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Ichthyopterygia ("fish flippers") was a designation introduced by
Sir Richard Owen Sir Richard Owen (20 July 1804 – 18 December 1892) was an English biologist, comparative anatomist and paleontologist. Owen is generally considered to have been an outstanding naturalist with a remarkable gift for interpreting fossils. Ow ...
in 1840 to designate the Jurassic
ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, alt ...
s that were known at the time, but the term is now used more often for both true Ichthyosauria and their more primitive early and middle
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest per ...
ancestors. Basal ichthyopterygians (prior to and ancestral to true Ichthyosauria) were mostly small (a meter or less in length) with elongated bodies and long, spool-shaped
vertebra The spinal column, a defining synapomorphy shared by nearly all vertebrates, Hagfish are believed to have secondarily lost their spinal column is a moderately flexible series of vertebrae (singular vertebra), each constituting a characteristi ...
e, indicating that they swam in a sinuous, eel-like manner. This allowed for quick movements and maneuverability that were advantages in shallow-water hunting. Even at this early stage, they were already very specialised animals with proper flippers, and would have been incapable of movement on land. These animals seem to have been widely distributed around the coast of the northern half of
Pangea Pangaea or Pangea () was a supercontinent that existed during the late Paleozoic and early Mesozoic eras. It assembled from the earlier continental units of Gondwana, Euramerica and Siberia during the Carboniferous approximately 335 million y ...
, as they are known the Late
Olenekian In the geologic timescale, the Olenekian is an age in the Early Triassic epoch; in chronostratigraphy, it is a stage in the Lower Triassic series. It spans the time between Ma and Ma (million years ago). The Olenekian is sometimes divided i ...
and Early
Anisian In the geologic timescale, the Anisian is the lower stage or earliest age of the Middle Triassic series or epoch and lasted from million years ago until million years ago. The Anisian Age succeeds the Olenekian Age (part of the Lower Triassic ...
(early part of the
Triassic The Triassic ( ) is a geologic period and system which spans 50.6 million years from the end of the Permian Period 251.902 million years ago ( Mya), to the beginning of the Jurassic Period 201.36 Mya. The Triassic is the first and shortest per ...
period) of
Japan Japan ( ja, 日本, or , and formally , ''Nihonkoku'') is an island country in East Asia. It is situated in the northwest Pacific Ocean, and is bordered on the west by the Sea of Japan, while extending from the Sea of Okhotsk in the n ...
,
China China, officially the People's Republic of China (PRC), is a country in East Asia. It is the world's List of countries and dependencies by population, most populous country, with a Population of China, population exceeding 1.4 billion, slig ...
,
Canada Canada is a country in North America. Its ten provinces and three territories extend from the Atlantic Ocean to the Pacific Ocean and northward into the Arctic Ocean, covering over , making it the world's second-largest country by to ...
, and
Spitsbergen Spitsbergen (; formerly known as West Spitsbergen; Norwegian: ''Vest Spitsbergen'' or ''Vestspitsbergen'' , also sometimes spelled Spitzbergen) is the largest and the only permanently populated island of the Svalbard archipelago in northern Nor ...
(
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of ...
). By the later part of the Middle Triassic, they were extinct, having been replaced by their descendants, the true ichthyosaurs.


Taxonomy

* Superorder Ichthyopterygia **? Genus '' Isfjordosaurus'' **? Family Omphalosauridae ** Family
Parvinatatoridae ''Parvinatator'', from Latin, “parvus” little and “natator” swimmer, is an extinct genus of small ichthyopterygian marine reptile that lived during the Early to Middle Triassic. Its fossils have been found in British Columbia, Canada ...
** Family
Thaisauridae ''Thaisaurus'' is an extinct genus of ichthyopterygian marine reptile that lived during the Spathian (late Olenekian, Early Triassic). Fossils have been found in Thailand.New Material of ''Qianichtyosaurus'' Li, 1999 (Reptilia, Ichthyosauria) fr ...
** Family
Utatsusauridae ''Utatsusaurus hataii'' is the earliest-known ichthyopterygian which lived in the Early Triassic period (c. 245–250 million years ago). It was nearly long with a slender body. The first specimen was found in Utatsu-cho (now part of Minamisa ...
** Eoichthyosauria *** Order
Grippidia Ichthyopterygia ("fish flippers") was a designation introduced by Sir Richard Owen in 1840 to designate the Jurassic ichthyosaurs that were known at the time, but the term is now used more often for both true Ichthyosauria and their more primit ...
*** Order
Ichthyosaur Ichthyosaurs (Ancient Greek for "fish lizard" – and ) are large extinct marine reptiles. Ichthyosaurs belong to the order known as Ichthyosauria or Ichthyopterygia ('fish flippers' – a designation introduced by Sir Richard Owen in 1842, alt ...
ia


Phylogeny

Below is a
cladogram A cladogram (from Greek ''clados'' "branch" and ''gramma'' "character") is a diagram used in cladistics to show relations among organisms. A cladogram is not, however, an evolutionary tree because it does not show how ancestors are related to ...
modified from Cuthbertson ''et al.'', 2013.


References


General references

* Ellis, Richard, (2003) ''Sea Dragons - Predators of the Prehistoric Oceans''. University Press of Kansas * McGowan, C & Motani, R. (2003) Ichthyopterygia, '' Handbook of Paleoherpetology'', Part 8, Verlag Dr. Friedrich Pfeil


External links


Ryosuke Motani's detailed Ichthyosaur homepage, with vivid graphics
{{Taxonbar, from=Q2583869 Ichthyosauromorphs Chordate superorders Early Triassic first appearances Turonian extinctions simple:Ichthyopterygia