Harris hawk
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The Harris's hawk (''Parabuteo unicinctus''), formerly known as the bay-winged hawk, dusky hawk, and sometimes a wolf hawk, and known in Latin America as peuco, is a medium-large
bird of prey Birds of prey or predatory birds, also known as raptors, are hypercarnivorous bird species that actively hunt and feed on other vertebrates (mainly mammals, reptiles and other smaller birds). In addition to speed and strength, these predat ...
that breeds from the
southwestern United States The Southwestern United States, also known as the American Southwest or simply the Southwest, is a geographic and cultural region of the United States that generally includes Arizona, New Mexico, and adjacent portions of California, Colorado, N ...
south to
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, central
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, and
Brazil Brazil ( pt, Brasil; ), officially the Federative Republic of Brazil (Portuguese: ), is the largest country in both South America and Latin America. At and with over 217 million people, Brazil is the world's fifth-largest country by area ...
. Birds are sometimes reported at large in
Western Europe Western Europe is the western region of Europe. The region's countries and territories vary depending on context. The concept of "the West" appeared in Europe in juxtaposition to "the East" and originally applied to the ancient Mediterranean ...
, especially
Britain Britain most often refers to: * The United Kingdom, a sovereign state in Europe comprising the island of Great Britain, the north-eastern part of the island of Ireland and many smaller islands * Great Britain, the largest island in the United King ...
, but it is a popular species in
falconry Falconry is the hunting of wild animals in their natural state and habitat by means of a trained bird of prey. Small animals are hunted; squirrels and rabbits often fall prey to these birds. Two traditional terms are used to describe a person ...
and these records almost invariably all refer to escapes from
captivity Captivity, or being held captive, is a state wherein humans or other animals are confined to a particular space and prevented from leaving or moving freely. An example in humans is imprisonment. Prisoners of war are usually held in captivity by a ...
. The name is derived from the
Greek Greek may refer to: Greece Anything of, from, or related to Greece, a country in Southern Europe: *Greeks, an ethnic group. *Greek language, a branch of the Indo-European language family. **Proto-Greek language, the assumed last common ancestor ...
''para'', meaning beside, near or like, and the
Latin Latin (, or , ) is a classical language belonging to the Italic languages, Italic branch of the Indo-European languages. Latin was originally a dialect spoken in the lower Tiber area (then known as Latium) around present-day Rome, but through ...
''buteo'', referring to a kind of buzzard; ''uni'' meaning once; and ''cinctus'' meaning girdled, referring to the white band at the tip of the tail.
John James Audubon John James Audubon (born Jean-Jacques Rabin; April 26, 1785 – January 27, 1851) was an American self-trained artist, naturalist, and ornithologist. His combined interests in art and ornithology turned into a plan to make a complete pictori ...
gave this bird its English name in honor of his ornithological companion, financial supporter, and friend Edward Harris. The Harris's hawk is notable for its behavior of hunting cooperatively in packs consisting of tolerant groups, while other raptors often hunt alone. Harris hawks' social nature has been attributed to their intelligence, which makes them easy to train and have made them a popular bird for use in falconry.Beebe, Frank (1984). ''A Falconry Manual.'' Hancock House Publishers, , page 81.


Description

This medium-large hawk is roughly intermediate in size between a
peregrine falcon The peregrine falcon (''Falco peregrinus''), also known as the peregrine, and historically as the duck hawk in North America, is a cosmopolitan bird of prey (raptor) in the family Falconidae. A large, crow-sized falcon, it has a blue-grey bac ...
(''Falco peregrinus'') and a
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members wit ...
(''Buteo jamaicensis''). Harris's hawks range in length from and generally have a wingspan of about . These hawks have a brownish plumage, reddish shoulders and tail feathers with a white base and white tip. They exhibit
sexual dimorphism Sexual dimorphism is the condition where the sexes of the same animal and/or plant species exhibit different morphological characteristics, particularly characteristics not directly involved in reproduction. The condition occurs in most an ...
with the females being larger by about 35%. In the United States, the average weight for adult males is about , with a range of , while the adult female average is , with a range of .Hamerstrom, F. (1978). "External sex characters of Harris' Hawks in winter". ''Raptor Res.'' 12:1–14. They have dark brown plumage with chestnut shoulders, wing linings, and thighs, white on the base and tip of the tail, long, yellow legs and a yellow
cere The beak, bill, or rostrum is an external anatomical structure found mostly in birds, but also in turtles, non-avian dinosaurs and a few mammals. A beak is used for eating, preening, manipulating objects, killing prey, fighting, probing for food ...
. The vocalizations of the Harris's hawk are very harsh sounds. The lifespan of the Harris's Hawk is 10-–12 years in the wild, 20–25 years in captivity.


Juvenile

The juvenile Harris's hawk is mostly streaked with
buff Buff or BUFF may refer to: People * Buff (surname), a list of people * Buff (nickname), a list of people * Johnny Buff, ring name of American world champion boxer John Lisky (1888–1955) * Buff Bagwell, a ring name of American professional ...
, and appears much lighter than the dark adults. When in flight, the undersides of the juveniles' wings are buff-colored with brown streaking. They can look unlike adults at first glance, but the identical chestnut plumage is an aid for identification.


Subspecies

There are three subspecies of the Harris's hawk: * ''P. u. superior'': found in
Baja California Baja California (; 'Lower California'), officially the Free and Sovereign State of Baja California ( es, Estado Libre y Soberano de Baja California), is a state in Mexico. It is the northernmost and westernmost of the 32 federal entities of Mex ...
,
Arizona Arizona ( ; nv, Hoozdo Hahoodzo ; ood, Alĭ ṣonak ) is a state in the Southwestern United States. It is the 6th largest and the 14th most populous of the 50 states. Its capital and largest city is Phoenix. Arizona is part of the Fou ...
, Sonora, and Sinaloa. ''P. u. superior'' was believed to have longer tails and wings and to be more blackish than ''P. u. harrisi''. However, the sample size of the original study was quite small, with only five males and six females. Later research has concluded that there is not as strong a physical difference as was originally assumed. Other ecological differences, such as latitudinal cline were also brought up as arguments against the validity of the subspecies segmentation. * ''P. u. harrisi'': found in
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, eastern
Mexico Mexico (Spanish: México), officially the United Mexican States, is a country in the southern portion of North America. It is bordered to the north by the United States; to the south and west by the Pacific Ocean; to the southeast by Guatema ...
, and much of
Central America Central America ( es, América Central or ) is a subregion of the Americas. Its boundaries are defined as bordering the United States to the north, Colombia to the south, the Caribbean Sea to the east, and the Pacific Ocean to the west. ...
. * ''P. u. unicinctus'': found exclusively in
South America South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sout ...
. It is smaller than the North American subspecies.


Distribution and habitat

Harris's hawks live in sparse
woodland A woodland () is, in the broad sense, land covered with trees, or in a narrow sense, synonymous with wood (or in the U.S., the ''plurale tantum'' woods), a low-density forest forming open habitats with plenty of sunlight and limited shade (se ...
and semi-desert, as well as
marsh A marsh is a wetland that is dominated by herbaceous rather than woody plant species.Keddy, P.A. 2010. Wetland Ecology: Principles and Conservation (2nd edition). Cambridge University Press, Cambridge, UK. 497 p Marshes can often be found a ...
es (with some trees) in some parts of their
range Range may refer to: Geography * Range (geographic), a chain of hills or mountains; a somewhat linear, complex mountainous or hilly area (cordillera, sierra) ** Mountain range, a group of mountains bordered by lowlands * Range, a term used to i ...
(Howell and Webb 1995), including
mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...
swamps, as in parts of their South American range. Harris's hawks are permanent residents and do not migrate. Important perches and nest supports are provided by scattered larger trees or other features (e.g., power poles, woodland edges, standing dead trees, live trees, and boulders, and saguaros). The wild Harris's hawk population is declining due to
habitat loss Habitat destruction (also termed habitat loss and habitat reduction) is the process by which a natural habitat becomes incapable of supporting its native species. The organisms that previously inhabited the site are displaced or dead, thereby ...
; however, under some circumstances, they have been known to move into developed areas.


Behaviour

This species occurs in relatively stable groups. A dominance hierarchy occurs in Harris's hawks, wherein the mature female is the dominant bird, followed by the adult male and then the young of previous years. Groups typically include from two to seven birds. Not only do birds cooperate in hunting, they also assist in the nesting process. No other bird of prey is known to hunt in groups as routinely as this species.


Breeding

They nest in small trees, shrubby growth, or cacti. The
nests A nest is a structure built for certain animals to hold eggs or young. Although nests are most closely associated with birds, members of all classes of vertebrates and some invertebrates construct nests. They may be composed of organic materi ...
are often compact, made of sticks, plant roots, and stems, and are often lined with leaves, moss, bark and plant roots. They are built mainly by the female. There are usually two to four white to blueish white
eggs Humans and human ancestors have scavenged and eaten animal eggs for millions of years. Humans in Southeast Asia had domesticated chickens and harvested their eggs for food by 1,500 BCE. The most widely consumed eggs are those of fowl, especial ...
sometimes with a speckling of pale
brown Brown is a color. It can be considered a composite color, but it is mainly a darker shade of orange. In the CMYK color model used in printing or painting, brown is usually made by combining the colors orange and black. In the RGB color model us ...
or gray. The nestlings start out light buff, but in five to six days turn a rich brown. Very often, there will be three hawks attending one nest: two males and one female. Whether or not this is
polyandry Polyandry (; ) is a form of polygamy in which a woman takes two or more husbands at the same time. Polyandry is contrasted with polygyny, involving one male and two or more females. If a marriage involves a plural number of "husbands and wives ...
is debated, as it may be confused with backstanding (one bird standing on another's back). The female does most of the incubation. The eggs hatch in 31 to 36 days. The young begin to explore outside the nest at 38 days, and fledge, or start to fly, at 45 to 50 days. The female sometimes breeds two or three times in a year. Young may stay with their parents for up to three years, helping to raise later broods. Nests are known to be predated by coyotes (''Canis latrans''),
golden eagle The golden eagle (''Aquila chrysaetos'') is a bird of prey living in the Northern Hemisphere. It is the most widely distributed species of eagle. Like all eagles, it belongs to the family Accipitridae. They are one of the best-known birds of ...
s (''Aquila chrysaetos''),
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members wit ...
s (''Buteo jamaicensis''),
great horned owl The great horned owl (''Bubo virginianus''), also known as the tiger owl (originally derived from early naturalists' description as the "winged tiger" or "tiger of the air"), or the hoot owl, is a large owl native to the Americas. It is an extre ...
s (''Bubo virginianus'') and flocks of
common raven The common raven (''Corvus corax'') is a large all-black passerine bird. It is the most widely distributed of all corvids, found across the Northern Hemisphere. It is a raven known by many names at the subspecies level; there are at least ...
s (''Corvus corax''), predators possibly too formidable to be fully displaced by the Harris's hawk's cooperative nest defenses. No accounts show predation on adults in the United States and the Harris's hawk may be considered an
apex predator An apex predator, also known as a top predator, is a predator at the top of a food chain, without natural predators of its own. Apex predators are usually defined in terms of trophic dynamics, meaning that they occupy the highest trophic lev ...
, although presumably predators like eagles and great horned owls would be capable of killing them. In
Chile Chile, officially the Republic of Chile, is a country in the western part of South America. It is the southernmost country in the world, and the closest to Antarctica, occupying a long and narrow strip of land between the Andes to the eas ...
, black-chested buzzard-eagles (''Geranoaetus melanoleucus'') are likely predators.


Feeding

The diet of the Harris's hawk consists of small creatures including
bird Birds are a group of warm-blooded vertebrates constituting the class Aves (), characterised by feathers, toothless beaked jaws, the laying of hard-shelled eggs, a high metabolic rate, a four-chambered heart, and a strong yet lightweig ...
s, lizards, mammals, and large
insect Insects (from Latin ') are pancrustacean hexapod invertebrates of the class Insecta. They are the largest group within the arthropod phylum. Insects have a chitinous exoskeleton, a three-part body ( head, thorax and abdomen), three ...
s. Because it often hunts in groups, the Harris's hawk can also take down larger prey. Although not particularly common, the Harris's hawk may take prey weighing over , such as adult
jackrabbits Hares and jackrabbits are mammals belonging to the genus ''Lepus''. They are herbivores, and live solitarily or in pairs. They nest in slight depressions called forms, and their young are able to fend for themselves shortly after birth. The gen ...
, great blue heron (''Ardea herodias'') and half-grown
wild turkey The wild turkey (''Meleagris gallopavo'') is an upland ground bird native to North America, one of two extant species of turkey and the heaviest member of the order Galliformes. It is the ancestor to the domestic turkey, which was originally d ...
s (''Meleagris gallapavo''). The
desert cottontail The desert cottontail (''Sylvilagus audubonii''), also known as Audubon's cottontail, is a New World cottontail rabbit, and a member of the family Leporidae. Unlike the European rabbit (''Oryctolagus cuniculus''), they do not form social burrow s ...
(''Syvilagus auduboni''), the leading prey species in the north of the Harris's hawk's range, usually weighs or less. Undoubtedly because it pursues large prey often, this hawk has larger and stronger feet, with long talons, and a larger, more prominent hooked beak than most other raptors around its size. Locally, other buteonine hawks, including the
ferruginous hawk The ferruginous hawk, (''Buteo regalis''), is a large bird of prey and belongs to the broad-winged buteo hawks. An old colloquial name is ferrugineous rough-leg, due to its similarity to the closely related rough-legged hawk (''B. lagopus''). ...
, the
red-tailed hawk The red-tailed hawk (''Buteo jamaicensis'') is a bird of prey that breeds throughout most of North America, from the interior of Alaska and northern Canada to as far south as Panama and the West Indies. It is one of the most common members wit ...
and the
white-tailed hawk The white-tailed hawk (''Geranoaetus albicaudatus'') is a large bird of prey species found in tropical and subtropical environments of the Americas. Description The white-tailed hawk is a large, stocky hawk. It is close in size to the Swainso ...
also hunt primarily cottontails and jackrabbits, but each are bigger, weighing about , and , respectively, more on average than a Harris's hawk. In the Southwestern United States, the most common prey species (in descending order of prevalence) are
desert cottontail The desert cottontail (''Sylvilagus audubonii''), also known as Audubon's cottontail, is a New World cottontail rabbit, and a member of the family Leporidae. Unlike the European rabbit (''Oryctolagus cuniculus''), they do not form social burrow s ...
(''Syvilagus auduboni''),
eastern cottontail The eastern cottontail (''Sylvilagus floridanus'') is a New World cottontail rabbit, a member of the family Leporidae. It is the most common rabbit species in North America. Distribution The eastern cottontail can be found in meadows and shrubb ...
(''Syvilagus floridanus''),
black-tailed jackrabbit The black-tailed jackrabbit (''Lepus californicus''), also known as the American desert hare, is a common hare of the western United States and Mexico, where it is found at elevations from sea level up to . Reaching a length around , and a ...
(''Lepus californicus''),
ground squirrel Ground squirrels are members of the squirrel family of rodents ( Sciuridae), which generally live on or in the ground, rather than trees. The term is most often used for the medium-sized ground squirrels, as the larger ones are more commonly known ...
s (''Ammopsermophilus'' spp. and ''Spermophilus'' spp.), woodrats (''Neotoma'' spp.),
kangaroo rat Kangaroo rats, small mostly nocturnal rodents of genus ''Dipodomys'', are native to arid areas of western North America. The common name derives from their bipedal form. They hop in a manner similar to the much larger kangaroo, but developed t ...
s (''Dipodomys'' spp.),
pocket gopher Pocket gophers, commonly referred to simply as gophers, are burrowing rodents of the family Geomyidae. The roughly 41 speciesSearch results for "Geomyidae" on thASM Mammal Diversity Database are all endemic to North and Central America. They are ...
s (''Geomys'' and ''Thomomys'' spp.),
Gambel's quail Gambel's quail (''Callipepla gambelii'') is a small ground-dwelling bird in the New World quail family. It inhabits the desert regions of Arizona, California, Colorado, New Mexico, Nevada, Utah, Texas, and Sonora; also New Mexico-border Chihuahu ...
(''Callipepla gambelii''),
scaled quail The scaled quail (''Callipepla squamata''), also commonly called blue quail or cottontop, is a species of the New World quail family. It is a bluish gray bird found in the arid regions of the Southwestern United States to Central Mexico. This sp ...
(''C. squamata''),
northern bobwhite The northern bobwhite (''Colinus virginianus''), also known as the Virginia quail or (in its home range) bobwhite quail, is a ground-dwelling bird native to Canada, the United States, Mexico, and Cuba, with introduced populations elsewhere in th ...
(''Colinus virginianus''),
cactus wren The cactus wren (''Campylorhynchus brunneicapillus'') is a species of wren endemic to the deserts of the southwestern United States and northern and central Mexico. It is the state bird of Arizona, and the largest wren in the United States. ...
(''Campylorhynchus brunneicapillus''),
northern mockingbird The northern mockingbird (''Mimus polyglottos'') is a mockingbird commonly found in North America. This bird is mainly a permanent resident, but northern birds may move south during harsh weather. This species has rarely been observed in Europe ...
(''Mimus polyglottos''), desert spiny lizards (''Sceloporus magister''), and skinks (''Eumeces'' spp.) In the tropics, Harris's hawks have adapted to taking prey of several varieties, including those like
chicken The chicken (''Gallus gallus domesticus'') is a domesticated junglefowl species, with attributes of wild species such as the grey and the Ceylon junglefowl that are originally from Southeastern Asia. Rooster or cock is a term for an adu ...
s and
European rabbit The European rabbit (''Oryctolagus cuniculus'') or coney is a species of rabbit native to the Iberian Peninsula (including Spain, Portugal, and southwestern France), western France, and the northern Atlas Mountains in northwest Africa. It has b ...
s introduced by man. In Chile, the
common degu The common degu (''Octodon degus''; ), or, historically, the degu, is a small hystricomorpha rodent endemic to the Chilean matorral ecoregion of central Chile. The name ''degu'' on its own indicates either the entire genus ''Octodon'' or, more ...
(''Octodon degus'') makes up 67.5% of the prey.


Hunting

While most raptors are solitary, only coming together for breeding and migration, Harris's hawks will hunt in cooperative groups of two to six. This is believed to be an adaptation to the lack of prey in the desert climate in which they live. In one hunting technique, a small group flies ahead and scouts, then another group member flies ahead and scouts, and this continues until prey is bagged and shared. In another, all the hawks spread around the prey and one bird flushes it out. Harris's hawks will often chase prey on foot, and are quite fast on the ground and their long, yellow legs are adapted for this, as most hawks don't spend as much time on the ground. Groups of Harris's hawks tend to be more successful at capturing prey than lone hawks, with groups of two to four individuals having ~10% higher success rates per extra individual.


Relationship with humans


Falconry

Since about 1980, Harris's hawks have been increasingly used in
falconry Falconry is the hunting of wild animals in their natural state and habitat by means of a trained bird of prey. Small animals are hunted; squirrels and rabbits often fall prey to these birds. Two traditional terms are used to describe a person ...
and are now the most popular hawks in the West (outside of Asia) for that purpose, as they are one of the easiest to train and the most social. Trained Harris's hawks have been used to remove an unwanted
pigeon Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
population from
London London is the capital and List of urban areas in the United Kingdom, largest city of England and the United Kingdom, with a population of just under 9 million. It stands on the River Thames in south-east England at the head of a estuary dow ...
's
Trafalgar Square Trafalgar Square ( ) is a public square in the City of Westminster, Central London, laid out in the early 19th century around the area formerly known as Charing Cross. At its centre is a high column bearing a statue of Admiral Nelson comm ...
, and from the tennis courts at Wimbledon. Trained Harris's hawks have been used for
bird abatement Bird control or bird abatement involves the methods to eliminate or deter pest birds from landing, roosting and nesting. Bird control is important because pest birds can create health-related problems through their feces, including histoplasmos ...
by falconry experts in Canada and the United States at various locations including airports, resorts, landfill sites, and industrial sites. File:Harris's hawk in flight, Southern Ontario, Canada (captive).jpg, Harris's hawk in flight at a falconry centre File:Harris's hawk at a licensed falconry centre.jpg, Harris's hawk at a licensed falconry centre, Ontario, Canada


In art

John James Audubon John James Audubon (born Jean-Jacques Rabin; April 26, 1785 – January 27, 1851) was an American self-trained artist, naturalist, and ornithologist. His combined interests in art and ornithology turned into a plan to make a complete pictori ...
illustrated the Harris's hawk in ''
The Birds of America ''The Birds of America'' is a book by naturalist and painter John James Audubon, containing illustrations of a wide variety of birds of the United States. It was first published as a series in sections between 1827 and 1838, in Edinburgh and ...
'' (published in London, 1827–38) as Plate 392 with the title "Louisiana Hawk -''Buteo harrisi''". The image was
engraved Engraving is the practice of incising a design onto a hard, usually flat surface by cutting grooves into it with a burin. The result may be a decorated object in itself, as when silver, gold, steel, or glass are engraved, or may provide an in ...
and colored by the
Robert Havell The Havell family of Reading, Berkshire, England, included a number of notable engravers, etchers and painters, as well as writers, publishers, educators, and musicians. In particular, members of this family were among the foremost practitioners ...
, London workshops in 1837. The original
watercolor Watercolor (American English) or watercolour (British English; see spelling differences), also ''aquarelle'' (; from Italian diminutive of Latin ''aqua'' "water"), is a painting method”Watercolor may be as old as art itself, going back to ...
by Audubon was purchased by the New York History Society where it remains to this day (January 2009).


References


External links


Peregrine Fund page on Harris' hawk



Harris' hawks hunting in pack
(video)


Historical material

* John James Audubon.
Louisiana Hawk
, ''Ornithological Biography'' volume 5 (1839).
Illustration
from ''Birds of America'' octavo edition, 1840. {{Taxonbar, from=Q742698
Harris's hawk The Harris's hawk (''Parabuteo unicinctus''), formerly known as the bay-winged hawk, dusky hawk, and sometimes a wolf hawk, and known in Latin America as peuco, is a medium-large bird of prey that breeds from the southwestern United States south ...
Falconry Birds of Central America Birds of Mexico Birds of prey Birds of South America Native birds of the Southwestern United States
Harris's hawk The Harris's hawk (''Parabuteo unicinctus''), formerly known as the bay-winged hawk, dusky hawk, and sometimes a wolf hawk, and known in Latin America as peuco, is a medium-large bird of prey that breeds from the southwestern United States south ...
Taxa named by Coenraad Jacob Temminck