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The great albatrosses are
seabird Seabirds (also known as marine birds) are birds that are adapted to life within the marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent evolution, as the same envir ...
s in the
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Diomedea'' in the
albatross Albatrosses, of the biological family Diomedeidae, are large seabirds related to the procellariids, storm petrels, and diving petrels in the order Procellariiformes (the tubenoses). They range widely in the Southern Ocean and the North Pac ...
family Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
. The genus ''Diomedea'' formerly included all albatrosses except the
sooty albatross The sooty albatross, dark-mantled sooty albatross or dark-mantled albatross,BirdLife International (2008b) (''Phoebetria fusca''), is a species of bird in the albatross family. They breed on sub-Antarctic islands and range at sea across the S ...
es, but in 1996 the genus was split, with the
mollymawk The mollymawks are a group of medium-sized albatrosses that form the genus ''Thalassarche''. The name has sometimes been used for the genus ''Phoebetria'' as well, but these are usually called sooty albatrosses. They are restricted to the Sout ...
s and the North Pacific albatrosses both being elevated to separate genera. The great albatrosses themselves form two species complexes, the wandering and Amsterdam albatrosses, and the royal albatrosses. The splitting of the great albatrosses into six or seven
species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
has been accepted by most, though not all, authorities.


Description

The
wandering albatross The wandering albatross, snowy albatross, white-winged albatross or goonie (''Diomedea exulans'') is a large seabird from the family Diomedeidae, which has a circumpolar range in the Southern Ocean. It was the last species of albatross to be desc ...
and the southern royal albatross are the largest of the albatrosses and are among the largest of flying birds. They have the largest wingspans of any bird, being up to from tip to tip, although the average is a little over . Large adult males of these two species may exceed in weight, as heavy as a large
swan Swans are birds of the family Anatidae within the genus ''Cygnus''. The swans' closest relatives include the geese and ducks. Swans are grouped with the closely related geese in the subfamily Anserinae where they form the tribe Cygnini. Som ...
. The great albatrosses are predominantly white in
plumage Plumage ( "feather") is a layer of feathers that covers a bird and the pattern, colour, and arrangement of those feathers. The pattern and colours of plumage differ between species and subspecies and may vary with age classes. Within species, ...
as adults, with birds becoming whiter as they age. The two royal albatrosses at all ages and the larger, older male
wandering albatross The wandering albatross, snowy albatross, white-winged albatross or goonie (''Diomedea exulans'') is a large seabird from the family Diomedeidae, which has a circumpolar range in the Southern Ocean. It was the last species of albatross to be desc ...
es are totally white-bodied, while adult females and younger animals of the other species have dark pencilling marks on the edges of their feathers. Generally the smaller species or subspecies and the juveniles have more dark brown colour. The recently discovered
Amsterdam albatross The Amsterdam albatross or Amsterdam Island albatross, (''Diomedea amsterdamensis''), is a large albatross which breeds only on Amsterdam Island in the southern Indian Ocean. It was only described in 1983, and was thought by some researchers to ...
retains the dark brown plumage of juvenile birds into adulthood.


Habitat and range

The great albatrosses range across the
Southern Ocean The Southern Ocean, also known as the Antarctic Ocean, comprises the southernmost waters of the World Ocean, generally taken to be south of 60° S latitude and encircling Antarctica. With a size of , it is regarded as the second-smal ...
, and nest (for the most part) on isolated oceanic islands. The wandering albatrosses nest on islands around the Southern Ocean, from the
Atlantic Ocean The Atlantic Ocean is the second-largest of the world's five oceans, with an area of about . It covers approximately 20% of Earth's surface and about 29% of its water surface area. It is known to separate the " Old World" of Africa, Europe ...
(
South Georgia South Georgia ( es, Isla San Pedro) is an island in the South Atlantic Ocean that is part of the British Overseas Territory of South Georgia and the South Sandwich Islands. It lies around east of the Falkland Islands. Stretching in the east� ...
and
Tristan da Cunha Tristan da Cunha (), colloquially Tristan, is a remote group of volcano, volcanic islands in the South Atlantic Ocean. It is the Extreme points of Earth, most remote inhabited archipelago in the world, lying approximately from Cape Town, Sou ...
), to the
Indian Ocean The Indian Ocean is the third-largest of the world's five oceanic divisions, covering or ~19.8% of the water on Earth's surface. It is bounded by Asia to the north, Africa to the west and Australia to the east. To the south it is bounded by ...
and
New Zealand New Zealand ( mi, Aotearoa ) is an island country in the southwestern Pacific Ocean. It consists of two main landmasses—the North Island () and the South Island ()—and over 700 smaller islands. It is the sixth-largest island coun ...
's Subantarctic islands. The royal albatrosses nest only on New Zealand's Subantarctic islands, with one unusual colony on New Zealand's
Otago Peninsula The Otago Peninsula ( mi, Muaūpoko) is a long, hilly indented finger of land that forms the easternmost part of Dunedin, New Zealand. Volcanic in origin, it forms one wall of the eroded valley that now forms Otago Harbour. The peninsula lies sou ...
.


Taxonomy

The
genus Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nom ...
''Diomedea'' was introduced in 1758 by the Swedish naturalist
Carl Linnaeus Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
in the tenth edition of his ''
Systema Naturae ' (originally in Latin written ' with the ligature æ) is one of the major works of the Swedish botanist, zoologist and physician Carl Linnaeus (1707–1778) and introduced the Linnaean taxonomy. Although the system, now known as binomial ...
''. The genus name is from
Greek mythology A major branch of classical mythology, Greek mythology is the body of myths originally told by the ancient Greeks, and a genre of Ancient Greek folklore. These stories concern the origin and nature of the world, the lives and activities o ...
. When the hero
Diomedes Diomedes (Jones, Daniel; Roach, Peter, James Hartman and Jane Setter, eds. ''Cambridge English Pronouncing Dictionary''. 17th edition. Cambridge UP, 2006.) or Diomede (; grc-gre, Διομήδης, Diomēdēs, "god-like cunning" or "advised by ...
died his companions were turned into white seabirds. The
type species In zoological nomenclature, a type species (''species typica'') is the species name with which the name of a genus or subgenus is considered to be permanently taxonomically associated, i.e., the species that contains the biological type specim ...
was designated as the
wandering albatross The wandering albatross, snowy albatross, white-winged albatross or goonie (''Diomedea exulans'') is a large seabird from the family Diomedeidae, which has a circumpolar range in the Southern Ocean. It was the last species of albatross to be desc ...
(''Diomedea exulans'') by
George Robert Gray George Robert Gray FRS (8 July 1808 – 6 May 1872) was an English zoologist and author, and head of the ornithological section of the British Museum, now the Natural History Museum, in London for forty-one years. He was the younger brother ...
in 1840.


Species

The genus contains six species: * ''Diomedea epomophora'', southern royal albatross * ''Diomedea sanfordi'', northern royal albatross * ''Diomedea exulans'',
wandering albatross The wandering albatross, snowy albatross, white-winged albatross or goonie (''Diomedea exulans'') is a large seabird from the family Diomedeidae, which has a circumpolar range in the Southern Ocean. It was the last species of albatross to be desc ...
* ''Diomedea antipodensis'',
Antipodean albatross The Antipodean albatross (''Diomedea antipodensis'') is a large seabird in the albatross family. Antipodean albatrosses are smaller than wandering albatrosses, and breed in predominantly brown plumage, but are otherwise difficult to distinguish f ...
** ''Diomedea antipodensis gibsoni'',
Gibson's albatross Gibson's albatross (''Diomedea antipodensis gibsoni''), also known as the Auckland Islands wandering albatross or Gibson's wandering albatross, is a large seabird in the great albatross group of the albatross family. The common name and trinomi ...
* ''Diomedea dabbenena'', Tristan albatross * ''Diomedea amsterdamensis'',
Amsterdam albatross The Amsterdam albatross or Amsterdam Island albatross, (''Diomedea amsterdamensis''), is a large albatross which breeds only on Amsterdam Island in the southern Indian Ocean. It was only described in 1983, and was thought by some researchers to ...
The earliest known fossils of the genus are from the Middle
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, about 12–15 mya. By that time, the genera ''Phoebastria'' and ''Diomedea'' had already diverged. Fossil species * ''Diomedea milleri'' (Round Mountain Silt Middle Miocene of Sharktooth Hill and possibly Astoria Middle Miocene of Oregon, US) * ''Diomedea'' sp. (Late Miocene of Valdez Peninsula, Antarctica) * ''Diomedea'' sp. (Early Pliocene of South Africa) * ''Diomedea'' sp. (Early Pliocene of Bone Valley, Florida, US) * ''Diomedea thyridata'' Wilkinson, 1969 (Upper
Miocene The Miocene ( ) is the first geological epoch of the Neogene Period and extends from about (Ma). The Miocene was named by Scottish geologist Charles Lyell; the name comes from the Greek words (', "less") and (', "new") and means "less recen ...
, Beaumaris Bay Fossil Site, Australia) Beaumaris Bay Fossil Site, Beaumaris, VIC Profile
aussieheritage.com.au
At least four species were found in the Early Pliocene deposits of Lee Creek Mine,
North Carolina North Carolina () is a U.S. state, state in the Southeastern United States, Southeastern region of the United States. The state is the List of U.S. states and territories by area, 28th largest and List of states and territories of the United ...
in the US. These may in part be identical with the forms mentioned above. Assignment of the undescribed taxa to ''Diomedea'' is tentative since most of them were discovered before the splitting of this genus. Especially the Southern Hemisphere species probably belong to other genera.


See also

* List of albatross breeding locations


References


Further reading

* * Brooke, Michael (2004): ''Albatrosses and Petrels Across the World''. Oxford University Press, Oxford, New York. * Tickell, W.L.N. (2000): ''Albatrosses''. Pica Press, Sussex. {{DEFAULTSORT:Great Albatross