Honeydew is a

sugar Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose Glucose is a sugar with the Chemical formula#Molecular formula, molecul ...

-rich sticky

liquid Liquid is a state of matter with a definite volume but no fixed shape. Liquids adapt to the shape of their container and are nearly incompressible, maintaining their volume even under pressure. The density of a liquid is usually close to th ...

, secreted by

aphid Aphids are small sap-sucking insects in the Taxonomic rank, family Aphididae. Common names include greenfly and blackfly, although individuals within a species can vary widely in color. The group includes the fluffy white Eriosomatinae, woolly ...

s, some

scale insect Scale insects are small insects of the Order (biology), order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient g ...

s, and many other

true bugs Hemiptera (; ) is an order (biology), order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, Cimex, bed bugs, and shield bugs. They ...

and some other insects as they feed on

plant sap Sap is a fluid transported in the xylem cells (vessel elements or tracheids) or phloem sieve tube elements of a plant. These cells transport water and nutrients throughout the plant. Sap is distinct from latex, resin, or cell sap; it is a sep ...

. When their mouthpart penetrates the

phloem Phloem (, ) is the living tissue in vascular plants that transports the soluble organic compounds made during photosynthesis and known as ''photosynthates'', in particular the sugar sucrose, to the rest of the plant. This transport process is ...

, the sugary, high-pressure liquid is forced out of the

anus In mammals, invertebrates and most fish, the anus (: anuses or ani; from Latin, 'ring' or 'circle') is the external body orifice at the ''exit'' end of the digestive tract (bowel), i.e. the opposite end from the mouth. Its function is to facil ...

of the insects, allowing them to rapidly process the large volume of sap required to extract essential nutrients present at low concentrations. Honeydew is particularly common as a secretion in

hemiptera Hemiptera (; ) is an order of insects, commonly called true bugs, comprising more than 80,000 species within groups such as the cicadas, aphids, planthoppers, leafhoppers, assassin bugs, bed bugs, and shield bugs. They range in size from ...

n insects and is often the basis for

trophobiosis Trophobiosis is a symbiosis, symbiotic association between organisms where food is obtained or provided. The provider of food in the association is referred to as a trophobiont. The name is derived from the Ancient Greek τροφή (''trophē''), ...

. Some

caterpillar Caterpillars ( ) are the larval stage of members of the order Lepidoptera (the insect order comprising butterflies and moths). As with most common names, the application of the word is arbitrary, since the larvae of sawflies (suborder ...

s of

Lycaenidae Lycaenidae is the second-largest family (biology), family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 6,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of ...

butterflies and some moths also produce honeydew. In addition to various sugars, honeydew contains small amounts of

amino acid Amino acids are organic compounds that contain both amino and carboxylic acid functional groups. Although over 500 amino acids exist in nature, by far the most important are the 22 α-amino acids incorporated into proteins. Only these 22 a ...

s, other organic compounds, and inorganic

salt In common usage, salt is a mineral composed primarily of sodium chloride (NaCl). When used in food, especially in granulated form, it is more formally called table salt. In the form of a natural crystalline mineral, salt is also known as r ...

s with its precise makeup affected by factors such as insect species, host plant species, and whether a symbiotic organism is present. Honeydew-producing insects, like cicadas, pierce phloem ducts to access the sugar rich sap; the excess fluid released by cicadas as honeydew is called "cicada rain". The sap continues to bleed after the insects have moved on, leaving a white sugar crust called manna.

Ant Ants are Eusociality, eusocial insects of the Family (biology), family Formicidae and, along with the related wasps and bees, belong to the Taxonomy (biology), order Hymenoptera. Ants evolved from Vespoidea, vespoid wasp ancestors in the Cre ...

s may collect, or "milk", honeydew directly from aphids and other honeydew producers, which benefit from their presence due to their driving away predators such as lady beetles or parasitic wasps—see '' Crematogaster peringueyi''. Animals and plants in a mutually symbiotic arrangement with ants are called Myrmecophiles. In Madagascar, some gecko species in the genera '' Phelsuma'' and ''

Lygodactylus ''Lygodactylus'' is a genus of Diurnality, diurnal geckos with 82 species. They are commonly referred to as dwarf geckos. They are mainly found in Africa and Madagascar although two species are found in South America. ''Lygodactylus picturatus'' ...

'' are known to approach flatid plant-hoppers on tree-trunks from below and induce them to excrete honeydew by head nodding behaviour. The plant-hopper then raises its abdomen and excretes a drop of honeydew almost right onto the snout of the gecko. Honeydew can cause

sooty mold Sooty mold (also spelled sooty mould) is a collective term for different Ascomycete fungi, which includes many genera, commonly ''Cladosporium'' and ''Alternaria''. It grows on plants and their fruit, but also environmental objects, like fences, ...

on many

ornamental plant Ornamental plants or ''garden plants'' are plants that are primarily grown for their beauty but also for qualities such as scent or how they shape physical space. Many flowering plants and garden varieties tend to be specially bred cultivars th ...

s. It also contaminates vehicles parked beneath trees, and can then be difficult to remove from glass and bodywork. Honeydew is also secreted by certain fungi, particularly ergot. Honeydew is collected by certain species of birds, mosquitoes,

wasp A wasp is any insect of the narrow-waisted suborder Apocrita of the order Hymenoptera which is neither a bee nor an ant; this excludes the broad-waisted sawflies (Symphyta), which look somewhat like wasps, but are in a separate suborder ...

stingless bee Stingless bees (SB), sometimes called stingless honey bees or simply meliponines, are a large group of bees (from about 462 to 552 described species), comprising the Tribe (biology), tribe Meliponini (or subtribe Meliponina according to other aut ...

s and

honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the ...

s, which process it into a dark, strong

honey Honey is a sweet and viscous substance made by several species of bees, the best-known of which are honey bees. Honey is made and stored to nourish bee colonies. Bees produce honey by gathering and then refining the sugary secretions of pl ...

( honeydew honey). This is highly prized in parts of

Europe Europe is a continent located entirely in the Northern Hemisphere and mostly in the Eastern Hemisphere. It is bordered by the Arctic Ocean to the north, the Atlantic Ocean to the west, the Mediterranean Sea to the south, and Asia to the east ...

and

Asia Asia ( , ) is the largest continent in the world by both land area and population. It covers an area of more than 44 million square kilometres, about 30% of Earth's total land area and 8% of Earth's total surface area. The continent, which ...

for its reputed medicinal value. '' Parachartergus fraternus'', a

eusocial Eusociality ( Greek 'good' and social) is the highest level of organization of sociality. It is defined by the following characteristics: cooperative brood care (including care of offspring from other individuals), overlapping generations wit ...

wasp species, collects honeydew to feed to their growing larvae. Recent research has also documented the use of honeydew by over 40 species of wild, native, mostly solitary bees in California.

Mythology

Norse mythology Norse, Nordic, or Scandinavian mythology, is the body of myths belonging to the North Germanic peoples, stemming from Old Norse religion and continuing after the Christianization of Scandinavia as the Nordic folklore of the modern period. The ...

dew Dew is water in the form of droplets that appears on thin, exposed objects in the morning or evening due to condensation. As the exposed surface cools by thermal radiation, radiating its heat, atmospheric moisture condenses at a rate grea ...

falls from the ash tree Yggdrasil to the earth, and according to the ''

Prose Edda The ''Prose Edda'', also known as the ''Younger Edda'', ''Snorri's Edda'' () or, historically, simply as ''Edda'', is an Old Norse textbook written in Iceland during the early 13th century. The work is often considered to have been to some exten ...

'' book ''

Gylfaginning ''Gylfaginning'' (Old Norse: 'The Beguiling of Gylfi' or 'The Deluding of Gylfi'; 13th century Old Norse pronunciation ) is the first main part of the 13th century ''Prose Edda'', after the initial Prologue. The ''Gylfaginning'' takes the form of ...

'', "this is what people call honeydew and from it bees feed." In

Greek mythology Greek mythology is the body of myths originally told by the Ancient Greece, ancient Greeks, and a genre of ancient Greek folklore, today absorbed alongside Roman mythology into the broader designation of classical mythology. These stories conc ...

, ''méli'', or "honey", drips from the Manna–ash (''

Fraxinus ornus ''Fraxinus ornus'', the manna ash or South European flowering ash, is a species of ''Fraxinus'' native to Southern Europe and Southwestern Asia, from Spain and Italy north to Austria and the Czech Republic, and east through the Balkans, Turkey, a ...

''), with which the ''

Meliae In Greek mythology, the Meliae (also called Meliads) (; or ) were usually considered to be the nymphs of the ash tree, whose name they shared. Mythology According to Hesiod, the Meliae (probably meaning all tree-nymphs) were born from the dr ...

'', or "ash tree

nymph A nymph (; ; sometimes spelled nymphe) is a minor female nature deity in ancient Greek folklore. Distinct from other Greek goddesses, nymphs are generally regarded as personifications of nature; they are typically tied to a specific place, land ...

s", nursed the infant god

Zeus Zeus (, ) is the chief deity of the List of Greek deities, Greek pantheon. He is a sky father, sky and thunder god in ancient Greek religion and Greek mythology, mythology, who rules as king of the gods on Mount Olympus. Zeus is the child ...

on the island of

Crete Crete ( ; , Modern Greek, Modern: , Ancient Greek, Ancient: ) is the largest and most populous of the Greek islands, the List of islands by area, 88th largest island in the world and the List of islands in the Mediterranean#By area, fifth la ...

(as in the ''Hymn to Zeus'' by

Callimachus Callimachus (; ; ) was an ancient Greek poet, scholar, and librarian who was active in Alexandria during the 3rd century BC. A representative of Ancient Greek literature of the Hellenistic period, he wrote over 800 literary works, most of which ...

). Honey-dew is referenced in the last lines of

Samuel Coleridge Samuel Taylor Coleridge ( ; 21 October 177225 July 1834) was an English poet, literary critic, philosopher, and theologian who was a founder of the Romantic Movement in England and a member of the Lake Poets with his friend William Wordswort ...

's poem '' Kubla Khan'', perhaps because of its mythological connotations:

Nectar producing trees

Dates

'' Ommatissus lybicus'' is attracted to certain

cultivars A cultivar is a kind of cultivated plant that people have selected for desired traits and which retains those traits when propagated. Methods used to propagate cultivars include division, root and stem cuttings, offsets, grafting, tissue cult ...

of the

date palm tree ''Phoenix dactylifera'', commonly known as the date palm, is a flowering-plant species in the palm family Arecaceae, cultivated for its edible sweet #Fruits, fruit called dates. The species is widely cultivated across North Africa, northern A ...

. The honeydew producing insects preferred the Medjhool variety to the Deglet Noor in

Israel Israel, officially the State of Israel, is a country in West Asia. It Borders of Israel, shares borders with Lebanon to the north, Syria to the north-east, Jordan to the east, Egypt to the south-west, and the Mediterranean Sea to the west. Isr ...

, where they have been observed in the Arava Valley. Very dense insect populations may have some adverse effects. Different methods of controlling the insects, including natural and chemical, have been studied.

Eucalyptus

eucalypt Eucalypt is any woody plant with Capsule (fruit), capsule fruiting bodies belonging to one of seven closely related genera (of the tribe Eucalypteae) found across Australia: ''Eucalyptus'', ''Corymbia'', ''Angophora'', ''Stockwellia'', ''Allosyn ...

forests, production of both the honeydew nectar and manna tends to increase in spring and autumn. ''

Eucalyptus ''Eucalyptus'' () is a genus of more than 700 species of flowering plants in the family Myrtaceae. Most species of ''Eucalyptus'' are trees, often Mallee (habit), mallees, and a few are shrubs. Along with several other genera in the tribe Eucalyp ...

'' can produce even more manna than honeydew nectar. The sugar glider eats both, licking the nectar from branches. Other species attracted to the nectar include the

feathertail glider The feathertail glider (''Acrobates pygmaeus''), also known as the pygmy gliding possum, pygmy glider, pygmy phalanger, flying phalanger and flying mouse, is a species of marsupial native to eastern Australia. It is the world's smallest gliding ...

, brush-tailed phascogale, and brown antechinus. Most trees are not able to produce sap if the phloem duct becomes damaged by mechanical processes.

Oaks

The acorn weevil ('' Curculio glandium'') habitually bores into the young

acorn The acorn is the nut (fruit), nut of the oaks and their close relatives (genera ''Quercus'', ''Notholithocarpus'' and ''Lithocarpus'', in the family Fagaceae). It usually contains a seedling surrounded by two cotyledons (seedling leaves), en ...

s of oak trees. This injury can cause the tree to release a sweet honeydew, thought to attract wasps that can parasitize the weevil within its acorns.

Honey bee A honey bee (also spelled honeybee) is a eusocial flying insect within the genus ''Apis'' of the bee clade, all native to mainland Afro-Eurasia. After bees spread naturally throughout Africa and Eurasia, humans became responsible for the ...

s sometimes collect this substance, in addition to honeydew from aphids feeding in oak forests, and use it to produce oak

, an unusual varietal sold by specialist

beekeeper A beekeeper is a person who keeps honey bees, a profession known as beekeeping. The term beekeeper refers to a person who keeps honey bees in beehives, boxes, or other receptacles. The beekeeper does not control the creatures. The beekeeper ow ...

Tamarisk

Two

scale insects Scale insects are small insects of the order Hemiptera, suborder Sternorrhyncha. Of dramatically variable appearance and extreme sexual dimorphism, they comprise the infraorder Coccomorpha which is considered a more convenient grouping than the ...

in the Sinai, '' Trabutina mannipara'' and '' Najacoccus serpentinus'', feed on Tamarisk trees. They secrete a sugary nectar that turns white when it hardens.

Honeydew honey

The kind of

made from when bees harvest off the honeydew of sap-eating insects is marked as honeydew honey or forest honey. It is notably darker and more viscous than typical honey. Honeydew honey typically contains more melezitose. It is prized in parts of Europe and

New Zealand New Zealand () is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and List of islands of New Zealand, over 600 smaller islands. It is the List of isla ...

References

External links

* {{Authority control Body fluids Insect products Sternorrhyncha