was an English zoologist and author, and head of the ornithological section of the British Museum, now the Natural History Museum, in London for forty-one years. He was the younger brother ...
in 1840 with the ivory-billed woodpecker as the
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
. Ornithologists currently recognize two subspecies of this bird: *American ivory-billed woodpecker (''C. p. principalis''), native to the southeastern United States *
Cuban ivory-billed woodpecker The Cuban ivory-billed woodpecker ( es, carpintero real, link=no) (''Campephilus principalis bairdii'') is a subspecies of the ivory-billed woodpecker native to Cuba. Originally classified as a separate species, recent research has indicated th ...
(''C. p. bairdii''), native to Cuba, including
Isla de la Juventud Isla de la Juventud (; en, Isle of Youth) is the second-largest Cuban island (after Cuba's mainland) and the seventh-largest island in the West Indies (after mainland Cuba itself, Hispaniola, Jamaica, Puerto Rico, Trinidad, and Andros Isl ...
The two look similar, with the Cuban bird somewhat smaller, some variations in
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
with the white dorsal strips extending to the bill, and the red crest feathers of the male being longer than its black crest feathers, while the two are of the same length in the American subspecies. Some controversy exists over whether the Cuban ivory-billed woodpecker is more appropriately recognized as a separate species. A 2006 study compared DNA samples taken from specimens of both ivory-billed woodpeckers, along with the imperial woodpecker (''Campephilus imperialis''), a larger but otherwise very similar bird. It concluded not only that the Cuban and American ivory-billed woodpeckers are genetically distinct, but also that they and the imperial form a North American clade within ''Campephilus'' that appeared in the mid-Pleistocene. The study does not attempt to define a lineage linking the three birds, although it does imply that the Cuban bird is more closely related to the imperial. The
American Ornithologists' Union The American Ornithological Society (AOS) is an ornithological organization based in the United States. The society was formed in October 2016 by the merger of the American Ornithologists' Union (AOU) and the Cooper Ornithological Society. Its ...
Committee on Classification and Nomenclature has said it is not yet ready to list the American and Cuban birds as separate species. Lovette, a member of the committee, said that more testing is needed to support that change, but concluded, "These results will likely initiate an interesting debate on how we should classify these birds." "Ivory-billed woodpecker" is the official name given to the species by the
International Ornithologists' Union The International Ornithologists' Union, formerly known as the International Ornithological Committee, is a group of about 200 international ornithologists, and is responsible for the International Ornithological Congress and other international ...
. Older common names included Log Cock, Log God, Good Lord, Indian Hen, Kent, Kate, (Wood Hen in
Cajun French Louisiana French ( frc, français de la Louisiane; lou, françé la lwizyàn) is an umbrella term for the dialects and varieties of the French language spoken traditionally by French Louisianians in colonial Lower Louisiana. As of today Louis ...
), and (Wood Cock in
Seminole The Seminole are a Native American people who developed in Florida in the 18th century. Today, they live in Oklahoma and Florida, and comprise three federally recognized tribes: the Seminole Nation of Oklahoma, the Seminole Tribe of Florida, ...
). Some modern authors refer to the species as the "Holy Grail bird" or "Grail Bird" because of its extreme rarity and elusiveness to birders.


Description

The ivory-billed woodpecker is one of the largest
woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions ...
s in the world at roughly long and in wingspan. It is the largest woodpecker in its range. The closely related imperial woodpecker (''C. imperialis'') of western Mexico is the largest woodpecker in the world. The ivory-billed woodpecker has a total length of , and based on scant information, weighs approximately . Its wingspan is typically . Standard measurements obtained include a wing chord length of , a tail length of , a bill length of , and a tarsus length of . The plumage of the ivory-billed woodpecker is predominated by a shiny black or purple tint. There are white lines extending from the cheeks down the neck, meeting on the back. The ends of the inner
primary feathers Flight feathers (''Pennae volatus'') are the long, stiff, asymmetrically shaped, but symmetrically paired pennaceous feathers on the wings or tail of a bird; those on the wings are called remiges (), singular remex (), while those on the tai ...
are white, as well as the whole of the outer secondary feathers. This creates extensive white on the trailing edge of both the upper- and underwing. The underwing also is white along its forward edge, resulting in a black line running along the middle of the underwing, expanding to more extensive black at the wingtip. Some birds have been recorded with more extensive amounts of white on the primary feathers. Ivory-bills have a prominent crest, although it is ragged in juveniles. The bird is somewhat
sexually dimorphic Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most ani ...
, as seen in the image to the right, the crest is black along its forward edge, changing abruptly to red on the side and rear in males, but solid black in females, as well as in juvenile males. When perched with the wings folded, birds of both genders present a large patch of white on the lower back, roughly triangular in shape. Like all woodpeckers, the ivory-billed woodpecker has a strong and straight bill and a long, mobile, hard-tipped, barbed tongue. In adults, the bill is ivory in color, while it is chalky white in juveniles. Among North American woodpeckers, the ivory-billed woodpecker is unique in having a bill whose tip is quite flattened laterally, shaped much like a beveled wood chisel. Its flight is strong and direct, and has been likened to that of a duck. These characteristics distinguish ivory-bills from the smaller and darker-billed pileated woodpecker. The pileated woodpecker normally is brownish-black, smoky, or slaty black in color. It also has a white neck stripe, but normally its back is black. Pileated woodpecker juveniles and adults have a red crest and a white chin. Pileated woodpeckers normally have no white on the trailing edges of their wings and when perched, normally show only a small patch of white on each side of the body, near the edge of the wing. However, aberrant individual pileated woodpeckers have been reported with white trailing edges on the wings, forming a white triangular patch on the lower back when perched. The drum of the ivory-billed woodpecker is a single or double rap. Four fairly distinct calls are reported in the literature and two were recorded in the 1930s. The most common, a ''kent'' or ''hant'', sounds like a toy
trumpet The trumpet is a brass instrument commonly used in classical and jazz ensembles. The trumpet group ranges from the piccolo trumpet—with the highest register in the brass family—to the bass trumpet, pitched one octave below the standard ...
often repeated in a series. When the bird is disturbed, the pitch of the ''kent'' note rises, it is repeated more frequently, and often doubled. A conversational call, also recorded, is given between individuals at the nest, and has been described as ''kent-kent-kent''.


Habitat and diet

No attempts to comprehensively estimate the range of the ivory-billed woodpecker were made until after its range already had been severely reduced by deforestation and hunting. The first range map produced for the species was made by Edwin M. Hasbrouck in 1891. The second range map produced was that made by James Tanner in 1942.Jackson (2004), page 47 Both authors reconstructed the original range of the species from historical records they considered reliable, in many cases from specimens with clear records of where they were obtained. The two authors produced broadly similar range estimates, finding that before deforestation and hunting began to shrink its range, the ivory-billed woodpecker had ranged from eastern
Texas Texas (, ; Spanish: ''Texas'', ''Tejas'') is a state in the South Central region of the United States. At 268,596 square miles (695,662 km2), and with more than 29.1 million residents in 2020, it is the second-largest U.S. state by ...
to
North Carolina North Carolina () is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. The state is the List of U.S. states and territories by area, 28th largest and List of states and territories of the United ...
, and from southern
Illinois Illinois ( ) is a state in the Midwestern United States. Its largest metropolitan areas include the Chicago metropolitan area, and the Metro East section, of Greater St. Louis. Other smaller metropolitan areas include, Peoria and Rock ...
to Florida and Cuba, typically from the coast inland to where the elevation is approximately . A few significant differences in their reconstructions exist, however. Based on the reports of Wells Woodbridge Cooke from Kansas City and Fayette, Hasbrouck's range map extended up the Missouri River and approximately to
Kansas City The Kansas City metropolitan area is a bi-state metropolitan area anchored by Kansas City, Missouri. Its 14 counties straddle the border between the U.S. states of Missouri (9 counties) and Kansas (5 counties). With and a population of more th ...
. which Tanner rejected as a possible accidental or unproven report.Tanner (1942), page 3 Similarly, Hasbrouck's range estimate extended up the
Ohio river The Ohio River is a long river in the United States. It is located at the boundary of the Midwestern and Southern United States, flowing southwesterly from western Pennsylvania to its mouth on the Mississippi River at the southern tip of ...
valley to
Franklin county, Indiana Franklin County is a county on the eastern border of the U.S. state of Indiana. In the 2020 United States Census, the county population was 22,785. The county seat is the town of Brookville. Franklin County is part of the Cincinnati, OH–KY� ...
based on a record from the E. T. Cox, which Tanner likewise rejected as unproven or accidental. Tanner's range estimate extended farther up the
Arkansas River The Arkansas River is a major tributary of the Mississippi River. It generally flows to the east and southeast as it traverses the U.S. states of Colorado, Kansas, Oklahoma, and Arkansas. The river's source basin lies in the western United ...
and
Canadian River The Canadian River is the longest tributary of the Arkansas River in the United States. It is about long, starting in Colorado and traveling through New Mexico, the Texas Panhandle, and Oklahoma. The drainage area is about .Fort Smith, Arkansas Fort Smith is the third-largest city in Arkansas and one of the two county seats of Sebastian County. As of the 2020 Census, the population was 89,142. It is the principal city of the Fort Smith, Arkansas–Oklahoma Metropolitan Statistical Are ...
, and Edwin James at the falls of the Canadian River, which were not mentioned by, and possibly, unknown to Hasbrouck. Tanner's range map is now generally accepted as the original range of the bird, but a number of records exist outside of both ranges, that were either overlooked or rejected by Tanner, or that surfaced after his analysis. Southwest of Tanner's range estimate, the species was reported along the
San Marcos River The San Marcos River rises from the San Marcos Springs, the location of Aquarena Springs, in San Marcos, Texas. The springs are home to several threatened or endangered species, including the Texas blind salamander, fountain darter, and Texas ...
and Guadalupe River, as well as near
New Braunfels New Braunfels ( ) is a city in Comal and Guadalupe counties in the U.S. state of Texas known for its German Texan heritage. It is the seat of Comal County. The city covers and had a population of 90,403 as of the 2020 Census. A suburb just north ...
, around 1900. Farther along the Ohio River Valley, William Fleming reported shooting an ivory-billed woodpecker at
Logan's Fort Stanford is a home rule-class city in Lincoln County, Kentucky, United States. It is one of the oldest settlements in Kentucky, having been founded in 1775. Its population was 3,487 at the 2010 census and an estimated 3,686 in 2018. It is the ...
, Kentucky in 1780. Ivory-billed woodpecker remains were found in middens in Scioto County, Ohio and were inferred to come from a bird locally hunted. Similar inferences were drawn from remains found near
Wheeling, West Virginia Wheeling is a city in the U.S. state of West Virginia. Located almost entirely in Ohio County, of which it is the county seat, it lies along the Ohio River in the foothills of the Appalachian Mountains and also contains a tiny portion extending ...
. There is also a report of a bird shot and eaten in Doddridge county, West Virginia around 1900.Jackson (2004), page 264 Based on reports that did not include specimens, Hasbrouck set the northern limit of the range along the Atlantic Coast to around Fort Macon, North Carolina, which was rejected as unproven by Tanner, who used the record of a bird shot north of
Wilmington, North Carolina Wilmington is a port city in and the county seat of New Hanover County in coastal southeastern North Carolina, United States. With a population of 115,451 at the 2020 census, it is the eighth most populous city in the state. Wilmington is t ...
by Alexander Wilson to set the northern limit of the range. Records exist of the ivory-billed woodpecker farther north along the Atlantic Coast;
Thomas Jefferson Thomas Jefferson (April 13, 1743 – July 4, 1826) was an American statesman, diplomat, lawyer, architect, philosopher, and Founding Fathers of the United States, Founding Father who served as the third president of the United States from 18 ...
included it as a bird of Virginia in ''
Notes on the State of Virginia ''Notes on the State of Virginia'' (1785) is a book written by the American statesman, philosopher, and planter Thomas Jefferson. He completed the first version in 1781 and updated and enlarged the book in 1782 and 1783. It originated in Jeffers ...
'', where it is listed as the "White bill woodpecker" with the designation of ''Picus principalis''. Audubon reported the bird could occasionally be found as far north as Maryland.
Pehr Kalm Pehr Kalm (6 March 1716 – 16 November 1779), also known as Peter Kalm, was a Swedish explorer, botanist, naturalist, and agricultural economist. He was one of the most important apostles of Carl Linnaeus. In 1747, he was commissioned by th ...
reported it was present seasonally in
Swedesboro, New Jersey Swedesboro is a borough in Gloucester County in the U.S. state of New Jersey. As of the 2010 U.S. census, the borough's population was 2,584,Winchester, Virginia Winchester is the most north western independent city in the Commonwealth of Virginia. It is the county seat of Frederick County, although the two are separate jurisdictions. The Bureau of Economic Analysis combines the city of Winchester wit ...
. Bones recovered from the Etowah Mounds in Georgia are generally believed to come from birds hunted locally. Within its range, the ivory-billed woodpecker is not smoothly distributed, but highly locally concentrated in areas where the habitat is suitable and where large quantities of appropriate food may be found. Knowledge of the ecology and behavior of ivory-billed woodpeckers is largely derived from James Tanner's study of several birds in a tract of forest along the Tensas River in the late 1930s. The extent to which those data can be extrapolated to ivory-bills as a whole, remains an open question. Ivory-billed woodpeckers have been found in habitat including dense swampland, comparatively open old-growth forest, and the upland pine forests of Cuba, but whether that is a complete list of suitable habitat is somewhat unclear. In the Tensas river region, Tanner estimate there was one pair of birds per . From historical data he estimated there was one pair of birds per in the California swamp in northern Florida and one pair per along the Wacissa river, he produced an understanding that these birds need large amounts of suitable territory to find enough food to feed themselves and their young, and thus they should be expected occur at low densities even in healthy populations. After the Civil War, the
timber industry Lumber is wood that has been processed into dimensional lumber, including beams and planks or boards, a stage in the process of wood production. Lumber is mainly used for construction framing, as well as finishing (floors, wall panels, w ...
deforested Deforestation or forest clearance is the removal of a forest or stand of trees from land that is then converted to non-forest use. Deforestation can involve conversion of forest land to farms, ranches, or urban use. The most concentrated d ...
millions of acres in the South, leaving only sparse, isolated tracts of appropriate habitat. Combined with the large range needs, this became the general understanding of the reason for the population decline of the species in the South. This picture has been disputed by Noel Snyder, who contended that hunting rather than habitat loss had been the primary cause of the population decline. He argued that Tanner's population estimates were made of an already depleted population, and the home range needs were significantly smaller. The preferred food of the ivory-billed woodpecker is beetle larvae, with roughly half of recorded stomach contents composed of large beetle larvae, particularly of species from the family
Cerambycidae The longhorn beetles (Cerambycidae), also known as long-horned or longicorns, are a large family of beetles, with over 35,000 species described. Most species are characterized by extremely long antennae, which are often as long as or longer than ...
,Jackson (2004), page 24 with
Scolytidae A bark beetle is the common name for the subfamily of beetles Scolytinae. Previously, this was considered a distinct family (Scolytidae), but is now understood to be a specialized clade of the "true weevil" family (Curculionidae). Although th ...
beetles also recorded. The bird also eats significant vegetable matter, with recorded stomach contents including the fruit of the southern magnolia, pecans,
acorn The acorn, or oaknut, is the nut of the oaks and their close relatives (genera '' Quercus'' and '' Lithocarpus'', in the family Fagaceae). It usually contains one seed (occasionally two seeds), enclosed in a tough, leathery shell, and b ...
s,
hickory nuts Hickory is a common name for trees composing the genus ''Carya'', which includes around 18 species. Five or six species are native to China, Indochina, and India (Assam), as many as twelve are native to the United States, four are found in Mex ...
, and poison ivy seeds. They also have been observed feeding on wild grapes, persimmons, and hackberries. To hunt woodboring grubs, the bird uses its enormous bill to hammer, wedge, and peel the bark off dead trees in order to access their tunnels. The species has no real competitors in hunting these grubs. No other species present in its range are able to remove tightly bound bark, as the ivory-billed woodpecker does.Tanner (1942), page 54 Ivory-billed woodpeckers are diurnal birds, spending their nights in individual roost holes that often are reused. The birds typically leave their roost holes around dawn, feeding and engaging in other activities during the early morning. They are generally inactive during the mid-day and resume feeding activities in the late afternoon before returning to the roosts around dusk.


Breeding biology and lifecycle

The ivory-billed woodpecker is thought to mate for life. Pairs are known to travel together. These paired birds breed every year between January and May. Both parents work together to excavate a cavity in a tree approximately from the ground for the nest in which their young will be raised. Limited data indicates a preference for living trees, or partially dead trees, with rotten ones avoided. Nest cavities are typically in or just below broken off stumps in living trees, where the wood is easier to excavate, and the overhanging stump can provide protection against rain and leave the opening in shadow, providing some protection against predators. There are no clear records of nest cavities being reused, and ivory-bills, like most woodpeckers, likely excavate a new nest each year.
Nest A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
openings are typically oval to rectangular in shape, and measure approximately tall by wide. The typical nest depth is roughly , with nests as shallow as and as deep as reported. Typically, eggs are laid in April or May, with a few records of eggs laid as early as mid-February. A second clutch has only been observed when the first one failed. Up to three glossy, china-white eggs are laid, measuring on average , although clutches of up to six eggs, and broods of up to four young, have been observed. No nest has been observed for the length of incubation so it remains unknown, although Tanner estimated it to be roughly 20 days. Parents incubate the eggs cooperatively, with the male observed to incubate overnight, and the two birds typically exchanging places every two hours during the day, with one foraging and one incubating. Once the young hatch, both parents forage to bring food to them. Young learn to fly about 7 to 8 weeks after hatching. The parents continue feeding them for another two months. The family eventually splits up in late fall or early winter. Ivory-billed woodpeckers are not migratory, and pairs are frequently observed to nest within a few hundred meters of previous nests, year after year. Although ivory-billed woodpeckers thus feed within a semiregular territory within a few kilometers of their nest or roost, they are not territorial; no records are known of ivory-bills protecting their territories from other ivory-bills when encountering one another. Indeed, in many instances the ivory-billed woodpecker has been observed acting as a social bird, with groups of four or five feeding together on a single tree, and as many as 11 observed feeding in the same location. Similarly, ivory-billed woodpeckers have been observed feeding on the same tree as the pileated woodpecker, the only other large woodpecker with which they share a range, without any hostile interactions. Although not migratory, sometimes the ivory-billed woodpecker is described as nomadic; birds relocate from time to time to areas where disasters such as fires or floods have created large amounts of dead wood, and subsequently large numbers of beetle larva upon which they prefer to feed. The maximum lifespan of an Ivory-billed woodpecker is not known, but other ''Campephilus'' woodpeckers are not known to live longer than 15 years, so this value is sometimes used as an estimate. No species (other than humans) are known to be predators of ivory-billed woodpeckers. However, they have been observed to exhibit predator response behaviors toward
Cooper's hawk Cooper's hawk (''Accipiter cooperii'') is a medium-sized hawk native to the North American continent and found from southern Canada to Mexico. This species is a member of the genus ''Accipiter'', sometimes referred to as true hawks, which are f ...
s and
red-shouldered hawk The red-shouldered hawk (''Buteo lineatus'') is a medium-sized buteo. Its breeding range spans eastern North America and along the coast of California and northern to northeastern-central Mexico. It is a permanent resident throughout most of its ...
s.


Status

Heavy logging activity exacerbated by
hunting Hunting is the human activity, human practice of seeking, pursuing, capturing, or killing wildlife or feral animals. The most common reasons for humans to hunt are to harvest food (i.e. meat) and useful animal products (fur/hide (skin), hide, ...
by collectors devastated the population of ivory-billed woodpeckers in the late 19th century. The species was generally considered extremely rare and some ornithologists believed it extinct by the 1920s. In 1924, Arthur Augustus Allen found a nesting pair in Florida, which local taxidermists shot for specimens. In 1932, a Louisiana state representative, Mason Spencer of Tallulah, killed an ivory-billed woodpecker along the Tensas River and took the specimen to his state wildlife office in
Baton Rouge Baton Rouge ( ; ) is a city in and the capital of the U.S. state of Louisiana. Located the eastern bank of the Mississippi River, it is the parish seat of East Baton Rouge Parish, Louisiana's most populous parish—the equivalent of counti ...
. As a result, Arthur Allen, fellow Cornell Ornithology professor Peter Paul Kellogg, Ph.D. student James Tanner, and avian artist
George Miksch Sutton George Miksch Sutton (May 16, 1898, Bethany, Nebraska – December 7, 1982) was an American ornithologist and bird artist. He published numerous technical papers in ornithology as well as more popular works illustrated with his own art. His early ...
organized an expedition to that part of Louisiana as part of a larger expedition to record images and sounds of endangered birds across the United States. The team located a population of woodpeckers in
Madison Parish Madison Parish (French: ''Paroisse de Madison'') is a parish located on the northeastern border of the U.S. state of Louisiana, in the delta lowlands along the Mississippi River. As of the 2010 census, the population was 12,093. Its paris ...
in northeastern Louisiana, in a section of the
old-growth forest An old-growth forestalso termed primary forest, virgin forest, late seral forest, primeval forest, or first-growth forestis a forest that has attained great age without significant disturbance, and thereby exhibits unique ecological feature ...
called the Singer tract, owned by the Singer Sewing Company, where logging rights were held by the Chicago Mill and Lumber Company. The team made the only universally accepted audio and motion picture recordings of the ivory-billed woodpecker. The
National Audubon Society The National Audubon Society (Audubon; ) is an American non-profit environmental organization dedicated to conservation of birds and their habitats. Located in the United States and incorporated in 1905, Audubon is one of the oldest of such orga ...
attempted to buy the logging rights to the tract so the habitat and birds could be preserved, but the company rejected their offer. Tanner spent 1937-1939 studying the ivory-billed woodpeckers on the Singer tract and travelling across the southern United States searching for other populations as part of his thesis work. At that time, he estimated there were 22–24 birds remaining, of which 6–8 were on the Singer tract. The last universally accepted sighting of an ivory billed woodpecker in the United States was made on the Singer tract by Audubon Society artist Don Eckelberry in April 1944, when logging of the tract was nearly complete. The ivory-billed woodpecker was listed as an endangered species on March 11, 1967, by the
United States Fish and Wildlife Service The United States Fish and Wildlife Service (USFWS or FWS) is an agency within the United States Department of the Interior dedicated to the management of fish, wildlife, and natural habitats. The mission of the agency is "working with othe ...
. It has been assessed as critically endangered by the
International Union for Conservation of Nature and Natural Resources The International Union for Conservation of Nature (IUCN; officially International Union for Conservation of Nature and Natural Resources) is an international organization working in the field of nature conservation and sustainable use of natur ...
, and is categorized as probably extinct or extinct by the
American Birding Association The American Birding Association (ABA) is a nonprofit organization, founded in 1969, dedicated to recreational birding in Canada and the United States. It has been called "the standard-bearer for serious birding in North America." Originally con ...
. A 2019 five-year review by the United States Fish and Wildlife Service recommended that the ivory-billed woodpecker be removed from the Endangered Species List due to extinction, and in September 2021 the U.S. Fish and Wildlife Service proposed that the species be declared extinct, pending a period of public comment that had been extended to February 10, 2022. A public hearing was held on January 26, 2022. On July 11, 2022 the U.S. Fish and Wildlife Service announced it is extending its review period by 6 months to determine the status of the ivory-billed woodpecker.


Evidence of persistence in the United States past 1944

Since 1944, regular reports have been made of ivory-billed woodpeckers being seen or heard across the southeastern United States, particularly in Louisiana, Florida, Texas, and South Carolina. In many instances, sightings were clearly misidentified pileated woodpeckers or red-headed woodpeckers. Similarly, in many cases, reports of hearing the ''kent'' call of the ivory-billed woodpecker were misidentifications of a similar call sometimes made by
blue jay The blue jay (''Cyanocitta cristata'') is a passerine bird in the family Corvidae, native to eastern North America. It lives in most of the eastern and central United States; some eastern populations may be migratory. Resident populations are ...
s. It also may be possible to mistake wing collisions in flying duck flocks for the characteristic double knock. However, a significant number of reports were accompanied by physical evidence or made by experienced ornithologists and could not be easily dismissed. In 1950, the Audubon Society established a wildlife sanctuary along the
Chipola River The Chipola River is a tributary of the Apalachicola River in western Florida. It is part of the ACF River Basin watershed. The U.S. Geological Survey. National Hydrography Dataset high-resolution flowline dataThe National Map, accessed April 1 ...
after a group led by
University of Florida The University of Florida (Florida or UF) is a public land-grant research university in Gainesville, Florida. It is a senior member of the State University System of Florida, traces its origins to 1853, and has operated continuously on its ...
graduate student Whitney Eastman reported a pair of ivory-billed woodpeckers with a roost hole. The sanctuary was terminated in 1952 when the woodpeckers could no longer be located. In 1967, ornithologist
John Dennis John Dennis may refer to: *John Dennis (dramatist) (1658–1734), English dramatist *John Dennis (1771–1806), Maryland congressman * John Dennis (1807–1859), his son, Maryland congressman * John Stoughton Dennis (1820–1885), Canadian surveyor ...
, sponsored by the U.S. Fish and Wildlife Service, reported sightings of ivory-billed woodpeckers along the
Neches River The Neches River () begins in Van Zandt County west of Rhine Lake and flows for through the piney woods of east Texas, defining the boundaries of 14 counties on its way to its mouth on Sabine Lake near the Rainbow Bridge. Two major reservoirs, ...
in Texas. Previously, Dennis had rediscovered the Cuban species in 1948. Dennis produced audio recording of possible ''kent'' calls that were found to be a good match to ivory-billed woodpecker calls, but possibly, also compatible with blue jays. At least 20 people reported sightings of one or more ivory-billed woodpeckers in the same area in the late 1960s, and several photographs, ostensibly showing an ivory-billed woodpecker in a roost, were produced by Neil Wright. Copies of two of his photographs were given to the
Academy of Natural Sciences of Drexel University The Academy of Natural Sciences of Drexel University, formerly the Academy of Natural Sciences of Philadelphia, is the oldest natural science research institution and museum in the Americas. It was founded in 1812, by many of the leading natura ...
. These sightings formed part of the basis for the creation of the
Big Thicket National Preserve The Big Thicket is the name given to a somewhat imprecise region of a heavily-forested area of Southeast Texas in the United States. This area represents a portion of the mixed pine-hardwood forests of southeast US. The National Park Service esta ...
. H. N. Agey and G. M. Heinzmann reported observing one or two ivory-billed woodpeckers in
Highlands County, Florida Highlands County is a county located in the Florida Heartland region of the U.S. state of Florida. As of the 2020 census, the population was 101,235. Its county seat is Sebring. Highlands County comprises the Sebring-Avon Park, FL Metropol ...
, on 11 occasions from 1967 to 1969. A tree in which the birds had been observed roosting was damaged during a storm and they were able to obtain a feather from the roost that was identified as an inner secondary feather of an ivory-billed woodpecker by A. Wetmore. The feather is stored at the
Florida Museum of Natural History The Florida Museum of Natural History (FLMNH) is Florida's official state-sponsored and chartered natural-history museum. Its main facilities are located at 3215 Hull Road on the campus of the University of Florida in Gainesville. The main pub ...
. The feather was described as "fresh, not worn", but as it could not be conclusively dated, it has not been universally accepted as proof that ivory-billed woodpeckers persisted to the date the feather was collected.
Louisiana State University Louisiana State University (officially Louisiana State University and Agricultural and Mechanical College, commonly referred to as LSU) is a public land-grant research university in Baton Rouge, Louisiana. The university was founded in 1860 n ...
museum director
George Lowery George Hines Lowery Jr. (October 2, 1913 – January 19, 1978) was an American ornithologist and professor of zoology at Louisiana State University, who pioneered a technique for studying nocturnal bird migration by large-scale observation through ...
presented two photographs at the 1971 annual meeting of the American Ornithologists' Union that show what appeared to be a male ivory-billed woodpecker. The photographs were taken by outdoorsman Fielding Lewis in the
Atchafalaya Basin The Atchafalaya Basin, or Atchafalaya Swamp (; Louisiana French: ''L'Atchafalaya'', ), is the largest wetland and swamp in the United States. Located in south central Louisiana, it is a combination of wetlands and river delta area where the At ...
of Louisiana, with an
Instamatic : ''For the film formats associated with the ''Instamatic'' and ''Pocket Instamatic'' camera ranges, see 126 film and 110 film respectively.'' The Instamatic is a series of inexpensive, easy-to-load 126 and 110 cameras made by Kodak beginning ...
camera. Although the photographs had the correct field markings for an ivory-billed woodpecker, their quality was not sufficient for other ornithologists to be confident that they did not depict a mounted specimen, and they were greeted with general skepticism. In 1999, a Louisiana State University forestry student reported an extended viewing of a pair of birds at close range in the
Pearl River The Pearl River, also known by its Chinese name Zhujiang or Zhu Jiang in Mandarin pinyin or Chu Kiang and formerly often known as the , is an extensive river system in southern China. The name "Pearl River" is also often used as a catch-a ...
region of southeast Louisiana, which some experts found very compelling. In 2002, an expedition, composed of researchers from Louisiana State University and Cornell University, was sent into the area. Six researchers spent 30 days searching the area, finding indications of large woodpeckers, but none that could be clearly ascribed to ivory-billed woodpeckers rather than pileated woodpeckers. Gene Sparling reported seeing an ivory-billed woodpecker in the
Cache River National Wildlife Refuge Cache, caching, or caché may refer to: Places United States * Cache, Idaho, an unincorporated community * Cache, Illinois, an unincorporated community * Cache, Oklahoma, a city in Comanche County * Cache, Utah, Cache County, Utah * Cache Coun ...
in 2004, prompting
Tim Gallagher Tim Gallagher is a writer and wildlife photographer and the author of six books: ''Parts Unknown, a Naturalist's Journey in Search of Birds and Wild Places''; ''The Grail Bird, Hot on the Trail of the Ivory-billed Woodpecker''; ''Falcon Fever, A Fa ...
and Bobby Harrison to investigate. They also observed a bird they identified as an ivory-billed woodpecker. An expedition led by John W. Fitzpatrick of the Cornell Laboratory of Ornithology followed and reported seven convincing sightings of an ivory-billed woodpecker. The team also heard and recorded possible double-knock and ''kent'' calls, and they produced a video with four seconds of footage of a large woodpecker that they identified as an ivory-billed woodpecker due to its size, field marks, and flight pattern. The sighting was accepted by the Bird Records Committee of the Arkansas Audubon Society. A team headed by David A. Sibley published a response arguing the bird in the video has a morphology that could be consistent with that of a pileated woodpecker, and a second team argued that flight characteristics may not be diagnostic. The original team published a rebuttal, but the identity of the bird in the video remains disputed. Following the publication of the video,
The Nature Conservancy The Nature Conservancy (TNC) is a global environmental organization headquartered in Arlington, Virginia. it works via affiliates or branches in 79 countries and territories, as well as across every state in the US. Founded in 1951, The Nat ...
bought of land to enlarge the protected areas that housed suitable habitat for the bird. However, a second search in 2005-2006 produced no unambiguous encounters with ivory-billed woodpeckers. The Lousiana State University-Cornell University collaboration team subsequently conducted searches in Arkansas, Florida, Illinois, Louisiana, Mississippi, North Carolina, South Carolina, Tennessee, and Texas, but found no clear indications of ivory-billed woodpeckers in any of those searches, at which point they concluded their efforts. Scientists from
Auburn University Auburn University (AU or Auburn) is a public land-grant research university in Auburn, Alabama. With more than 24,600 undergraduate students and a total enrollment of more than 30,000 with 1,330 faculty members, Auburn is the second largest ...
and the
University of Windsor , mottoeng = Goodness, Discipline and Knowledge , established = , academic_affiliations = CARL, COU, Universities Canada , former_names = Assumption College (1857-1956)Assumption University of Windsor (1956-1963) , type = Public universi ...
published a paper describing a search for ivory-billed woodpeckers along the Choctawhatchee River from 2005 to 2006, during which they recorded 14 sightings of ivory-billed woodpeckers, 41 occasions on which double-knocks or ''kent'' calls were heard, and 244 occasions on which double-knocks or ''kent'' calls were recorded. They analysed those recordings and conducted examinations of tree cavities and bark stripping by woodpeckers seen during the search and determined them to be consistent with the behavior of ivory-billed woodpeckers, but inconsistent with the behavior of pileated woodpeckers. In 2008, the sightings and sound detections largely dried up and the team ended their search in 2009. The scientists' sightings were not accepted by the Florida Ornithological Society Records Committee. Mike Collins reported ten sightings of ivory-billed woodpeckers between 2006 and 2008. He obtained video evidence at the Pearl River in Louisiana in 2006 and 2008 and at the Choctawhatchee River in Florida in 2007. His analyses of these sightings and videos were published in peer-reviewed journal articles.supplemental material
These reports, like all others since 1944, have encountered skepticism. Collins argues that the lack of clear photographs after 1944 is a function of species behavior and habitat, and that the expected time interval between clear photographs will be several orders of magnitude greater than it would be for a more typical species of comparable rarity. Project Principalis, a team of "researchers, community scientists, and nature enthusiasts" searching for the ivory-billed woodpecker, took photographs in November 2019, 9 January 2020, 1 October 2021, and 14 October 2021 that they argue depict ivory-billed woodpeckers.


Relationship with humans

Sometimes body parts of ivory-billed woodpeckers, particularly their bills, were used for trade, ceremonies, and decoration by various Native American groups from the western
Great Lakes The Great Lakes, also called the Great Lakes of North America, are a series of large interconnected freshwater lakes in the mid-east region of North America that connect to the Atlantic Ocean via the Saint Lawrence River. There are five lakes ...
and
Great Plains The Great Plains (french: Grandes Plaines), sometimes simply "the Plains", is a broad expanse of flatland in North America. It is located west of the Mississippi River and east of the Rocky Mountains, much of it covered in prairie, steppe, a ...
regions. For instance, bills marked with red pigment were found among
grave goods Grave goods, in archaeology and anthropology, are the items buried along with the body. They are usually personal possessions, supplies to smooth the deceased's journey into the afterlife or offerings to the gods. Grave goods may be classed as a ...
in burials at Ton won Tonga, a village of the Omaha people. The bills may have been part of "Wawaⁿ Pipes. Ivory-billed woodpecker bills and scalps were commonly incorporated into ceremonial pipes by the Iowa people, another Siouan-speaking people. The
Sauk people The Sauk or Sac are a group of Native Americans of the Eastern Woodlands culture group, who lived primarily in the region of what is now Green Bay, Wisconsin, when first encountered by the French in 1667. Their autonym is oθaakiiwaki, and th ...
and
Meskwaki The Meskwaki (sometimes spelled Mesquaki), also known by the European exonyms Fox Indians or the Fox, are a Native American people. They have been closely linked to the Sauk people of the same language family. In the Meskwaki language, th ...
used ivory-billed body parts in amulets, headbands, and sacred bundles. In many cases it is likely that the bills were acquired through trade. For instance, Ton won Tonga was located roughly from the farthest reported range of the ivory-billed woodpecker, and the bills were only found in the graves of wealthy adult men. Another bill was found in a grave in
Johnstown, Colorado Johnstown is a home rule municipality in Weld and Larimer counties in the U.S. state of Colorado. The population was 9,887 at the 2010 United States Census. History The Town of Johnstown began with the vision of Harvey J. Parish before its pl ...
. The bills were quite valuable, with Catesby reporting a north–south trade where bills were exchanged outside the bird's range for two or three deerskins. European settlers in the United States also used ivory-bill remains for adornment, often securing dried heads to their shot pouches, or employing them as
watch fob A pocket watch (or pocketwatch) is a watch that is made to be carried in a pocket, as opposed to a wristwatch, which is strapped to the wrist. They were the most common type of watch from their development in the 16th century until wristwatc ...
s. The presence of remains in
kitchen midden A midden (also kitchen midden or shell heap) is an old dump for domestic waste which may consist of animal bone, human excrement, botanical material, mollusc shells, potsherds, lithics (especially debitage), and other artifacts and ecofa ...
s has been used to infer that some Native American groups would hunt and eat the ivory-billed woodpecker. Such remains have been found in Illinois, Ohio, West Virginia, and Georgia. The hunting of ivory-billed woodpeckers for food by the residents of the Southeastern United States continued into the early 20th century, with reports of hunting ivory-billed woodpeckers for food continuing until at least the 1950s. In some instances, the flesh of ivory-billed woodpeckers was used as bait by trappers and fishermen. In the 19th and into the early 20th century, hunting for
bird collections Bird collections are curated repositories of scientific specimens consisting of birds and their parts. They are a research resource for ornithology, the science of birds, and for other scientific disciplines in which information about birds is us ...
was extensive, with 413 specimens now housed in museum and university collections. The largest collection is that of more than 60 skins at the
Harvard Museum of Comparative Zoology Harvard University is a Private university, private Ivy League research university in Cambridge, Massachusetts. Founded in 1636 as Harvard College and named for its first benefactor, the History of the Puritans in North America, Puritan cler ...
. The ivory-billed woodpecker has been a particular focus among
birdwatchers Birdwatching, or birding, is the observing of birds, either as a recreational activity or as a form of citizen science. A birdwatcher may observe by using their naked eye, by using a visual enhancement device like binoculars or a telescope, by ...
. It has been called Audubon's favorite bird.
Roger Tory Peterson Roger Tory Peterson (August 28, 1908 – July 28, 1996) was an American naturalist, ornithologist, illustrator and educator, and one of the founding inspirations for the 20th-century environmental movement. Background Peterson was born in James ...
called his unsuccessful search for the birds along the
Congaree River The Congaree River is a short but wide river in South Carolina in the United States; It flows for approximately 53 miles (85 km). The river serves an important role as the final outlet channel for the entire Lower Saluda and Lower Broad wa ...
in the 1930s his "most exciting bird experience". After the publication of the Fitzpatrick results, tourist attention was drawn to eastern Arkansas, with tourist spending up 30% in and around the city of Brinkley, Arkansas. Brinkley hosted "The Call of the Ivory-billed Woodpecker Celebration" in February 2006. The celebration included exhibits, birding tours, educational presentations, and a vendor market. By the 21st century, the ivory-billed woodpecker had achieved a near-mythic status among birdwatchers, most of whom would regard it as a prestigious entry on their
life list A life list, or life-list, is a List (information), list of all biological species seen by a person. The phrase is particularly common among bird watchers, some of whom compete with each other to have the most complete list. References

Bird ...
s. The rare and elusive status of the species has inspired rewards being offered for information that allows the location of live birds. During their searches for conclusive proof of continued existence of the species, Cornell University offered a reward of $50,000. The Louisiana Wilds project offered $12,000 for the location of an active roost or nest in 2020. The ivory-billed woodpecker has been the subject of various artistic works. Joseph Bartholomew Kidd produced a painting based on Audubon's plates that was intended for a travelling exhibition throughout the United Kingdom and United States. The exhibition never took place and the painting is displayed in the
Metropolitan Museum of Art The Metropolitan Museum of Art of New York City, colloquially "the Met", is the largest art museum in the Americas. Its permanent collection contains over two million works, divided among 17 curatorial departments. The main building at 1000 ...
. Based on interviews with residents of Brinkley, Arkansas,
Sufjan Stevens Sufjan Stevens ( ; born July 1, 1975) is an American singer, songwriter, and multi-instrumentalist. He has released nine solo studio albums and multiple collaborative albums with other artists. Stevens has received Grammy and Academy Award nom ...
wrote a song entitled "The Lord God Bird" on the ivory-billed woodpecker that was broadcast on
National Public Radio National Public Radio (NPR, stylized in all lowercase) is an American privately and state funded nonprofit media organization headquartered in Washington, D.C., with its NPR West headquarters in Culver City, California. It differs from other n ...
following the public reports of sightings there. The Alex Karpovsky film entitled ''Red Flag'' features Karpovsky as a filmmaker touring his documentary film about the ivory-billed woodpecker that is entitled ''
Woodpecker Woodpeckers are part of the bird family Picidae, which also includes the piculets, wrynecks, and sapsuckers. Members of this family are found worldwide, except for Australia, New Guinea, New Zealand, Madagascar, and the extreme polar regions ...
''. Arkansas has issued license plates featuring a graphic of an ivory-billed woodpecker.


Notes


References


Further reading

*Farrand, John Jr. and Bull, John, ''The Audubon Society Field Guide to North American Birds: Eastern Region'', National Audubon Society (1977)


External links


Ivory-billed Woodpecker (Northern)

The U.S. Fish and Wildlife Service draft recovery plan

U.S. Fish and Wildlife Service Delisting Proposal
{{Taxonbar, from=Q756063 ivory-billed woodpecker Native birds of the Southeastern United States Birds of the Caribbean Birds of Cuba ivory-billed woodpecker Articles containing video clips ESA endangered species ivory-billed woodpecker