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The Systellommatophora (synonym Gymnomorpha) is a
clade A clade (), also known as a monophyletic group or natural group, is a group of organisms that are monophyletic – that is, composed of a common ancestor and all its lineal descendants – on a phylogenetic tree. Rather than the English ter ...
of primitive, air-breathing
slug Slug, or land slug, is a common name for any apparently shell-less terrestrial gastropod mollusc. The word ''slug'' is also often used as part of the common name of any gastropod mollusc that has no shell, a very reduced shell, or only a ...
s, according to the
taxonomy of the Gastropoda (Bouchet & Rocroi, 2005) The taxonomy of the Gastropoda as it was revised in 2005 by Philippe Bouchet and Jean-Pierre Rocroi is a system for the scientific classification of gastropod mollusks. (Gastropods are a taxonomic class of animals which consists of snails and s ...
. They are
marine Marine is an adjective meaning of or pertaining to the sea or ocean. Marine or marines may refer to: Ocean * Maritime (disambiguation) * Marine art * Marine biology * Marine debris * Marine habitats * Marine life * Marine pollution Military ...
and
terrestrial Terrestrial refers to things related to land or the planet Earth. Terrestrial may also refer to: * Terrestrial animal, an animal that lives on land opposed to living in water, or sometimes an animal that lives on or near the ground, as opposed to ...
pulmonate Pulmonata or pulmonates, is an informal group (previously an Order (biology), order, and before that a Class (biology), subclass) of snails and slugs characterized by the ability to breathe air, by virtue of having a Respiratory system of gastro ...
gastropod The gastropods (), commonly known as snails and slugs, belong to a large taxonomic class of invertebrates within the phylum Mollusca called Gastropoda (). This class comprises snails and slugs from saltwater, from freshwater, and from land. T ...
s within the
Heterobranchia Heterobranchia, the ''heterobranchs'' (meaning "different-gilled snails"), is a taxonomic clade of snails and slugs, which includes marine, aquatic and terrestrial gastropod mollusks. Heterobranchia is one of the main clades of gastropods. Cur ...
. There are two superfamilies in this clade. Slugs in the superfamily Onchidioidea are primarily marine (except for five land-dwelling or freshwater species), whereas slugs in the superfamily
Veronicelloidea Veronicelloidea is a superfamily of air-breathing land slugs. They are terrestrial pulmonate gastropod molluscs in the clade Systellommatophora The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, accord ...
are primarily terrestrial.


Anatomy

No species in this clade have shells as adults. These slugs are distinguished by the location of the
anus The anus (Latin, 'ring' or 'circle') is an opening at the opposite end of an animal's digestive tract from the mouth. Its function is to control the expulsion of feces, the residual semi-solid waste that remains after food digestion, which, ...
at the rear of the body. Although neither superfamily bears shells as adults, the Onchidioidea do possess a vestigial, non-mineralized shell sac and possess a larval shell. It is not known whether or not the veronicellids bear a larval shell.


Taxonomy

According to the previous taxonomy of the Gastropoda (Ponder & Lindberg, 1997) Systellommatophora was considered to be a
suborder Order ( la, ordo) is one of the eight major hierarchical taxonomic ranks in Linnaean taxonomy. It is classified between family and class. In biological classification, the order is a taxonomic rank used in the classification of organisms and ...
or an order. The following two superfamilies and families have been recognized in the taxonomy of Bouchet & Rocroi (2005): * superfamily Onchidioidea
Rafinesque Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimat ...
, 1815 ** Onchidiidae
Rafinesque Constantine Samuel Rafinesque-Schmaltz (; October 22, 1783September 18, 1840) was a French 19th-century polymath born near Constantinople in the Ottoman Empire and self-educated in France. He traveled as a young man in the United States, ultimat ...
, 1815 * Superfamily
Veronicelloidea Veronicelloidea is a superfamily of air-breathing land slugs. They are terrestrial pulmonate gastropod molluscs in the clade Systellommatophora The Systellommatophora (synonym Gymnomorpha) is a clade of primitive, air-breathing slugs, accord ...
Gray, 1840 **
Veronicellidae The Veronicellidae, also known by their common name the leatherleaf slugs, are a family of pulmonate terrestrial slugs. The herbivorous molluscs occur mainly in the tropical and subtropical areas of America, Asia and Africa. They act as intermed ...
Gray, 1840 **
Rathouisiidae Rathouisiidae is a family of carnivorous air-breathing land slugs, terrestrial pulmonate gastropod mollusks in the superfamily Veronicelloidea, the leatherleaf slugs and their allies. This family has no subfamilies according to the ''Taxonomy of ...
Heude, 1885


References

{{Taxonbar, from=Q1319200 Panpulmonata