Homarus americanus
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The American lobster (''Homarus americanus'') is a
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of
lobster Lobsters are a family (Nephropidae, synonym Homaridae) of marine crustaceans. They have long bodies with muscular tails and live in crevices or burrows on the sea floor. Three of their five pairs of legs have claws, including the first pair, ...
found on the Atlantic coast of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, chiefly from
Labrador , nickname = "The Big Land" , etymology = , subdivision_type = Country , subdivision_name = Canada , subdivision_type1 = Province , subdivision_name1 ...
to
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delawa ...
. It is also known as Atlantic lobster, Canadian lobster, true lobster, northern lobster, Canadian Reds, or Maine lobster. It can reach a body length of , and a
mass Mass is an intrinsic property of a body. It was traditionally believed to be related to the quantity of matter in a physical body, until the discovery of the atom and particle physics. It was found that different atoms and different ele ...
of over , making it not only the heaviest
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
in the world, but also the heaviest of all living
arthropod Arthropods (, (gen. ποδός)) are invertebrate animals with an exoskeleton, a segmented body, and paired jointed appendages. Arthropods form the phylum Arthropoda. They are distinguished by their jointed limbs and cuticle made of chiti ...
species. Its closest relative is the European lobster ''
Homarus gammarus ''Homarus gammarus'', known as the European lobster or common lobster, is a species of clawed lobster from the eastern Atlantic Ocean, Mediterranean Sea and parts of the Black Sea. It is closely related to the American lobster, ''H. amer ...
'', which can be distinguished by its coloration and the lack of spines on the underside of the
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...
. American lobsters are usually bluish green to brown with red spines, but several color variations have been observed.


Distribution

''Homarus americanus'' is distributed along the
Atlantic The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
coast of
North America North America is a continent in the Northern Hemisphere and almost entirely within the Western Hemisphere. It is bordered to the north by the Arctic Ocean, to the east by the Atlantic Ocean, to the southeast by South America and th ...
, from
Labrador , nickname = "The Big Land" , etymology = , subdivision_type = Country , subdivision_name = Canada , subdivision_type1 = Province , subdivision_name1 ...
in the north to
Cape Hatteras Cape Hatteras is a cape located at a pronounced bend in Hatteras Island, one of the barrier islands of North Carolina. Long stretches of beach, sand dunes, marshes, and maritime forests create a unique environment where wind and waves shap ...
,
North Carolina North Carolina () is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. The state is the List of U.S. states and territories by area, 28th largest and List of states and territories of the United ...
, in the south. South of
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delawa ...
, the species is uncommon, and landings in
Delaware Delaware ( ) is a state in the Mid-Atlantic region of the United States, bordering Maryland to its south and west; Pennsylvania to its north; and New Jersey and the Atlantic Ocean to its east. The state takes its name from the adjacent ...
,
Maryland Maryland ( ) is a state in the Mid-Atlantic region of the United States. It shares borders with Virginia, West Virginia, and the District of Columbia to its south and west; Pennsylvania to its north; and Delaware and the Atlantic Ocean t ...
,
Virginia Virginia, officially the Commonwealth of Virginia, is a state in the Mid-Atlantic and Southeastern regions of the United States, between the Atlantic Coast and the Appalachian Mountains. The geography and climate of the Commonwealth are ...
and North Carolina usually make up less than 0.1% of all landings. A
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...
assigned to ''Homarus americanus'' was found at
Nantucket Nantucket () is an island about south from Cape Cod. Together with the small islands of Tuckernuck and Muskeget, it constitutes the Town and County of Nantucket, a combined county/town government that is part of the U.S. state of Massachuse ...
, dating from the
Pleistocene The Pleistocene ( , often referred to as the ''Ice age'') is the geological Epoch (geology), epoch that lasted from about 2,580,000 to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fina ...
. In 2013, an American lobster was caught at the
Farallon Islands The Farallon Islands, or Farallones (from the Spanish ''farallón'' meaning "pillar" or "sea cliff"), are a group of islands and sea stacks in the Gulf of the Farallones, off the coast of San Francisco, California, United States. The isl ...
off the coast of
California California is a state in the Western United States, located along the Pacific Coast. With nearly 39.2million residents across a total area of approximately , it is the most populous U.S. state and the 3rd largest by area. It is also the m ...
. It has been introduced to
Norway Norway, officially the Kingdom of Norway, is a Nordic countries, Nordic country in Northern Europe, the mainland territory of which comprises the western and northernmost portion of the Scandinavian Peninsula. The remote Arctic island of ...
and potentially
Iceland Iceland ( is, Ísland; ) is a Nordic island country in the North Atlantic Ocean and in the Arctic Ocean. Iceland is the most sparsely populated country in Europe. Iceland's capital and largest city is Reykjavík, which (along with its ...
.


Description

''Homarus americanus'' commonly reaches long and weighs in weight, but has been known to weigh as much as , making this the heaviest crustacean in the world. Together with ''
Sagmariasus verreauxi ''Sagmariasus verreauxi'' is a species of spiny lobster that lives around northern New Zealand, the Kermadec Islands the Chatham Islands and Australia from Queensland to Tasmania. It is probably the longest decapod crustacean in the world, ...
'', it is also the longest decapod crustacean in the world; an average adult is about long and weighs . The longest American lobsters have a body (excluding claws) long. According to
Guinness World Records ''Guinness World Records'', known from its inception in 1955 until 1999 as ''The Guinness Book of Records'' and in previous United States editions as ''The Guinness Book of World Records'', is a reference book published annually, listing world ...
, the heaviest
crustacean Crustaceans (Crustacea, ) form a large, diverse arthropod taxon which includes such animals as decapoda, decapods, ostracoda, seed shrimp, branchiopoda, branchiopods, argulidae, fish lice, krill, remipedes, isopoda, isopods, barnacles, copepods, ...
ever recorded was an American lobster caught off Nova Scotia, Canada, weighing . The closest relative of ''H. americanus'' is the European lobster, ''
Homarus gammarus ''Homarus gammarus'', known as the European lobster or common lobster, is a species of clawed lobster from the eastern Atlantic Ocean, Mediterranean Sea and parts of the Black Sea. It is closely related to the American lobster, ''H. amer ...
''. The two species are very similar, and can be crossed artificially, although hybrids are unlikely to occur in the wild since their ranges do not overlap. The two species can be distinguished by several characteristics: *The
rostrum Rostrum may refer to: * Any kind of a platform for a speaker: **dais **pulpit * Rostrum (anatomy), a beak, or anatomical structure resembling a beak, as in the mouthparts of many sucking insects * Rostrum (ship), a form of bow on naval ships * Ros ...
of ''H. americanus'' bears one or more spines on the underside, which are lacking in ''H. gammarus''. *The spines on the claws of ''H. americanus'' are red or red-tipped, while those of ''H. gammarus'' are white or white-tipped. *The underside of the claw of ''H. americanus'' is orange or red, while that of ''H. gammarus'' is creamy white or very pale red.


Head

The antennae measure about long and split into Y-shaped structures with pointed tips. Each tip exhibits a dense zone of hair tufts staggered in a zigzag arrangement. These hairs are covered with multiple nerve cells that can detect odors. Larger, thicker hairs found along the edges control the flow of water, containing odor molecules, to the inner sensory hairs. The shorter
antennule Antennae ( antenna), sometimes referred to as "feelers", are paired appendages used for sensing in arthropods. Antennae are connected to the first one or two segments of the arthropod head. They vary widely in form but are always made of one o ...
s provide a further sense of smell. By having a pair of
olfactory The sense of smell, or olfaction, is the special sense through which smells (or odors) are perceived. The sense of smell has many functions, including detecting desirable foods, hazards, and pheromones, and plays a role in taste. In humans, ...
organs, a lobster can locate the direction a smell comes from, much the same way humans can hear the direction a sound comes from. In addition to sensing smells, the antennules can judge water speed to improve direction finding. Lobsters have two
urinary bladder The urinary bladder, or simply bladder, is a hollow organ in humans and other vertebrates that stores urine from the kidneys before disposal by urination. In humans the bladder is a distensible organ that sits on the pelvic floor. Urine ente ...
s, located on either side of the head. Lobsters use scents to communicate what and where they are, and those scents are in the
urine Urine is a liquid by-product of metabolism in humans and in many other animals. Urine flows from the kidneys through the ureters to the urinary bladder. Urination results in urine being excreted from the body through the urethra. Cellul ...
. They project long plumes of urine in front of them, and do so when they detect a rival or a potential mate in the area.


Thorax

The first pair of
pereiopod The decapod ( crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in variou ...
s (legs) is armed with a large, asymmetric pair of
claw A claw is a curved, pointed appendage found at the end of a toe or finger in most amniotes (mammals, reptiles, birds). Some invertebrates such as beetles and spiders have somewhat similar fine, hooked structures at the end of the leg or tarsus ...
s. The larger one is the "crusher", and has rounded nodules used for crushing prey; the other is the "cutter" or "gripper", which has sharp inner edges and is used for holding or tearing the prey. Whether the crusher claw is on the left side or right side of its body determines whether a lobster is left or right handed.


Coloration

The normal coloration of ''Homarus americanus'' is bluish green to brown with red spines due to a mixture of yellow, blue, and red pigments that occur naturally in the shell. On rare occasions these colors are distorted due to genetic mutations or conditions creating a spectacle for those who catch them. In 2012 it was reported that there has been an increase in these "rare" catches due to unclear reasons. Social media influence making reporting and sharing more accessible to a drop in predator populations have been suggested as possible reasons. The lobsters mentioned below thus usually receive media coverage due to their rarity and eye appeal.


Life cycle

Mating In biology, mating is the pairing of either opposite- sex or hermaphroditic organisms for the purposes of sexual reproduction. ''Fertilization'' is the fusion of two gametes. ''Copulation'' is the union of the sex organs of two sexually reprod ...
only takes place shortly after the female has molted and her
exoskeleton An exoskeleton (from Greek ''éxō'' "outer" and ''skeletós'' "skeleton") is an external skeleton that supports and protects an animal's body, in contrast to an internal skeleton ( endoskeleton) in for example, a human. In usage, some of the ...
is still soft. The female releases a
pheromone A pheromone () is a secreted or excreted chemical factor that triggers a social response in members of the same species. Pheromones are chemicals capable of acting like hormones outside the body of the secreting individual, to affect the behavio ...
which causes the males to become less aggressive and to begin
courtship Courtship is the period wherein some couples get to know each other prior to a possible marriage. Courtship traditionally may begin after a betrothal and may conclude with the celebration of marriage. A courtship may be an informal and private m ...
, which involves a courtship dance with claws closed. Eventually, the male inserts
spermatophore A spermatophore or sperm ampulla is a capsule or mass containing spermatozoa created by males of various animal species, especially salamanders and arthropods, and transferred in entirety to the female's ovipore during reproduction. Spermatophore ...
s (
sperm Sperm is the male reproductive cell, or gamete, in anisogamous forms of sexual reproduction (forms in which there is a larger, female reproductive cell and a smaller, male one). Animals produce motile sperm with a tail known as a flagellum, ...
packets) into the female's seminal receptacle using his first
pleopod The decapod ( crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in variou ...
s; the female may store the sperm for up to 15 months. The female releases
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
through her oviducts, and they pass the seminal receptacle and are
fertilized Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a new individual organism or offspring and initiate its development. Proc ...
by the stored sperm. They are then attached to the female's pleopods (swimmerets) using an adhesive, where they are cared for until they are ready to hatch. The female cleans the eggs regularly and fans them with water to keep them oxygenated. The large telolecithal eggs may resemble the segments of a
raspberry The raspberry is the edible fruit of a multitude of plant species in the genus ''Rubus'' of the rose family, most of which are in the subgenus '' Idaeobatus''. The name also applies to these plants themselves. Raspberries are perennial with ...
, and a female carrying eggs is said to be "in berry". Since this period lasts 10–11 months, berried females can be found at any time of year. In the waters off
New England New England is a region comprising six states in the Northeastern United States: Connecticut, Maine, Massachusetts, New Hampshire, Rhode Island, and Vermont. It is bordered by the state of New York to the west and by the Canadian provinces ...
, the eggs are typically laid in July or August, and hatch the following May or June. The developing
embryo An embryo is an initial stage of development of a multicellular organism. In organisms that reproduce sexually, embryonic development is the part of the life cycle that begins just after fertilization of the female egg cell by the male spe ...
passes through several molts within the egg, before hatching as a
metanauplius Metanauplius is an early larval stage of some crustaceans such as krill. It follows the nauplius stage. In sac-spawning krill, there is an intermediary phase called pseudometanauplius, a newly hatched form distinguished from older metanauplii b ...
larva. When the eggs hatch, the female releases them by waving her tail in the water, setting batches of larvae free. The metanauplius of ''H. americanus'' is long, transparent, with large eyes and a long spine projecting from its head. It quickly molts, and the next three stages are similar, but larger. These molts take 10–20 days, during which the
plankton Plankton are the diverse collection of organisms found in water (or air) that are unable to propel themselves against a current (or wind). The individual organisms constituting plankton are called plankters. In the ocean, they provide a cruc ...
ic larvae are vulnerable to predation; only 1 in 1,000 is thought to survive to the juvenile stage. To reach the fourth stage – the post-larva – the larva undergoes
metamorphosis Metamorphosis is a biological process by which an animal physically develops including birth or hatching, involving a conspicuous and relatively abrupt change in the animal's body structure through cell growth and differentiation. Some inse ...
, and subsequently shows a much greater resemblance to the adult lobster, is around long, and swims with its
pleopod The decapod ( crustaceans such as a crab, lobster, shrimp or prawn) is made up of 20 body segments grouped into two main body parts: the cephalothorax and the pleon (abdomen). Each segment may possess one pair of appendages, although in variou ...
s. At this stage, the lobster's claws are still relatively small so they rely primarily on tail-flip escapes if threatened. After the next molt, the lobster sinks to the ocean floor and adopts a
benthic The benthic zone is the ecological region at the lowest level of a body of water such as an ocean, lake, or stream, including the sediment surface and some sub-surface layers. The name comes from ancient Greek, βένθος (bénthos), meaning " ...
lifestyle. It molts more and more infrequently, from an initial rate of ten times per year to once every few years. After one year it is around long, and after six years it may weigh . By the time it reaches the
minimum landing size The minimum landing size (MLS) is the smallest fish measurement at which it is legal to keep or sell a fish. The MLS depends on the species of fish. Sizes also vary around the world, as they are legal definitions which are defined by the local regu ...
, an individual may have molted 25–27 times, and thereafter each molt may signal a 40%–50% increase in weight, and a 14% increase in
carapace A carapace is a dorsal (upper) section of the exoskeleton or shell in a number of animal groups, including arthropods, such as crustaceans and arachnids, as well as vertebrates, such as turtles and tortoises. In turtles and tortoises, the unde ...
length. If threatened, adult lobsters will generally choose to fight unless they have lost their claws.


Ecology

The American lobster thrives in cold, shallow waters where there are many rocks and other places to hide from predators. It typically lives at a depth of , but can be found up to below the surface.


Diet

The natural diet of ''H. americanus'' is relatively consistent across different habitats. It is dominated by
mollusks Mollusca is the second-largest phylum of invertebrate animals after the Arthropoda, the members of which are known as molluscs or mollusks (). Around 85,000  extant species of molluscs are recognized. The number of fossil species is esti ...
(especially
mussel Mussel () is the common name used for members of several families of bivalve molluscs, from saltwater and freshwater habitats. These groups have in common a shell whose outline is elongated and asymmetrical compared with other edible clams, which ...
s),
echinoderm An echinoderm () is any member of the phylum Echinodermata (). The adults are recognisable by their (usually five-point) radial symmetry, and include starfish, brittle stars, sea urchins, sand dollars, and sea cucumbers, as well as the ...
s and
polychaete Polychaeta () is a paraphyletic class of generally marine annelid worms, commonly called bristle worms or polychaetes (). Each body segment has a pair of fleshy protrusions called parapodia that bear many bristles, called chaetae, which are made ...
s, although a wide range of other prey items may be eaten, including other crustaceans,
brittle star Brittle stars, serpent stars, or ophiuroids (; ; referring to the serpent-like arms of the brittle star) are echinoderms in the class Ophiuroidea, closely related to starfish. They crawl across the sea floor using their flexible arms for locomot ...
s and
cnidaria Cnidaria () is a phylum under kingdom Animalia containing over 11,000 species of aquatic animals found both in freshwater and marine environments, predominantly the latter. Their distinguishing feature is cnidocytes, specialized cells that ...
ns. Lobsters in Maine have been shown to gain 35–55% of their calories from herring, which is used as bait for lobster traps. Only 6% of lobsters entering lobster traps to feed are caught.


Diseases


Bacterial

Gaffkaemia or red-tail is an extremely virulent infectious disease of lobsters caused by the bacterium ''Aerococcus viridans''. It only requires a few bacterial cells to cause death of otherwise healthy lobsters. The "red tail" common name refers to a dark orange discoloration of the ventral abdomen of affected lobsters. This is, in fact, the
hemolymph Hemolymph, or haemolymph, is a fluid, analogous to the blood in vertebrates, that circulates in the interior of the arthropod (invertebrate) body, remaining in direct contact with the animal's tissues. It is composed of a fluid plasma in which ...
or blood seen through the thin ventral arthrodial membranes. The red discoloration comes from
astaxanthin Astaxanthin is a keto-carotenoid within a group of chemical compounds known as terpenes. Astaxanthin is a metabolite of zeaxanthin and canthaxanthin, containing both hydroxyl and ketone functional groups. It is a lipid-soluble pigment with red ...
, a
carotenoid Carotenoids (), also called tetraterpenoids, are yellow, orange, and red organic pigments that are produced by plants and algae, as well as several bacteria, and fungi. Carotenoids give the characteristic color to pumpkins, carrots, parsnips, ...
pigment exported to the blood during times of stress. The same sign is also seen in other diseases of lobsters and appears to be a nonspecific stress response, possibly relating to the antioxidant and immunostimulatory properties of the astaxanthin molecule. Epizootic shell disease is a bacterial infection which causes black lesions on the lobsters' dorsal carapaces, reducing their saleability and sometimes killing the lobsters. Limp lobster disease caused by systemic infection by the bacterium ''
Vibrio fluvialis ''Vibrio fluvialis'' is a water-borne bacterium first isolated from patients with severe diarrhoea in Bahrain in the 1970s by A. L. Furniss and his colleagues, and is considered to be an emerging pathogen with the potential to have a significant ...
'' (or similar species) causes lobsters to become lethargic and die.


Parasitic

Paramoebiasis is an infectious disease of lobsters caused by infection with the sarcomastigophoran (amoeba) ''
Neoparamoeba pemaquidensis Neoparamoeba pemaquidensis is a single-celled species of marine amoebozoan Amoebozoa is a major taxonomic group containing about 2,400 described species of amoeboid protists, often possessing blunt, fingerlike, lobose pseudopods and tubular mi ...
''. This organism also causes
amoebic gill disease Amoebic gill disease (AGD) is a potentially fatal disease of some marine fish. It is caused by ''Neoparamoeba perurans'', the most important amoeba in cultured fish. It primarily affects farm raised fish of the family Salmonidae, most notably affe ...
in farmed
Atlantic salmon The Atlantic salmon (''Salmo salar'') is a species of ray-finned fish in the family Salmonidae. It is the third largest of the Salmonidae, behind Siberian taimen and Pacific Chinook salmon, growing up to a meter in length. Atlantic salmon are ...
, ''Salmo salar''. Infection occurs throughout the tissues, causing
granuloma A granuloma is an aggregation of macrophages that forms in response to chronic inflammation. This occurs when the immune system attempts to isolate foreign substances that it is otherwise unable to eliminate. Such substances include infectious o ...
-like lesions, especially within the ventral nerve cord, the interstices of the hepatopancreas and the antennal gland. Paramoebiasis is strongly suspected to play a prominent role in the rapid die-off of American lobsters in
Long Island Sound Long Island Sound is a marine sound and tidal estuary of the Atlantic Ocean. It lies predominantly between the U.S. state of Connecticut to the north and Long Island in New York to the south. From west to east, the sound stretches from the Eas ...
that occurred in the summer of 1999.


Environmental

Excretory calcinosis in American lobsters in Long Island Sound was described in 2002. The disease causes mineralized calculi to form in the antennal glands and gills. These cause a loss of surface area around the gills, and the lobster eventually asphyxiates. Several reasons have been proposed for the cause of a recent outbreak of the disease. The most generally attributed factor is an increased duration of warmer temperatures in the bottom of the Long Island Sound.
Plastic pollution Plastic pollution is the accumulation of plastic objects and particles (e.g. plastic bottles, bags and microbeads) in the Earth's environment that adversely affects humans, wildlife and their habitat. Plastics that act as pollutants are cate ...
is harmful for American lobsters. Consumption of microplastic particles may be deadly to early-stage larvae. For later stage larvae, oxygen consumption rate decreases with high level of microplastic fibers.


Taxonomy

The American lobster was first described by
Thomas Say Thomas Say (June 27, 1787 – October 10, 1834) was an American entomologist, conchologist, and herpetologist. His studies of insects and shells, numerous contributions to scientific journals, and scientific expeditions to Florida, Georgia, the R ...
in 1817, with a type locality of " Long-branch, part of the coast of
New Jersey New Jersey is a state in the Mid-Atlantic and Northeastern regions of the United States. It is bordered on the north and east by the state of New York; on the east, southeast, and south by the Atlantic Ocean; on the west by the Delawa ...
". The name Say chose – "''Astacus marinus''" – was invalid as a
junior homonym In biology, a homonym is a name for a taxon that is identical in spelling to another such name, that belongs to a different taxon. The rule in the International Code of Zoological Nomenclature is that the first such name to be published is the se ...
of ''Astacus marinus'' Fabricius, 1775, which is in turn a
junior synonym The Botanical and Zoological Codes of nomenclature treat the concept of synonymy differently. * In botanical nomenclature, a synonym is a scientific name that applies to a taxon that (now) goes by a different scientific name. For example, Linn ...
of ''
Homarus gammarus ''Homarus gammarus'', known as the European lobster or common lobster, is a species of clawed lobster from the eastern Atlantic Ocean, Mediterranean Sea and parts of the Black Sea. It is closely related to the American lobster, ''H. amer ...
''. The American lobster was given its current
scientific name In taxonomy, binomial nomenclature ("two-term naming system"), also called nomenclature ("two-name naming system") or binary nomenclature, is a formal system of naming species of living things by giving each a name composed of two parts, bo ...
of ''Homarus americanus'' by
Henri Milne-Edwards Henri Milne-Edwards (23 October 1800 – 29 July 1885) was an eminent French zoologist. Biography Henri Milne-Edwards was the 27th child of William Edwards, an English planter and colonel of the militia in Jamaica and Elisabeth Vaux, a French ...
in his 1837 work ' (''"Natural History of the Crustacea"''). The
common name In biology, a common name of a taxon or organism (also known as a vernacular name, English name, colloquial name, country name, popular name, or farmer's name) is a name that is based on the normal language of everyday life; and is often contrast ...
preferred by the
Food and Agriculture Organization The Food and Agriculture Organization of the United Nations (FAO)french: link=no, Organisation des Nations unies pour l'alimentation et l'agriculture; it, Organizzazione delle Nazioni Unite per l'Alimentazione e l'Agricoltura is an intern ...
is "American lobster", but the species is also known locally as the "northern lobster", "Maine lobster" or simply "lobster".


As food

American lobsters are a popular food. They are commonly boiled or steamed. Hard-shells (lobsters that are several months past their last molt) can survive out of water for up to four or five days if kept refrigerated. Soft-shells (lobsters that have only recently molted) do not survive more than a few hours out of water. Lobsters are usually cooked alive, which may be illegal in certain areas and which some people consider inhumane. One common way of serving lobster 'tail' (actually the
abdomen The abdomen (colloquially called the belly, tummy, midriff, tucky or stomach) is the part of the body between the thorax (chest) and pelvis, in humans and in other vertebrates. The abdomen is the front part of the abdominal segment of the to ...
) is with
beef Beef is the culinary name for meat from cattle (''Bos taurus''). In prehistoric times, humankind hunted aurochs and later domesticated them. Since that time, numerous breeds of cattle have been bred specifically for the quality or quantit ...
, known as
surf and turf Surf and turf or surf 'n' turf is a main course combining seafood and red meat. A typical seafood component would be lobster (either lobster tail or a whole lobster), prawns, shrimp, squid or scallops, any of which could be steamed, grilled or ...
. Lobsters have a greenish or brownish organ called the
tomalley Tomalley (from the Carib word ', meaning a sauce of lobster liver), crab fat, or lobster paste is the soft, green substance found in the body cavity of lobsters, that fulfills the functions of both the liver and the pancreas. Tomalley correspo ...
, which, like the
liver The liver is a major organ only found in vertebrates which performs many essential biological functions such as detoxification of the organism, and the synthesis of proteins and biochemicals necessary for digestion and growth. In humans, it i ...
and
pancreas The pancreas is an organ of the digestive system and endocrine system of vertebrates. In humans, it is located in the abdomen behind the stomach and functions as a gland. The pancreas is a mixed or heterocrine gland, i.e. it has both an ...
in a human, filters out
toxin A toxin is a naturally occurring organic poison produced by metabolic activities of living cells or organisms. Toxins occur especially as a protein or conjugated protein. The term toxin was first used by organic chemist Ludwig Brieger (1849 ...
s from the body. Some diners consider it a delicacy, but others avoid it because they consider it a toxin source; dislike eating innards; or are put off by its texture and appearance, that of a grainy greenish paste. A set of nutcrackers and a long, thin tool for pulling meat from inaccessible areas are suggested as basics, although more experienced diners can eat the animal with their bare hands or a simple tool (a fork, knife or rock). Eating a lobster can get messy, and most restaurants offer a lobster bib. Meat is generally contained in the larger claws and tails, and stays warm quite a while after being served. There is some meat in the legs and in the arms that connect the large claws to the body. There is also some small amount of meat just below the carapace around the thorax and in the smaller legs.


North American lobster industry

Most lobsters come from the northeastern coast of North America, with the
Atlantic Provinces Atlantic Canada, also called the Atlantic provinces (french: provinces de l'Atlantique), is the region of Eastern Canada comprising the provinces located on the Atlantic coast, excluding Quebec. The four provinces are New Brunswick, Newfoundlan ...
of Canada and the
U.S. state In the United States, a state is a constituent political entity, of which there are 50. Bound together in a political union, each state holds governmental jurisdiction over a separate and defined geographic territory where it shares its sove ...
of
Maine Maine () is a state in the New England and Northeastern regions of the United States. It borders New Hampshire to the west, the Gulf of Maine to the southeast, and the Canadian provinces of New Brunswick and Quebec to the northeast and nor ...
being the largest producers. They are caught primarily using
lobster trap A lobster trap or lobster pot is a portable trap that traps lobsters or crayfish and is used in lobster fishing. In Scotland (chiefly in the north), the word creel is used to refer to a device used to catch lobsters and other crustaceans. A lo ...
s, although lobsters are also harvested as
bycatch Bycatch (or by-catch), in the fishing industry, is a fish or other marine species that is caught unintentionally while fishing for specific species or sizes of wildlife. Bycatch is either the wrong species, the wrong sex, or is undersized or juve ...
by bottom trawlers, fishermen using
gillnet Gillnetting is a fishing method that uses gillnets: vertical panels of netting that hang from a line with regularly spaced floaters that hold the line on the surface of the water. The floats are sometimes called "corks" and the line with corks is ...
s, and by scuba divers in some areas. Maine prohibits scuba divers from catching lobsters; violations are punishable by fines of up to $1000. Maine also prohibits the landing of lobsters caught by bottom trawlers and other "mobile gear". Massachusetts offers scuba divers lobster licenses for a fee, and they are only available to state residents. Rhode Island also requires divers to acquire a permit. Lobster traps are rectangular cages made of vinyl-coated galvanized steel mesh or wood, with woven mesh entrances. These are baited and lowered to the sea floor. They allow a lobster to enter, but make it difficult for the larger specimens to turn around and exit. This allows the creatures to be captured alive. The traps, sometimes referred to as "pots", have a
buoy A buoy () is a floating device that can have many purposes. It can be anchored (stationary) or allowed to drift with ocean currents. Types Navigational buoys * Race course marker buoys are used for buoy racing, the most prevalent form of y ...
floating on the surface, and lobstermen check their traps between one and seven days after setting them. The inefficiency of the trapping system has inadvertently prevented the lobster population from being overfished. Lobsters can easily escape the trap, and will defend the trap against other lobsters because it is a source of food. An estimated 10% of lobsters that encounter a trap enter, and of those that enter 6% will be caught.


United States

In the United States, the lobster industry is
regulated Regulation is the management of complex systems according to a set of rules and trends. In systems theory, these types of rules exist in various fields of biology and society, but the term has slightly different meanings according to context. ...
. Every lobster fisher is required to use a lobster gauge to measure the distance from the lobster's eye socket to the end of its carapace: if the lobster is less than long, it is too young to be sold and must be released back to the sea. There is also a legal maximum size of in Maine, meant to ensure the survival of a healthy breeding stock of adult males, but in parts of some states, such as Massachusetts, there is none. Also, traps must contain an escape hole or "vent", which allows juvenile lobsters and bycatch species to escape. The law in Maine and other states dictates a second large escape hole or "ghost panel" must be installed. This hole is held shut through use of degradable clips made of ferrous metal. Should the trap become lost, the trap eventually opens, allowing the catch to escape. To protect known breeding females, lobsters caught carrying eggs are to be notched on a tail flipper (second from the right, if the lobster is right-side up and the tail is fully extended). Following this, the female cannot be kept or sold, and is commonly referred to as a "punch-tail" or as "v-notched". This notch remains for two molts of the lobster exoskeleton, providing harvest protection and continued breeding availability for up to five years.


Canada

In Canada, the
Department of Fisheries and Oceans Fisheries and Oceans Canada (DFO; french: Pêches et Océans Canada, MPO), is a department of the Government of Canada that is responsible for developing and implementing policies and programs in support of Canada's economic, ecological and sc ...
is responsible for the governance of fisheries under the authority of th
Fisheries Act
The governance structure also includes various other acts, regulations, orders and policies. American Lobster is fished in Canada by lobster licence holders hailing from ports located in provinces on Canada's east coast. Lobster is Canada's most valuable seafood export, worth over CAD$2 billion in 2016.


Management

American lobster tends to have a stable stock in colder northern waters, but gradually decreases in abundance moving southward. To manage lobster populations, more regulations and restrictions, geared towards achieving sustainable populations, are implemented gradually southward.


Genetics

Currently there is no published
genome In the fields of molecular biology and genetics, a genome is all the genetic information of an organism. It consists of nucleotide sequences of DNA (or RNA in RNA viruses). The nuclear genome includes protein-coding genes and non-coding ...
for the American lobster, although a
transcriptome The transcriptome is the set of all RNA transcripts, including coding and non-coding, in an individual or a population of cells. The term can also sometimes be used to refer to all RNAs, or just mRNA, depending on the particular experiment. The t ...
was published in 2016.


See also

*


References


Further reading

* * *


External links

* {{Taxonbar, from=Q37668 Crustaceans described in 1837 Commercial crustaceans Crustaceans of North America Crustaceans of the Atlantic Ocean Crustaceans of the United States Edible crustaceans Extant Pleistocene first appearances Least concern biota of North America Least concern biota of the United States Natural history of Maine Pleistocene animals of North America True lobsters