Harold Munro Fox (; 28 September 1889 – 29 January 1967) was an English

zoologist Zoology ( , ) is the scientific study of animals. Its studies include the structure, embryology, classification, habits, and distribution of all animals, both living and extinct, and how they interact with their ecosystems. Zoology is one ...

Education and early life

He was born Harold Munro Fuchs in

Clapham Clapham () is a district in south London, south west London, England, lying mostly within the London Borough of Lambeth, but with some areas (including Clapham Common) extending into the neighbouring London Borough of Wandsworth. History Ea ...

, London, in 1889 to George Gotthilf Fuchs, a former captain in the Prussian Army, and Margaret Isabella Campbell Munro, daughter of Lieutenant Colonel Andrew Munro of the

Yorkshire Regiment The Royal Yorkshire Regiment (14th/15th, 19th and 33rd/76th Foot) (abbreviated R YORKS) is an infantry regiment of the British Army, created by the amalgamation of three historic regiments in 2006. It lost one battalion as part of the Future of ...

. However, his parents separated when he was just a few years old. Fox was educated at

Brighton College Brighton College is a fee-charging, co-educational, boarding and day public school for boys and girls aged 3 to 18 in Brighton and Hove, England. The school has three sites: Brighton College (the senior school, ages 11 to 18), Brighton Co ...

and

Gonville and Caius College, Cambridge Gonville and Caius College, commonly known as Caius ( ), is a constituent college of the University of Cambridge in Cambridge, England. Founded in 1348 by Edmund Gonville, it is the fourth-oldest of the University of Cambridge's 31 colleges and ...

, where he read for the

Natural Sciences Tripos The Natural Sciences Tripos is the framework within which most of the science at the University of Cambridge is taught. The tripos includes a wide range of Natural Sciences from physics, astronomy, and geoscience, to chemistry and biology, whi ...

(1908–1911).

Career

After graduation he went to the Plymouth Laboratory of the

Marine Biological Association of the United Kingdom The Marine Biological Association of the United Kingdom (MBA) is a learned society with a scientific laboratory that undertakes research in marine biology. The organisation was founded in 1884 and has been based in Plymouth since the Citadel Hil ...

(1911–1912), where he worked with Cresswell Shearer and Walter de Morgan on the genetics of

sea urchin Sea urchins or urchins () are echinoderms in the class (biology), class Echinoidea. About 950 species live on the seabed, inhabiting all oceans and depth zones from the intertidal zone to deep seas of . They typically have a globular body cove ...

hybrids. After his year in

Plymouth Plymouth ( ) is a port city status in the United Kingdom, city and unitary authority in Devon, South West England. It is located on Devon's south coast between the rivers River Plym, Plym and River Tamar, Tamar, about southwest of Exeter and ...

, he went to

Naples Naples ( ; ; ) is the Regions of Italy, regional capital of Campania and the third-largest city of Italy, after Rome and Milan, with a population of 908,082 within the city's administrative limits as of 2025, while its Metropolitan City of N ...

Italy Italy, officially the Italian Republic, is a country in Southern Europe, Southern and Western Europe, Western Europe. It consists of Italian Peninsula, a peninsula that extends into the Mediterranean Sea, with the Alps on its northern land b ...

, in 1912, where he worked on

fertilisation Fertilisation or fertilization (see spelling differences), also known as generative fertilisation, syngamy and impregnation, is the fusion of gametes to give rise to a zygote and initiate its development into a new individual organism or of ...

at the

Stazione Zoologica The Stazione Zoologica Anton Dohrn is a research institute in Naples, Italy, devoted to basic research in biology. Research is largely interdisciplinary involving the fields of evolution, biochemistry, molecular biology, neurobiology, cell biol ...

for ten months. In 1913 he was appointed lecturer in zoology at the Royal College of Science, London, by Ernest William MacBride. When the

First World War World War I or the First World War (28 July 1914 – 11 November 1918), also known as the Great War, was a World war, global conflict between two coalitions: the Allies of World War I, Allies (or Entente) and the Central Powers. Fighting to ...

broke out, he enlisted in the

Royal Army Service Corps The Royal Army Service Corps (RASC) was a corps of the British Army responsible for land, coastal and lake transport, air despatch, barracks administration, the Army Fire Service, staffing headquarters' units, supply of food, water, fuel and do ...

and served with the City of London Yeomanry in the

Balkans The Balkans ( , ), corresponding partially with the Balkan Peninsula, is a geographical area in southeastern Europe with various geographical and historical definitions. The region takes its name from the Balkan Mountains that stretch throug ...

Egypt Egypt ( , ), officially the Arab Republic of Egypt, is a country spanning the Northeast Africa, northeast corner of Africa and Western Asia, southwest corner of Asia via the Sinai Peninsula. It is bordered by the Mediterranean Sea to northe ...

Salonika Thessaloniki (; ), also known as Thessalonica (), Saloniki, Salonika, or Salonica (), is the second-largest city in Greece (with slightly over one million inhabitants in its Thessaloniki metropolitan area, metropolitan area) and the capital cit ...

(Thessaloniki), and

Palestine Palestine, officially the State of Palestine, is a country in West Asia. Recognized by International recognition of Palestine, 147 of the UN's 193 member states, it encompasses the Israeli-occupied West Bank, including East Jerusalem, and th ...

. It was then when he changed his name from Fuchs to Fox by

deed poll A deed poll (plural: deeds poll) is a legal document binding on a single person or several persons acting jointly to express an intention or create an obligation. It is a deed, and not a contract, because it binds only one party. Etymology Th ...

. While he was stationed in Egypt, he met his first wife, Léonie Thérèse Roger, the daughter of Henri Roger, a

French people French people () are a nation primarily located in Western Europe that share a common Culture of France, French culture, History of France, history, and French language, language, identified with the country of France. The French people, esp ...

official of the

Suez Canal Company Suez (, , , ) is a seaport city with a population of about 800,000 in north-eastern Egypt, located on the north coast of the Gulf of Suez on the Red Sea, near the southern terminus of the Suez Canal. It is the capital and largest city of the ...

. They married in 1917. After the war, Fox spent six months back in London working with MacBride again, and also worked at the Marine Biological Association again for some time. However, in 1919 he returned to Cairo on invitation by Edward Hindle to join his staff at the Cairo School of Medicine as lecturer (1919–1923). During this time he finished a thesis on the

flagellate A flagellate is a cell or organism with one or more whip-like appendages called flagella. The word ''flagellate'' also describes a particular construction (or level of organization) characteristic of many prokaryotes and eukaryotes and the ...

protozoa Protozoa (: protozoan or protozoon; alternative plural: protozoans) are a polyphyletic group of single-celled eukaryotes, either free-living or parasitic, that feed on organic matter such as other microorganisms or organic debris. Historically ...

n ''

Bodo Bodo may refer to: Ethnicity * Boro people, also called ''Bodo'', an ethno-linguistic group mainly from Northwest Assam, India * Bodo-Kachari people, an umbrella group from Nepal, India and Bangladesh that includes the Boro people Culture an ...

'', the research on which he had begun at Plymouth, and presented it to

, to apply for

fellowship A fellow is a title and form of address for distinguished, learned, or skilled individuals in academia, medicine, research, and industry. The exact meaning of the term differs in each field. In learned or professional societies, the term refers ...

. His application was accepted in 1920, but he took up the post only in 1923. In Cambridge, he and John Stanley Gardiner organised an expedition to study the

fauna Fauna (: faunae or faunas) is all of the animal life present in a particular region or time. The corresponding terms for plants and fungi are ''flora'' and '' funga'', respectively. Flora, fauna, funga and other forms of life are collectively ...

of the

Suez Canal The Suez Canal (; , ') is an artificial sea-level waterway in Egypt, Indo-Mediterranean, connecting the Mediterranean Sea to the Red Sea through the Isthmus of Suez and dividing Africa and Asia (and by extension, the Sinai Peninsula from the rest ...

during 1924–1925. In 1927, Fox was appointed head of the Department of Zoology and Comparative Anatomy at the

University of Birmingham The University of Birmingham (informally Birmingham University) is a Public university, public research university in Birmingham, England. It received its royal charter in 1900 as a successor to Queen's College, Birmingham (founded in 1825 as ...

, a post he held until 1941. It was during his time in Birmingham that he and his wife became increasingly estranged from each other and eventually divorced. In 1931, Fox married his second wife, Natalia 'Natasha' Lvovna. He was Professor of Zoology,

Bedford College, London Bedford College was founded in London in 1849 as the first higher education college for women in the United Kingdom. In 1900, it became a constituent of the University of London. Having played a leading role in the advancement of women in highe ...

, from 1941 to 1954. Because the college had moved from London to Cambridge due to the outbreak of

World War II World War II or the Second World War (1 September 1939 – 2 September 1945) was a World war, global conflict between two coalitions: the Allies of World War II, Allies and the Axis powers. World War II by country, Nearly all of the wo ...

, he was able to work with his former colleagues from Gonville and Caius College again. On retirement he moved to Queen Mary College, London as a

research associate Research associates are researchers (scholars and professionals) that usually have an advanced degree beyond a Bachelor's degree such as a master's degree or a PhD. In some universities/research institutes, such as Harvard/Harvard Medical Scho ...

. His research focused on researching marine

invertebrate Invertebrates are animals that neither develop nor retain a vertebral column (commonly known as a ''spine'' or ''backbone''), which evolved from the notochord. It is a paraphyletic grouping including all animals excluding the chordata, chordate s ...

s such as

ostracod Ostracods, or ostracodes, are a Class (biology), class of the crustacean, Crustacea (class Ostracoda), sometimes known as seed shrimp. Some 33,000 species (only 13,000 of which are extant taxon, extant) have been identified,Brandão, S.N.; Antoni ...

crustacea Crustaceans (from Latin meaning: "those with shells" or "crusted ones") are invertebrate animals that constitute one group of arthropods that are traditionally a part of the subphylum Crustacea (), a large, diverse group of mainly aquatic arthrop ...

. He was elected a

Fellow of the Royal Society Fellowship of the Royal Society (FRS, ForMemRS and HonFRS) is an award granted by the Fellows of the Royal Society of London to individuals who have made a "substantial contribution to the improvement of natural science, natural knowledge, incl ...

in 1937 and won the society's

Darwin Medal The Darwin Medal is one of the medals awarded by the Royal Society for "distinction in evolution, biological diversity and developmental, population and organismal biology". In 1885, International Darwin Memorial Fund was transferred to the ...

in 1966. Fox died in St George's Hospital, London, on 29 January 1967.

Selected publications

*''The Personality of Animals'' (1941)

References

{{DEFAULTSORT:Fox, Harold Munro 1889 births 1967 deaths 20th-century English zoologists Academics of Bedford College, London Academics of Queen Mary University of London Academics of the University of Birmingham Alumni of Gonville and Caius College, Cambridge British Army personnel of World War I Fellows of the Royal Society Fullerian Professors of Physiology People educated at Brighton College People from Clapham Scientists from London Royal Army Service Corps soldiers City of London Yeomanry (Rough Riders) officers