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In
botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
, an infraspecific name is the scientific name for any
taxon In biology, a taxon ( back-formation from '' taxonomy''; plural taxa) is a group of one or more populations of an organism or organisms seen by taxonomists to form a unit. Although neither is required, a taxon is usually known by a particular n ...
below the
rank Rank is the relative position, value, worth, complexity, power, importance, authority, level, etc. of a person or object within a ranking, such as: Level or position in a hierarchical organization * Academic rank * Diplomatic rank * Hierarchy * ...
of
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriate s ...
, i.e. an infraspecific taxon or infraspecies. (A "taxon", plural "taxa", is a group of organisms to be given a particular name.) The scientific names of botanical taxa are regulated by the '' International Code of Nomenclature for algae, fungi, and plants'' (ICN). This specifies a 'three part name' for infraspecific taxa, plus a 'connecting term' to indicate the rank of the name., Art. 24 An example of such a name is ''Astrophytum myriostigma'' subvar. ''glabrum'', the name of a subvariety of the species '' Astrophytum myriostigma'' (bishop's hat cactus). Names below the rank of species of cultivated kinds of plants and of animals are regulated by different codes of nomenclature and are formed somewhat differently.


Construction of infraspecific names

Article 24 of the ICN describes how infraspecific names are constructed. The order of the three parts of an infraspecific name is: :genus name, specific epithet, connecting term indicating the rank (not part of the name, but required), infraspecific epithet. It is customary to italicize all three parts of such a name, but not the connecting term. For example: * ''Acanthocalycium klimpelianum'' var. ''macranthum'' :genus name = ''Acanthocalycium'', specific epithet = ''klimpelianum'', connecting term = var. (short for "varietas" or
variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
), infraspecific epithet = ''macranthum'' * ''Astrophytum myriostigma'' subvar. ''glabrum'' :genus name = ''Astrophytum'', specific epithet = ''myriostigma'', connecting term = subvar. (short for "subvarietas" or subvariety), infraspecific epithet = ''glabrum'' The recommended abbreviations for ranks below species are: * subspecies - recommended abbreviation: subsp. (but "ssp." is also in use although not recognised by Art 26) * varietas (
variety Variety may refer to: Arts and entertainment Entertainment formats * Variety (radio) * Variety show, in theater and television Films * ''Variety'' (1925 film), a German silent film directed by Ewald Andre Dupont * ''Variety'' (1935 film), ...
) - recommended abbreviation: var. * subvarietas ( subvariety) - recommended abbreviation: subvar. * forma ( form) - recommended abbreviation: f. * subforma (subform) - recommended abbreviation: subf. Although the "connecting terms" mentioned above to indicate rank, such as "var.", are the recommended ones, the ICN allows for other connecting terms in validly published infraspecific taxa. It specifically mentions that Greek letters α, β, γ, etc. can be used in this way in the original document, Note 1 to Art. 32.1 and further ranks may be added without limit., Art. 4.3 and Art. 2 Names which use these connecting terms are now deprecated (though still legal), but they have an importance because they can be basionyms of current species. The commonest cases use "β" and "b"; examples mentioned in the ICN are ''Cynoglossum cheirifolium'' β Anchusa (''lanata''), Art. 24.4 Ex. 9 and ''Polyporus fomentarius'' β ''applanatus'', entry ''Boletus applanatus'' of Appendix III whilst other examples (coming from the fungus database
Index Fungorum ''Index Fungorum'' is an international project to index all formal names (scientific names) in the fungus kingdom. the project is based at the Royal Botanic Gardens, Kew, one of three partners along with Landcare Research and the Institute of M ...
) are ''Agaricus plexipes'' b ''fuliginaria'' and ''Peziza capula'' ß ''cernua''. The ICN allows the possibility that a validly published name could have no defined rank and uses " nranked as the connecting term in such cases., for instance ''Agaricus compactus'' nranked''sarcocephalus'' in Art. 50E.Ex 7, or ''Geranium andicola'' nranked''longipedicellatum'' in Art. 37.3.Ex. 5


Abbreviation of infraspecific names

Like specific epithets, infraspecific epithets cannot be used in isolation as names., Art. 24.1 Thus the name of a particular species of '' Acanthocalycium'' is '' Acanthocalycium klimpelianum'', which can be abbreviated to ''A. klimpelianum'' where the context makes the genus clear. The species cannot be referred to as just ''klimpelianum''. In the same way, the name of a particular variety of ''Acanthocalycium klimpelianum'' is ''Acanthocalycium klimpelianum'' var. ''macranthum'', which can be abbreviated to ''A. k.'' var. ''macranthum'' where the context makes the species clear. The variety cannot be referred to as just ''macranthum''. Sometimes more than three parts will be given; strictly speaking, this is not a name, but a classification. The ICN gives the example of ''Saxifraga aizoon'' var. ''aizoon'' subvar. ''brevifolia'' f. ''multicaulis'' subf. ''surculosa''; the name of the subform would be ''Saxifraga aizoon'' subf. ''surculosa''.


Legitimate infraspecific names

For a proposed infraspecific name to be legitimate it must be in accordance with all the rules of the ICN. Only some of the main points are described here. A key concept in botanical names is that of a type. In many cases the type will be a particular preserved specimen stored in a herbarium, although there are other kinds of type. Like other names, an infraspecific name is attached to a type. Whether a plant should be given a particular infraspecific name can then be decided by comparing it to the type. There is no requirement for a species to be divided into infraspecific taxa, of whatever rank; in other words, a species does not have to have subspecies, varieties, forms, etc. However, if infraspecific ranks are created, then the name of the type of the species must repeat the specific epithet as its infraspecific epithet. The type acquires this name automatically as soon as any infraspecific rank is created., Art. 26 As an example, consider ''
Poa secunda ''Poa secunda'' (variously known by the common names of Sandberg bluegrass, alkali bluegrass, big bluegrass, Canby's bluegrass, Nevada bluegrass, one-sided bluegrass, Pacific bluegrass, pine blugrass, slender bluegrass, wild bluegrass, and curly ...
'' J.Presl, whose type specimen is in the Wisconsin State Herbarium. * As soon as a subspecies of ''Poa secunda'' was created, then the type specimen of ''P. secunda'' immediately became the type specimen of ''Poa secunda'' subsp. ''secunda''. The name "''Poa secunda'' subsp. ''secunda''" was automatically created (it is an "autonym"). Soreng created the subspecies ''Poa secunda'' subsp. ''juncifolia'' (whose type specimen is also in the Wisconsin State Herbarium), thereby making the type specimen of ''P. secunda'' also the type specimen of ''Poa secunda'' subsp. ''secunda''. * If in addition to the subspecies any variety of ''Poa secunda'' were to be created, then the type specimen of ''P. secunda'' would automatically become the type specimen of ''Poa secunda'' var. ''secunda''. The type specimen would then have the classification ''Poa secunda'' subsp. ''secunda'' var. ''secunda''. The same epithet can be used again within a species, at whatever level, only if the names with the re-used epithet are attached to the same type. Thus there can be a form called ''Poa secunda'' f. ''juncifolia'' as well as the subspecies ''Poa secunda'' subsp. ''juncifolia'' if, and only if, the type specimen of ''Poa secunda'' f. ''juncifolia'' is the same as the type specimen of ''Poa secunda'' subsp. ''juncifolia'' (in other words, if there is a single type specimen whose classification is ''Poa secunda'' subsp. ''juncifolia'' f. ''juncifolia''). If two infraspecific taxa which have different types are accidentally given the same epithet, then a
homonym In linguistics, homonyms are words which are homographs (words that share the same spelling, regardless of pronunciation), or homophones ( equivocal words, that share the same pronunciation, regardless of spelling), or both. Using this definiti ...
has been created. The earliest published name is the legitimate one and the other must be changed., Art. 53.3


Specifying authors

When indicating authors for infraspecific names, it is possible to show either just the author(s) of the final, infraspecific epithet, or the authors of both the specific and the infraspecific epithets. Examples: * ''Adenia aculeata'' subsp. ''inermis'' de Wilde :This identifies de Wilde as the author who published this name for the subspecies (i.e. who created the epithet ''inermis''). Note that here it was decided not to indicate authority for the species. * ''Pinus nigra'' J.F.Arnold subsp. ''salzmannii'' (Dunal) Franco :Here, J.F.Arnold is the author who gave the species,
European black pine ''Pinus nigra'', the Austrian pine or black pine, is a moderately variable species of pine, occurring across Southern Europe from the Iberian Peninsula to the eastern Mediterranean, on the Anatolian peninsula of Turkey, Corsica and Cyprus, as wel ...
, its botanical name ''Pinus nigra''; Dunal is the author who was the first to publish the epithet ''salzmanii'' for this taxon (as the species ''Pinus salzmanii''); Franco is the author who reduced the taxon to a subspecies of ''Pinus nigra''.


Difference from zoological nomenclature

In
zoological nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the In ...
, names of taxa below species rank are formed somewhat differently, using a
trinomen In biology, trinomial nomenclature refers to names for taxa below the rank of species. These names have three parts. The usage is different in zoology and botany. In zoology In zoological nomenclature, a trinomen (), trinominal name, or ternary ...
or 'trinomial name'. No connecting term is required as there is only one rank below species, the subspecies.


Cultivated plants

The ''ICN'' does not regulate the names of cultivated plants, of
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
s, i.e. plants specifically created for use in agriculture or horticulture. Such names are regulated by the ''
International Code of Nomenclature for Cultivated Plants The ''International Code of Nomenclature for Cultivated Plants'' (ICNCP), is a guide to the rules and regulations for naming cultigens, plants whose origin or selection is primarily due to intentional human activity. It is also known as Cultivat ...
'' (ICNCP). Although logically below the rank of species (and hence "infraspecific"), a
cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
name may be attached to any scientific name at the genus level or below. The minimum requirement is to specify a genus name. For example, ''Achillea'' 'Cerise Queen' is a cultivar; ''Pinus nigra'' 'Arnold Sentinel' is a cultivar of the species ''P. nigra'' (which is propagated vegetatively, by
cloning Cloning is the process of producing individual organisms with identical or virtually identical DNA, either by natural or artificial means. In nature, some organisms produce clones through asexual reproduction. In the field of biotechnology, c ...
).


See also

* ''
International Code of Zoological Nomenclature The International Code of Zoological Nomenclature (ICZN) is a widely accepted convention in zoology that rules the formal scientific naming of organisms treated as animals. It is also informally known as the ICZN Code, for its publisher, the ...
'' * ''
International Code of Nomenclature for Cultivated Plants The ''International Code of Nomenclature for Cultivated Plants'' (ICNCP), is a guide to the rules and regulations for naming cultigens, plants whose origin or selection is primarily due to intentional human activity. It is also known as Cultivat ...
'' *
Cultivar A cultivar is a type of cultivated plant that people have selected for desired traits and when propagated retain those traits. Methods used to propagate cultivars include: division, root and stem cuttings, offsets, grafting, tissue culture ...
* ''
Forma specialis ''Forma specialis'' (plural: ''formae speciales''), abbreviated f. sp. (plural ff. spp.) without italics, is an informal taxonomic grouping allowed by the International Code of Nomenclature for algae, fungi, and plants, that is applied to a par ...
'', an informal naming system for parasites that is included in the botanical code of nomenclature *
Pathovar A pathovar is a bacterial strain or set of strains with the same or similar characteristics, that is differentiated at infrasubspecific level from other strains of the same species or subspecies on the basis of distinctive pathogenicity to one o ...
, a system for naming parasitic bacteria


References


Bibliography

* {{DEFAULTSORT:Infraspecific Name Botanical nomenclature Plant taxonomy Biology terminology Biological classification