Poaceae () or Gramineae () is a large and nearly ubiquitous
family
Family (from la, familia) is a group of people related either by consanguinity (by recognized birth) or affinity (by marriage or other relationship). The purpose of the family is to maintain the well-being of its members and of society. Idea ...
of
monocotyledon
Monocotyledons (), commonly referred to as monocots, ( Lilianae '' sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one ...
ous
flowering plant
Flowering plants are plants that bear flowers and fruits, and form the clade Angiospermae (), commonly called angiosperms. The term "angiosperm" is derived from the Greek words ('container, vessel') and ('seed'), and refers to those plants t ...
s commonly known as grasses. It includes the
cereal
A cereal is any grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, germ, and bran. Cereal grain crops are grown in greater quantities and provide more food ...
grasses,
bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...
s and the grasses of natural
grassland
A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
and species cultivated in
lawn
A lawn is an area of soil-covered land planted with grasses and other durable plants such as clover which are maintained at a short height with a lawnmower (or sometimes grazing animals) and used for aesthetic and recreational purposes. ...
s and
pasture
Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, cattle, sheep, or sw ...
. The latter are commonly referred to collectively as grass.
With around 780
genera
Genus ( plural genera ) is a taxonomic rank used in the biological classification of living and fossil organisms as well as viruses. In the hierarchy of biological classification, genus comes above species and below family. In binomial nomenclat ...
and around 12,000 species,
the Poaceae is the fifth-largest
plant family, following the
Asteraceae
The family Asteraceae, alternatively Compositae, consists of over 32,000 known species of flowering plants in over 1,900 genera within the order Asterales. Commonly referred to as the aster, daisy, composite, or sunflower family, Compositae ...
,
Orchidaceae
Orchids are plants that belong to the family Orchidaceae (), a diverse and widespread group of flowering plants with blooms that are often colourful and fragrant.
Along with the Asteraceae, they are one of the two largest families of flowerin ...
,
and
Rubiaceae
The Rubiaceae are a family of flowering plants, commonly known as the coffee, madder, or bedstraw family. It consists of terrestrial trees, shrubs, lianas, or herbs that are recognizable by simple, opposite leaves with interpetiolar stipules a ...
.
The Poaceae are the most economically important plant family, providing
staple food
A staple food, food staple, or simply a staple, is a food that is eaten often and in such quantities that it constitutes a dominant portion of a standard Diet (nutrition), diet for a given person or group of people, supplying a large fraction of ...
s from domesticated
cereal
A cereal is any grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, germ, and bran. Cereal grain crops are grown in greater quantities and provide more food ...
crops such as
maize
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
,
wheat
Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
,
rice
Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and '' Porteresia'', both wild and domesticat ...
,
barley
Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley p ...
, and
millet
Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most species generally referred to as millets belong to the tribe Paniceae, but some millets a ...
as well as
feed for meat-producing animals. They provide, through direct human consumption, just over one-half (51%) of all dietary energy; rice provides 20%, wheat supplies 20%, maize (corn) 5.5%, and other
grain
A grain is a small, hard, dry fruit ( caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
s 6%. Some members of the Poaceae are used as building materials (
bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...
,
thatch
Thatching is the craft of building a roof with dry vegetation such as straw, water reed, sedge (''Cladium mariscus''), rushes, heather, or palm branches, layering the vegetation so as to shed water away from the inner roof. Since the bulk of ...
, and
straw
Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
); others can provide a source of
biofuel
Biofuel is a fuel that is produced over a short time span from biomass, rather than by the very slow natural processes involved in the formation of fossil fuels, such as oil. According to the United States Energy Information Administration ...
, primarily via the conversion of maize to
ethanol
Ethanol (abbr. EtOH; also called ethyl alcohol, grain alcohol, drinking alcohol, or simply alcohol) is an organic compound. It is an alcohol with the chemical formula . Its formula can be also written as or (an ethyl group linked to a ...
.
Grasses have
stem
Stem or STEM may refer to:
Plant structures
* Plant stem, a plant's aboveground axis, made of vascular tissue, off which leaves and flowers hang
* Stipe (botany), a stalk to support some other structure
* Stipe (mycology), the stem of a mushro ...
s that are hollow except at the
nodes and narrow alternate leaves borne in two ranks. The lower part of each leaf encloses the stem, forming a leaf-sheath. The leaf grows from the base of the blade, an adaptation allowing it to cope with frequent grazing.
Grassland
A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s such as
savannah
A savanna or savannah is a mixed woodland-grassland (i.e. grassy woodland) ecosystem characterised by the trees being sufficiently widely spaced so that the canopy does not close. The open canopy allows sufficient light to reach the ground to ...
and
prairie
Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
where grasses are dominant are estimated to constitute 40.5% of the land area of the
Earth
Earth is the third planet from the Sun and the only astronomical object known to harbor life. While large volumes of water can be found throughout the Solar System, only Earth sustains liquid surface water. About 71% of Earth's sur ...
, excluding
Greenland
Greenland ( kl, Kalaallit Nunaat, ; da, Grønland, ) is an island country in North America that is part of the Kingdom of Denmark. It is located between the Arctic and Atlantic oceans, east of the Canadian Arctic Archipelago. Greenland ...
and
Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
. Grasses are also an important part of the vegetation in many other habitats, including
wetland
A wetland is a distinct ecosystem that is flooded or saturated by water, either permanently (for years or decades) or seasonally (for weeks or months). Flooding results in oxygen-free (Anoxic waters, anoxic) processes prevailing, especially in t ...
s,
forest
A forest is an area of land dominated by trees. Hundreds of definitions of forest are used throughout the world, incorporating factors such as tree density, tree height, land use, legal standing, and ecological function. The United Nations' ...
s and
tundra
In physical geography, tundra () is a type of biome where tree growth is hindered by frigid temperatures and short growing seasons. The term ''tundra'' comes through Russian (') from the Kildin Sámi word (') meaning "uplands", "treeless mou ...
.
Though they are commonly called "grasses", groups such as the
seagrasses
Seagrasses are the only flowering plants which grow in marine environments. There are about 60 species of fully marine seagrasses which belong to four families (Posidoniaceae, Zosteraceae, Hydrocharitaceae and Cymodoceaceae), all in the or ...
,
rushes and
sedges fall outside this family. The rushes and sedges are related to the Poaceae, being members of the
order
Order, ORDER or Orders may refer to:
* Categorization, the process in which ideas and objects are recognized, differentiated, and understood
* Heterarchy, a system of organization wherein the elements have the potential to be ranked a number of ...
Poales
The Poales are a large order of flowering plants in the monocotyledons, and includes families of plants such as the grasses, bromeliads, and sedges. Sixteen plant families are currently recognized by botanists to be part of Poales.
Descriptio ...
, but the seagrasses are members of order
Alismatales
The Alismatales (alismatids) are an order of flowering plants including about 4,500 species. Plants assigned to this order are mostly tropical or aquatic. Some grow in fresh water, some in marine habitats.
Description
The Alismatales compr ...
. However, all of them belong to the
monocot
Monocotyledons (), commonly referred to as monocots, (Lilianae '' sensu'' Chase & Reveal) are grass and grass-like flowering plants (angiosperms), the seeds of which typically contain only one embryonic leaf, or cotyledon. They constitute one ...
group of plants.
Description
Grasses may be
annual
Annual may refer to:
*Annual publication, periodical publications appearing regularly once per year
** Yearbook
** Literary annual
*Annual plant
*Annual report
*Annual giving
*Annual, Morocco, a settlement in northeastern Morocco
*Annuals (band), ...
or
perennial
A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
herbs
In general use, herbs are a widely distributed and widespread group of plants, excluding vegetables and other plants consumed for macronutrients, with savory or aromatic properties that are used for flavoring and garnishing food, for medicina ...
,
[ generally with the following characteristics (the image gallery can be used for reference):
The stems of grasses, called culms, are usually cylindrical (more rarely flattened, but not 3-angled) and are hollow, plugged at the nodes, where the leaves are attached.] Grass leaves
A leaf ( : leaves) is any of the principal appendages of a vascular plant stem, usually borne laterally aboveground and specialized for photosynthesis. Leaves are collectively called foliage, as in "autumn foliage", while the leaves, st ...
are nearly always alternate and distichous (in one plane), and have parallel veins.[ Each leaf is differentiated into a lower sheath hugging the stem and a blade with entire (i.e., smooth) margins.][ The leaf blades of many grasses are hardened with ]silica
Silicon dioxide, also known as silica, is an oxide of silicon with the chemical formula , most commonly found in nature as quartz and in various living organisms. In many parts of the world, silica is the major constituent of sand. Silica is ...
phytolith
Phytoliths (from Greek, "plant stone") are rigid, microscopic structures made of silica, found in some plant tissues and persisting after the decay of the plant. These plants take up silica from the soil, whereupon it is deposited within different ...
s, which discourage grazing animals; some, such as sword grass, are sharp enough to cut human skin. A membranous appendage or fringe of hairs called the ligule A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf and leafstalk of many grasses (Poaceae) and sedges. A ligule is also a strap-shaped extension of the corolla, such as that of a ...
lies at the junction between sheath and blade, preventing water or insects from penetrating into the sheath.[
]Flower
A flower, sometimes known as a bloom or blossom, is the reproductive structure found in flowering plants (plants of the division Angiospermae). The biological function of a flower is to facilitate reproduction, usually by providing a mechanis ...
s of Poaceae are characteristically arranged in spikelet
A spikelet, in botany, describes the typical arrangement of the flowers of grasses, sedges and some other Monocots.
Each spikelet has one or more florets. The spikelets are further grouped into panicles or spikes. The part of the spikelet that ...
s, each having one or more florets.[ The spikelets are further grouped into panicles or spikes. The part of the spikelet that bears the florets is called the rachilla. A spikelet consists of two (or sometimes fewer) ]bracts
In botany, a bract is a modified or specialized leaf, especially one associated with a reproductive structure such as a flower, inflorescence axis or cone scale. Bracts are usually different from foliage leaves. They may be smaller, larger, or ...
at the base, called glumes
In botany, a glume is a bract (leaf-like structure) below a spikelet in the inflorescence (flower cluster) of grasses (Poaceae) or the flowers of sedges (Cyperaceae). There are two other types of bracts in the spikelets of grasses: the lemma an ...
, followed by one or more florets.[ A floret consists of the flower surrounded by two bracts, one external—the ]lemma
Lemma may refer to:
Language and linguistics
* Lemma (morphology), the canonical, dictionary or citation form of a word
* Lemma (psycholinguistics), a mental abstraction of a word about to be uttered
Science and mathematics
* Lemma (botany), ...
—and one internal—the palea. The flowers are usually hermaphroditic
In reproductive biology, a hermaphrodite () is an organism that has both kinds of reproductive organs and can produce both gametes associated with male and female sexes.
Many taxonomic groups of animals (mostly invertebrates) do not have s ...
—maize
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
being an important exception—and mainly anemophilous or wind-pollinated, although insects occasionally play a role. The perianth
The perianth (perigonium, perigon or perigone in monocots) is the non-reproductive part of the flower, and structure that forms an envelope surrounding the sexual organs, consisting of the calyx (sepals) and the corolla ( petals) or tepals when ...
is reduced to two scales, called '' lodicules'',[ that expand and contract to spread the lemma and palea; these are generally interpreted to be modified sepals. The ]fruit
In botany, a fruit is the seed-bearing structure in flowering plants that is formed from the ovary after flowering.
Fruits are the means by which flowering plants (also known as angiosperms) disseminate their seeds. Edible fruits in partic ...
of grasses is a caryopsis
In botany, a caryopsis (plural caryopses) is a type of simple fruit—one that is monocarpellate (formed from a single carpel) and indehiscent (not opening at maturity) and resembles an achene, except that in a caryopsis the pericarp is fuse ...
, in which the seed coat is fused to the fruit wall.[
A ]tiller
A tiller or till is a lever used to steer a vehicle. The mechanism is primarily used in watercraft, where it is attached to an outboard motor, rudder post or stock to provide leverage in the form of torque for the helmsman to turn the rudder ...
is a leafy shoot other than the first shoot produced from the seed.[
]
Growth and development
Grass blades grow at the base of the blade and not from elongated stem tips. This low growth point evolved in response to grazing animals and allows grasses to be grazed or mown regularly without severe damage to the plant.
Three general classifications of growth habit present in grasses: bunch-type (also called caespitose), stolon
In biology, stolons (from Latin '' stolō'', genitive ''stolōnis'' – "branch"), also known as runners, are horizontal connections between organisms. They may be part of the organism, or of its skeleton; typically, animal stolons are external s ...
iferous, and rhizomatous
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ho ...
.
The success of the grasses lies in part in their morphology and growth processes and in part in their physiological diversity. There are both C3 and C4 grasses, referring to the photosynthetic pathway for carbon fixation. The C4 grasses have a photosynthetic pathway, linked to specialized Kranz leaf anatomy, which allows for increased water use efficiency
Water-use efficiency (WUE) refers to the ratio of water used in plant metabolism to water lost by the plant through transpiration. Two types of water-use efficiency are referred to most frequently:
*photosynthetic water-use efficiency (also cal ...
, rendering them better adapted to hot, arid environments.
The C3 grasses are referred to as "cool-season" grasses, while the C4 plants are considered "warm-season" grasses.[
* Annual cool-season – ]wheat
Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
, rye, annual bluegrass (annual meadowgrass, '' Poa annua''), and oat
* Perennial cool-season – orchardgrass (cocksfoot, ''Dactylis glomerata
''Dactylis Glomerata'' is the sixth studio album by Swedish doom metal band Candlemass, released in 1998. This was their first album released since their split in 1994.
The album was originally destined to be the second release by Candlemass m ...
''), fescue (''Festuca
''Festuca'' (fescue) is a genus of flowering plants belonging to the grass family Poaceae (subfamily Pooideae). They are evergreen or herbaceous perennial tufted grasses with a height range of and a cosmopolitan distribution, occurring on ever ...
'' spp.), Kentucky bluegrass
''Poa pratensis'', commonly known as Kentucky bluegrass (or blue grass), smooth meadow-grass, or common meadow-grass, is a perennial species of grass native to practically all of Europe, North Asia and the mountains of Algeria and Morocco. Altho ...
and perennial ryegrass (''Lolium perenne
''Lolium perenne'', common name perennial ryegrass, English ryegrass, winter ryegrass, or ray grass, is a grass from the family Poaceae. It is native to Europe, Asia and northern Africa, but is widely cultivated and naturalised around the world.
...
'')
* Annual warm-season – maize
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
, sudangrass, and pearl millet
Pearl millet (''Cenchrus americanus'', commonly known as the synonym ''Pennisetum glaucum''; also known as 'Bajra' in Hindi, 'Sajje' in Kannada, 'Kambu' in Tamil, 'Bajeer' in Kumaoni and 'Maiwa' in Hausa, 'Mexoeira' in Mozambique) is the most ...
* Perennial warm-season – big bluestem
''Andropogon gerardi'', commonly known as big bluestem, is a species of tall grass native to much of the Great Plains and grassland regions of central and eastern North America. It is also known as tall bluestem, bluejoint, and turkeyfoot.
Taxon ...
, Indiangrass
''Sorghastrum nutans'', commonly known as either Indiangrass or yellow Indiangrass, is a North American prairie grass found in the central and eastern United States and Canada, especially in the Great Plains and tallgrass prairies.
Descripti ...
, Bermudagrass
''Cynodon dactylon'', commonly known as Bermuda grass, is a grass found worldwide. It is native to Europe, Africa, Australia and much of Asia. It has been introduced to the Americas. Although it is not native to Bermuda, it is an abundant invasi ...
and switchgrass
''Panicum virgatum'', commonly known as switchgrass, is a perennial warm season bunchgrass native to North America, where it occurs naturally from 55°N latitude in Canada southwards into the United States and Mexico. Switchgrass is one of the ...
.
Although the C4 species are all in the PACMAD clade (see diagram above), it seems that various forms of C4 have arisen some twenty or more times, in various subfamilies or genera. In the '' Aristida'' genus for example, one species (''A. longifolia'') is C3 but the approximately 300 other species are C4. As another example, the whole tribe of Andropogoneae
The Andropogoneae, sometimes called the sorghum tribe, are a large tribe of grasses (family Poaceae) with roughly 1,200 species in 90 genera, mainly distributed in tropical and subtropical areas. They include such important crops as maize (corn), ...
, which includes maize
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
, sorghum
''Sorghum'' () is a genus of about 25 species of flowering plants in the grass family (Poaceae). Some of these species are grown as cereals for human consumption and some in pastures for animals. One species is grown for grain, while many other ...
, sugar cane
Sugarcane or sugar cane is a species of (often hybrid) tall, perennial grass (in the genus '' Saccharum'', tribe Andropogoneae) that is used for sugar production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fibrous stalk ...
, " Job's tears", and bluestem grass
Bluestem can refer to various grasses:
* Little bluestem (''Schizachyrium scoparium'')
* Big bluestem (''Andropogon gerardii'') and other species of the genus '' Andropogon''
* Species in the genus ''Dichanthium''
* Cane bluestem and Caucasian b ...
es, is C4. Around 46 percent of grass species are C4 plants.
Taxonomy
The name Poaceae was given by John Hendley Barnhart
John Hendley Barnhart (October 4, 1871 – November 11, 1949) was an American botanist and author, specializing in biographies of botanists.Gleaston, H. A. John Hendley Barnhart—An appreciation. '' Journal of the New York Botanical Garden'' Augu ...
in 1895, based on the tribe Poeae described in 1814 by Robert Brown, and the type genus '' Poa'' described in 1753 by Carl Linnaeus
Carl Linnaeus (; 23 May 1707 – 10 January 1778), also known after his ennoblement in 1761 as Carl von Linné Blunt (2004), p. 171. (), was a Swedish botanist, zoologist, taxonomist, and physician who formalised binomial nomenclature, ...
. The term is derived from the Ancient Greek πόα (póa, "fodder").
Evolutionary history
Grasses include some of the most versatile plant life-forms. They became widespread toward the end of the Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
period, and fossilized dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
dung (coprolite
A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is ...
s) have been found containing phytolith
Phytoliths (from Greek, "plant stone") are rigid, microscopic structures made of silica, found in some plant tissues and persisting after the decay of the plant. These plants take up silica from the soil, whereupon it is deposited within different ...
s of a variety that include grasses that are related to modern rice
Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and '' Porteresia'', both wild and domesticat ...
and bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...
. Grasses have adapted to conditions in lush rain forest
Rainforests are characterized by a closed and continuous tree canopy, moisture-dependent vegetation, the presence of epiphytes and lianas and the absence of wildfire. Rainforest can be classified as tropical rainforest or temperate rainforest ...
s, dry desert
A desert is a barren area of landscape where little precipitation occurs and, consequently, living conditions are hostile for plant and animal life. The lack of vegetation exposes the unprotected surface of the ground to denudation. About on ...
s, cold mountains and even intertidal habitats, and are currently the most widespread plant type; grass is a valuable source of food and energy for all sorts of wildlife.
A cladogram shows subfamilies and approximate species numbers in brackets:[ ]
Before 2005, fossil findings indicated that grasses evolved around 55 million years ago. Finds of grass-like phytolith
Phytoliths (from Greek, "plant stone") are rigid, microscopic structures made of silica, found in some plant tissues and persisting after the decay of the plant. These plants take up silica from the soil, whereupon it is deposited within different ...
s in Cretaceous
The Cretaceous ( ) is a geological period that lasted from about 145 to 66 million years ago (Mya). It is the third and final period of the Mesozoic Era, as well as the longest. At around 79 million years, it is the longest geological period of ...
dinosaur coprolite
A coprolite (also known as a coprolith) is fossilized feces. Coprolites are classified as trace fossils as opposed to body fossils, as they give evidence for the animal's behaviour (in this case, diet) rather than morphology. The name is ...
s from the latest Cretaceous (Maastrichtian
The Maastrichtian () is, in the ICS geologic timescale, the latest age (uppermost stage) of the Late Cretaceous Epoch or Upper Cretaceous Series, the Cretaceous Period or System, and of the Mesozoic Era or Erathem. It spanned the inte ...
) aged Lameta Formation of India have pushed this date back to 66 million years ago. In 2011, fossils from the same deposit were found to belong to the modern rice tribe Oryzeae, suggesting substantial diversification of major lineages by this time.
Wu, You & Li (2018) described grass microfossils extracted from the teeth of the hadrosauroid dinosaur
Dinosaurs are a diverse group of reptiles of the clade Dinosauria. They first appeared during the Triassic period, between 243 and 233.23 million years ago (mya), although the exact origin and timing of the evolution of dinosaurs is t ...
'' Equijubus normani'' from northern China, dating to the Albian
The Albian is both an age of the geologic timescale and a stage in the stratigraphic column. It is the youngest or uppermost subdivision of the Early/Lower Cretaceous Epoch/ Series. Its approximate time range is 113.0 ± 1.0 Ma to 100.5 ± 0 ...
stage of the Early Cretaceous
The Early Cretaceous (geochronology, geochronological name) or the Lower Cretaceous (chronostratigraphy, chronostratigraphic name), is the earlier or lower of the two major divisions of the Cretaceous. It is usually considered to stretch from 145& ...
approximately 113–100 million years ago, which were found to belong to primitive lineages within Poaceae, similar in position to the Anomochlooideae. These are currently the oldest known grass fossils.
The relationships among the three subfamilies Bambusoideae, Oryzoideae and Pooideae in the BOP clade have been resolved: Bambusoideae and Pooideae are more closely related to each other than to Oryzoideae. This separation occurred within the relatively short time span of about 4 million years.
According to Lester Charles King
Lester Charles King (1907–1989) was an English geologist and geomorphologist known for his theories on scarp retreat. He offered a very different view of the origin of continental landscaping than that of William Morris Davis. Studying at univ ...
the spread of grasses in the Late Cenozoic would have changed patterns of hillslope evolution favouring slopes that are convex upslope and concave downslope and lacking a free face
Free may refer to:
Concept
* Freedom, having the ability to do something, without having to obey anyone/anything
* Freethought, a position that beliefs should be formed only on the basis of logic, reason, and empiricism
* Emancipate, to procur ...
were common. King argued that this was the result of more slowly acting surface wash caused by carpets of grass which in turn would have resulted in relatively more soil creep
Downhill creep, also known as soil creep or commonly just creep, is a type of creep characterized by the slow, downward progression of rock and soil down a low grade slope; it can also refer to slow deformation of such materials as a result of ...
.
Subdivisions
There are about 12,000 grass species in about 771 genera that are classified into 12 subfamilies.[ ] See the full list of Poaceae genera
The true grasses (Poaceae) are one of the largest plant families, with around 12,000 species and roughly 800 genera. They contain, among others, the cereal crop species and other plants of economic importance, such as the bamboos, and several imp ...
.
* Anomochlooideae
Anomochlooideae is a subfamily of the true grass family Poaceae. It is sister to all the other grasses. It includes perennial herbs that grow on the shaded floor of forests in the Neotropics
The Neotropical realm is one of the eight biogeogr ...
Pilg. ex Potztal, a small lineage of broad-leaved grasses that includes two genera (''Anomochloa'', ''Streptochaeta'')
* Pharoideae
Pharoideae is a subfamily of the true grass family Poaceae, containing the single tribe Phareae with the two genera '' Leptaspis'' and '' Pharus''. Its members grow on the shaded floors of tropical to warm temperate forests.
This subfamily i ...
L.G.Clark & Judz., a small lineage of grasses of three genera, including ''Pharus'' and ''Leptaspis''
* Puelioideae
The Puelioideae is a subfamily of the true grass family Poaceae with two genera, '' Guaduella'' and '' Puelia'', each in its own tribe. Its members grow in the understory of rainforests.
This subfamily is one of the earliest-diverging grass lin ...
L.G.Clark, M.Kobay., S.Mathews, Spangler
Spangler is a surname. Notable people with the surname include:
* Aaron Spangler (born 1971), American sculptor and printmaker
* Al Spangler (born 1933), American baseball player
*Amy Spangler (born 1949), American breastfeeding activist and consu ...
& E.A.Kellogg, a small lineage of the African genus ''Puelia''
* Pooideae
The Pooideae are the largest subfamily of the grass family Poaceae, with about 4,000 species in 15 tribes and roughly 200 genera. They include some major cereals such as wheat, barley, oat, rye and many lawn and pasture grasses. They are often ...
, including wheat
Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
, barley
Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley p ...
, oats, brome-grass (''Bromus
''Bromus'' is a large genus of grasses, classified in its own tribe Bromeae. They are commonly known as bromes, brome grasses, cheat grasses or chess grasses. Estimates in the scientific literature of the number of species have ranged from 100 ...
''), reed-grasses (''Calamagrostis'') and many lawn and pasture grasses such as bluegrass (''Poa'')
* Bambusoideae
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...
, including bamboo
Bamboos are a diverse group of evergreen perennial flowering plants making up the subfamily Bambusoideae of the grass family Poaceae. Giant bamboos are the largest members of the grass family. The origin of the word "bamboo" is uncertain, ...
* Ehrhartoideae
Oryzoideae ( syn. Ehrhartoideae) is a subfamily of the true grass family Poaceae. It has around 120 species in 19 genera, notably including the major cereal crop rice. Within the grasses, this subfamily is one of three belonging to the specie ...
, including rice
Rice is the seed of the grass species '' Oryza sativa'' (Asian rice) or less commonly ''Oryza glaberrima'' (African rice). The name wild rice is usually used for species of the genera '' Zizania'' and '' Porteresia'', both wild and domesticat ...
and wild rice
Wild rice, also called manoomin, Canada rice, Indian rice, or water oats, is any of four species of grasses that form the genus ''Zizania'', and the grain that can be harvested from them. The grain was historically gathered and eaten in both ...
* Aristidoideae
The Aristideae is the sole tribe of grasses in the monotypic subfamily Aristidoideae of the true grass family Poaceae. Its members are herbaceous annuals or perennials found in the tropics, subtropics and temperate zones. The tribe has over 300 ...
, including '' Aristida''
* Arundinoideae
The Arundinoideae are a subfamily of the true grass family Poaceae with around 40 species, including giant reed and common reed. Unlike many other members of the PACMAD clade of grasses, the Arundinoideae all use C3 photosynthesis. Their sister ...
, including giant reed and common reed
* Chloridoideae
Chloridoideae is one of the largest subfamilies of grasses, with roughly 150 genera and 1,600 species, mainly found in arid tropical or subtropical grasslands. Within the PACMAD clade, their sister group are the Danthonioideae. The subfamily ...
, including the lovegrasses (''Eragrostis'', about 350 species, including teff), dropseeds (''Sporobolus'', some 160 species), finger millet
''Eleusine coracana'', or finger millet, also known as ragi in India, kodo in Nepal, is an annual herbaceous plant widely grown as a cereal crop in the arid and semiarid areas in Africa and Asia. It is a tetraploid and self-pollinating species ...
(''Eleusine coracana'' (L.) Gaertn.), and the muhly grasses (''Muhlenbergia'', about 175 species)
* Panicoideae
Panicoideae is the second-largest subfamily of the grasses with over 3,500 species, mainly distributed in warm temperate and tropical regions. It comprises some important agricultural crops, including sugarcane, maize (or corn), sorghum, and swi ...
, including panic grass
''Panicum'' (panicgrass) is a large genus of about 450 species of grasses native throughout the tropical regions of the world, with a few species extending into the northern temperate zone. They are often large, annual or perennial grasses, growi ...
, maize
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American English, North American and Australian English), is a cereal grain first domesticated by indigenous peoples of Mexico, indigenous ...
, sorghum
''Sorghum'' () is a genus of about 25 species of flowering plants in the grass family (Poaceae). Some of these species are grown as cereals for human consumption and some in pastures for animals. One species is grown for grain, while many other ...
, sugarcane
Sugarcane or sugar cane is a species of (often hybrid) tall, perennial grass (in the genus '' Saccharum'', tribe Andropogoneae) that is used for sugar production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fibrous stalk ...
, most millet
Millets () are a highly varied group of small-seeded grasses, widely grown around the world as cereal crops or grains for fodder and human food. Most species generally referred to as millets belong to the tribe Paniceae, but some millets a ...
s, fonio, " Job's tears", and bluestem grass
Bluestem can refer to various grasses:
* Little bluestem (''Schizachyrium scoparium'')
* Big bluestem (''Andropogon gerardii'') and other species of the genus '' Andropogon''
* Species in the genus ''Dichanthium''
* Cane bluestem and Caucasian b ...
es
* Micrairoideae
* Danthonioideae, including pampas grass Pampas grass or pampas-grass is a common name which may refer to any of several similar-looking, tall-growing species of grass:
* Species of '' Cortaderia'' including:
:* ''Cortaderia selloana'' and its selected cultivars
:* ''Cortaderia jubata'' ( ...
Distribution
The grass family is one of the most widely distributed and abundant groups of plants on Earth
Earth is the third planet from the Sun and the only astronomical object known to harbor life. While large volumes of water can be found throughout the Solar System, only Earth sustains liquid surface water. About 71% of Earth's sur ...
. Grasses are found on every continent, including Antarctica
Antarctica () is Earth's southernmost and least-populated continent. Situated almost entirely south of the Antarctic Circle and surrounded by the Southern Ocean, it contains the geographic South Pole. Antarctica is the fifth-largest cont ...
. The Antarctic hair grass, '' Deschampsia antarctica'' is one of only two plant species native to the western Antarctic Peninsula
The Antarctic Peninsula, known as O'Higgins Land in Chile and Tierra de San Martín in Argentina, and originally as Graham Land in the United Kingdom and the Palmer Peninsula in the United States, is the northernmost part of mainland Antarctic ...
.
Ecology
Grasses are the dominant vegetation in many habitats, including grassland
A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
, salt-marsh
A salt marsh or saltmarsh, also known as a coastal salt marsh or a tidal marsh, is a coastal ecosystem in the upper coastal intertidal zone between land and open saltwater or brackish water that is regularly flooded by the tides. It is dominate ...
, reedswamp
A reedbed or reed bed is a natural habitat found in floodplains, waterlogged depressions and
estuaries. Reedbeds are part of a succession from young reeds colonising open water or wet ground through a gradation of increasingly dry ground. A ...
and steppes
In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes.
Steppe biomes may include:
* the montane grasslands and shrublands biome
* the temperate grassland ...
. They also occur as a smaller part of the vegetation in almost every other terrestrial habitat.
Grass-dominated biome
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
s are called grasslands. If only large, contiguous areas of grasslands are counted, these biomes cover 31% of the planet's land. Grasslands include pampas, steppe
In physical geography, a steppe () is an ecoregion characterized by grassland plains without trees apart from those near rivers and lakes.
Steppe biomes may include:
* the montane grasslands and shrublands biome
* the temperate gras ...
s, and prairie
Prairies are ecosystems considered part of the temperate grasslands, savannas, and shrublands biome by ecologists, based on similar temperate climates, moderate rainfall, and a composition of grasses, herbs, and shrubs, rather than trees, as the ...
s.
Grasses provide food to many grazing
In agriculture, grazing is a method of animal husbandry whereby domestic livestock are allowed outdoors to roam around and consume wild vegetations in order to convert the otherwise indigestible (by human gut) cellulose within grass and ot ...
mammals, as well as to many species
In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit of biodiversity. A species is often defined as the largest group of organisms in which any two individuals of the appropriat ...
of butterflies
Butterflies are insects in the macrolepidopteran clade Rhopalocera from the order Lepidoptera, which also includes moths. Adult butterflies have large, often brightly coloured wings, and conspicuous, fluttering flight. The group comprises ...
and moth
Moths are a paraphyletic group of insects that includes all members of the order Lepidoptera that are not butterflies, with moths making up the vast majority of the order. There are thought to be approximately 160,000 species of moth, many of w ...
s.
Many types of animals eat grass as their main source of food, and are called '' graminivores'' – these include cattle
Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult ...
, sheep
Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticate ...
, horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s, rabbit
Rabbits, also known as bunnies or bunny rabbits, are small mammals in the family Leporidae (which also contains the hares) of the order Lagomorpha (which also contains the pikas). ''Oryctolagus cuniculus'' includes the European rabbit sp ...
s and many invertebrate
Invertebrates are a paraphyletic group of animals that neither possess nor develop a vertebral column (commonly known as a ''backbone'' or ''spine''), derived from the notochord. This is a grouping including all animals apart from the chorda ...
s, such as grasshopper
Grasshoppers are a group of insects belonging to the suborder Caelifera. They are among what is possibly the most ancient living group of chewing herbivorous insects, dating back to the early Triassic around 250 million years ago.
Grasshopp ...
s and the caterpillars of many brown butterflies. Grasses are also eaten by omnivorous
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
or even occasionally by primarily carnivorous
A carnivore , or meat-eater (Latin, ''caro'', genitive ''carnis'', meaning meat or "flesh" and ''vorare'' meaning "to devour"), is an animal or plant whose food and energy requirements derive from animal tissues (mainly muscle, fat and other ...
animals.
Grasses dominate certain biomes
A biome () is a biogeographical unit consisting of a biological community that has formed in response to the physical environment in which they are found and a shared regional climate. Biomes may span more than one continent. Biome is a broader ...
, especially temperate grasslands, because many species are adapted to grazing and fire.
Grasses are unusual in that the meristem
The meristem is a type of tissue found in plants. It consists of undifferentiated cells (meristematic cells) capable of cell division. Cells in the meristem can develop into all the other tissues and organs that occur in plants. These cells conti ...
is near the bottom of the plant; hence, grasses can quickly recover from cropping at the top.
The evolution of large grazing animals in the Cenozoic
The Cenozoic ( ; ) is Earth's current geological era, representing the last 66million years of Earth's history. It is characterised by the dominance of mammals, birds and flowering plants, a cooling and drying climate, and the current configu ...
contributed to the spread of grasses. Without large grazers, fire-cleared areas are quickly colonized by grasses, and with enough rain, tree seedlings. Trees eventually outcompete most grasses. Trampling grazers kill seedling trees but not grasses.
Uses
Grasses are, in human terms, perhaps the most economically important plant family. Their economic importance stems from several areas, including food production, industry, and lawn
A lawn is an area of soil-covered land planted with grasses and other durable plants such as clover which are maintained at a short height with a lawnmower (or sometimes grazing animals) and used for aesthetic and recreational purposes. ...
s. They have been grown as food for domesticated animals for up to 6,000 years and the grains of grasses such as wheat
Wheat is a grass widely cultivated for its seed, a cereal grain that is a worldwide staple food. The many species of wheat together make up the genus ''Triticum'' ; the most widely grown is common wheat (''T. aestivum''). The archaeologi ...
, rice, maize (corn) and barley
Barley (''Hordeum vulgare''), a member of the grass family, is a major cereal grain grown in temperate climates globally. It was one of the first cultivated grains, particularly in Eurasia as early as 10,000 years ago. Globally 70% of barley p ...
have been the most important human food crop
A crop is a plant that can be grown and harvested extensively for profit or subsistence. When the plants of the same kind are cultivated at one place on a large scale, it is called a crop. Most crops are cultivated in agriculture or hydropon ...
s. Grasses are also used in the manufacture of thatch
Thatching is the craft of building a roof with dry vegetation such as straw, water reed, sedge (''Cladium mariscus''), rushes, heather, or palm branches, layering the vegetation so as to shed water away from the inner roof. Since the bulk of ...
, paper
Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distribu ...
, fuel
A fuel is any material that can be made to react with other substances so that it releases energy as thermal energy or to be used for work. The concept was originally applied solely to those materials capable of releasing chemical energy b ...
, clothing
Clothing (also known as clothes, apparel, and attire) are items worn on the body. Typically, clothing is made of fabrics or textiles, but over time it has included garments made from animal skin and other thin sheets of materials and natura ...
, insulation, timber for fencing
Fencing is a group of three related combat sports. The three disciplines in modern fencing are the foil, the épée, and the sabre (also ''saber''); winning points are made through the weapon's contact with an opponent. A fourth discipline, ...
, furniture
Furniture refers to movable objects intended to support various human activities such as seating (e.g., stools, chairs, and sofas), eating ( tables), storing items, eating and/or working with an item, and sleeping (e.g., beds and hammocks) ...
, scaffolding
Scaffolding, also called scaffold or staging, is a temporary structure used to support a work crew and materials to aid in the construction, maintenance and repair of buildings, bridges and all other man-made structures. Scaffolds are widely use ...
and construction
Construction is a general term meaning the art and science to form objects, systems, or organizations,"Construction" def. 1.a. 1.b. and 1.c. ''Oxford English Dictionary'' Second Edition on CD-ROM (v. 4.0) Oxford University Press 2009 and ...
materials, floor matting, sports turf
Poaceae () or Gramineae () is a large and nearly ubiquitous family of monocotyledonous flowering plants commonly known as grasses. It includes the cereal grasses, bamboos and the grasses of natural grassland and species cultivated in lawns and ...
and baskets
A basket is a container that is traditionally constructed from stiff fibers and can be made from a range of materials, including wood splints, runners, and cane. While most baskets are made from plant materials, other materials such as horsehai ...
.
Food production
Of all crops grown, 70% are grasses. Agricultural grasses grown for their edible seeds are called ''cereal
A cereal is any grass cultivated for the edible components of its grain (botanically, a type of fruit called a caryopsis), composed of the endosperm, germ, and bran. Cereal grain crops are grown in greater quantities and provide more food ...
s'' or ''grain
A grain is a small, hard, dry fruit ( caryopsis) – with or without an attached hull layer – harvested for human or animal consumption. A grain crop is a grain-producing plant. The two main types of commercial grain crops are cereals and legum ...
s'' (although the latter term, when used agriculturally, refers to both cereals and similar seeds of other plant species, such as buckwheat
Buckwheat (''Fagopyrum esculentum''), or common buckwheat, is a flowering plant in the knotweed family Polygonaceae cultivated for its grain-like seeds and as a cover crop. The name "buckwheat" is used for several other species, such as ''Fagop ...
and legumes
A legume () is a plant in the family Fabaceae (or Leguminosae), or the fruit or seed of such a plant. When used as a dry grain, the seed is also called a pulse. Legumes are grown agriculturally, primarily for human consumption, for livestock for ...
). Three cereals—rice, wheat, and maize (corn)—provide more than half of all calories consumed by humans. Cereals constitute the major source of carbohydrate
In organic chemistry, a carbohydrate () is a biomolecule consisting of carbon (C), hydrogen (H) and oxygen (O) atoms, usually with a hydrogen–oxygen atom ratio of 2:1 (as in water) and thus with the empirical formula (where ''m'' may o ...
s for humans and perhaps the major source of protein; these include rice (in southern
Southern may refer to:
Businesses
* China Southern Airlines, airline based in Guangzhou, China
* Southern Airways, defunct US airline
* Southern Air, air cargo transportation company based in Norwalk, Connecticut, US
* Southern Airways Express, M ...
and eastern Asia
East Asia is the eastern region of Asia, which is defined in both geographical and ethno-cultural terms. The modern states of East Asia include China, Japan, Mongolia, North Korea, South Korea, and Taiwan. China, North Korea, South Korea ...
), maize (in Central
Central is an adjective usually referring to being in the center of some place or (mathematical) object.
Central may also refer to:
Directions and generalised locations
* Central Africa, a region in the centre of Africa continent, also known a ...
and South America
South America is a continent entirely in the Western Hemisphere and mostly in the Southern Hemisphere, with a relatively small portion in the Northern Hemisphere at the northern tip of the continent. It can also be described as the sou ...
), and wheat and barley (in Europe
Europe is a large peninsula conventionally considered a continent in its own right because of its great physical size and the weight of its history and traditions. Europe is also considered a Continent#Subcontinents, subcontinent of Eurasia ...
, northern Asia
North Asia or Northern Asia, also referred to as Siberia, is the northern region of Asia, which is defined in geographical terms and is coextensive with the Asian part of Russia, and consists of three Russian regions east of the Ural Mountain ...
and the Americas
The Americas, which are sometimes collectively called America, are a landmass comprising the totality of North America, North and South America. The Americas make up most of the land in Earth's Western Hemisphere and comprise the New World. ...
).
Sugarcane
Sugarcane or sugar cane is a species of (often hybrid) tall, perennial grass (in the genus '' Saccharum'', tribe Andropogoneae) that is used for sugar production. The plants are 2–6 m (6–20 ft) tall with stout, jointed, fibrous stalk ...
is the major source of sugar
Sugar is the generic name for sweet-tasting, soluble carbohydrates, many of which are used in food. Simple sugars, also called monosaccharides, include glucose, fructose, and galactose. Compound sugars, also called disaccharides or do ...
production. Additional food uses of sugarcane include sprouted grain, shoot
In botany, a plant shoot consists of any plant stem together with its appendages, leaves and lateral buds, flowering stems, and flower buds. The new growth from seed germination that grows upward is a shoot where leaves will develop. In the sp ...
s, and rhizome
In botany and dendrology, a rhizome (; , ) is a modified subterranean plant stem that sends out roots and shoots from its nodes. Rhizomes are also called creeping rootstalks or just rootstalks. Rhizomes develop from axillary buds and grow ...
s, and in drink they include sugarcane juice and plant milk
Plant milk is a plant beverage with a color resembling that of milk. Plant milks are non-dairy beverages made from a water-based plant extract for flavoring and aroma. Plant milks are consumed as alternatives to milk, and often provide a crea ...
, as well as rum
Rum is a liquor made by fermenting and then distilling sugarcane molasses or sugarcane juice. The distillate, a clear liquid, is usually aged in oak barrels. Rum is produced in nearly every sugar-producing region of the world, such as the Ph ...
, beer
Beer is one of the oldest and the most widely consumed type of alcoholic drink in the world, and the third most popular drink overall after water and tea. It is produced by the brewing and fermentation of starches, mainly derived from ce ...
, whisky
Whisky or whiskey is a type of distilled alcoholic beverage made from fermented grain mash. Various grains (which may be malted) are used for different varieties, including barley, corn, rye, and wheat. Whisky is typically aged in wooden ca ...
, and vodka
Vodka ( pl, wódka , russian: водка , sv, vodka ) is a clear distilled alcoholic beverage. Different varieties originated in Poland, Russia, and Sweden. Vodka is composed mainly of water and ethanol but sometimes with traces of impuriti ...
.
Bamboo shoots are used in numerous Asian dishes and broths, and are available in supermarkets in various sliced forms, in both fresh, fermented and canned versions.
Lemongrass
''Cymbopogon'', also known as lemongrass, barbed wire grass, silky heads, Cochin grass, Malabar grass, oily heads, citronella grass or fever grass, is a genus of Asian, African, Australian, and tropical island plants in the grass family.
Some ...
is a grass used as a culinary herb for its citrus-like flavor and scent.
Many species of grass are grown as pasture
Pasture (from the Latin ''pastus'', past participle of ''pascere'', "to feed") is land used for grazing. Pasture lands in the narrow sense are enclosed tracts of farmland, grazed by domesticated livestock, such as horses, cattle, sheep, or sw ...
for foraging or as fodder
Fodder (), also called provender (), is any agricultural foodstuff used specifically to feed domesticated livestock, such as cattle, rabbits, sheep, horses, chickens and pigs. "Fodder" refers particularly to food given to the animals (includ ...
for prescribed livestock
Livestock are the domesticated animals raised in an agricultural setting to provide labor and produce diversified products for consumption such as meat, eggs, milk, fur, leather, and wool. The term is sometimes used to refer solely to ani ...
feeds, particularly in the case of cattle
Cattle (''Bos taurus'') are large, domesticated, cloven-hooved, herbivores. They are a prominent modern member of the subfamily Bovinae and the most widespread species of the genus '' Bos''. Adult females are referred to as cows and adult ...
, horse
The horse (''Equus ferus caballus'') is a domesticated, one-toed, hoofed mammal. It belongs to the taxonomic family Equidae and is one of two extant subspecies of ''Equus ferus''. The horse has evolved over the past 45 to 55 million yea ...
s, and sheep
Sheep or domestic sheep (''Ovis aries'') are domesticated, ruminant mammals typically kept as livestock. Although the term ''sheep'' can apply to other species in the genus '' Ovis'', in everyday usage it almost always refers to domesticate ...
. Such grasses may be cut and stored for later feeding, especially for the winter, in the form of bales of hay or straw
Straw is an agricultural byproduct consisting of the dry stalks of cereal plants after the grain and chaff have been removed. It makes up about half of the yield of cereal crops such as barley, oats, rice, rye and wheat. It has a number ...
, or in silos as silage
Silage () is a type of fodder made from green foliage crops which have been preserved by fermentation to the point of acidification. It can be fed to cattle, sheep and other such ruminants (cud-chewing animals). The fermentation and storage ...
. Straw (and sometimes hay) may also be used as bedding for animals.
An example of a sod-forming perennial grass used in agriculture is ''Thinopyrum intermedium
''Thinopyrum intermedium'', known commonly as intermediate wheatgrass, is a sod-forming perennial grass in the Triticeae tribe of Pooideae native to Europe and Western Asia. It is part of a group of plants commonly called wheatgrasses because of ...
''.
Industry
Grasses are used as raw material for a multitude of purposes, including construction and in the composition of building materials such as cob, for insulation, in the manufacture of paper and board such as oriented structural straw board Oriented structural straw board (OSSB) is an engineered board that is made by splitting straw and formed by adding formaldehyde-free adhesives and then hot compressing layers of straw in specific orientations. Research and development for OSSB panel ...
. Grass fiber
Fiber or fibre (from la, fibra, links=no) is a natural or artificial substance that is significantly longer than it is wide. Fibers are often used in the manufacture of other materials. The strongest engineering materials often incorporate ...
can be used for making paper
Paper is a thin sheet material produced by mechanically or chemically processing cellulose fibres derived from wood, rags, grasses or other vegetable sources in water, draining the water through fine mesh leaving the fibre evenly distribu ...
, biofuel
Biofuel is a fuel that is produced over a short time span from biomass, rather than by the very slow natural processes involved in the formation of fossil fuels, such as oil. According to the United States Energy Information Administration ...
production, nonwoven fabrics, and as replacement for glass fibers used in reinforced plastics. Bamboo scaffolding is able to withstand typhoon-force winds that would break steel scaffolding. Larger bamboos and ''Arundo donax
''Arundo donax'' is a tall perennial cane. It is one of several so-called reed species. It has several common names including giant cane, elephant grass, carrizo, arundo, Spanish cane, Colorado river reed, wild cane, and giant reed. ''Arundo'' a ...
'' have stout culms that can be used in a manner similar to timber, ''Arundo'' is used to make reeds for woodwind instrument
Woodwind instruments are a family of musical instruments within the greater category of wind instruments. Common examples include flute, clarinet, oboe, bassoon, and saxophone. There are two main types of woodwind instruments: flutes and r ...
s, and bamboo is used for innumerable implements.
'' ''Phragmites australis'''' (common reed) is important for thatching
Thatching is the craft of building a roof with dry vegetation such as straw, water reed, sedge (''Cladium mariscus''), rushes, heather, or palm branches, layering the vegetation so as to shed water away from the inner roof. Since the bulk of ...
and wall construction of homes in Africa.
Grasses are used in water treatment
Water treatment is any process that improves the quality of water to make it appropriate for a specific end-use. The end use may be drinking, industrial water supply, irrigation, river flow maintenance, water recreation or many other uses, inc ...
systems, in wetland conservation
Wetland conservation is aimed at protecting and preserving areas where water exists at or near the Earth's surface, such as swamps, marshes and bogs. Wetlands cover at least six per cent of the Earth and have become a focal issue for conservation ...
and land reclamation
Land reclamation, usually known as reclamation, and also known as land fill (not to be confused with a waste landfill), is the process of creating new land from oceans, seas, riverbeds or lake beds. The land reclaimed is known as reclamat ...
, and used to lessen the erosional impact of urban storm water runoff.
Lawn and ornamental use
Grasses are the primary plant used in lawns, which themselves derive from grazed grassland
A grassland is an area where the vegetation is dominated by grasses ( Poaceae). However, sedge ( Cyperaceae) and rush ( Juncaceae) can also be found along with variable proportions of legumes, like clover, and other herbs. Grasslands occur na ...
s in Europe. They also provide an important means of erosion control (e.g., along roadsides), especially on sloping land. Grass lawns are an important covering of playing surfaces in many sports, including football (soccer)
Association football, more commonly known as football or soccer, is a team sport played between two teams of 11 players who primarily use their feet to propel the ball around a rectangular field called a pitch. The objective of the game is ...
, American football
American football (referred to simply as football in the United States and Canada), also known as gridiron, is a team sport played by two teams of eleven players on a rectangular field with goalposts at each end. The offense, the team wi ...
, tennis
Tennis is a racket sport that is played either individually against a single opponent (singles) or between two teams of two players each (doubles). Each player uses a tennis racket that is strung with cord to strike a hollow rubber ball cov ...
, golf
Golf is a club-and-ball sport in which players use various clubs to hit balls into a series of holes on a course in as few strokes as possible.
Golf, unlike most ball games, cannot and does not use a standardized playing area, and coping wi ...
, cricket
Cricket is a bat-and-ball game played between two teams of eleven players on a field at the centre of which is a pitch with a wicket at each end, each comprising two bails balanced on three stumps. The batting side scores runs by st ...
, softball
Softball is a game similar to baseball played with a larger ball on a smaller field. Softball is played competitively at club levels, the college level, and the professional level. The game was first created in 1887 in Chicago by George Hanc ...
and baseball
Baseball is a bat-and-ball sport played between two teams of nine players each, taking turns batting and fielding. The game occurs over the course of several plays, with each play generally beginning when a player on the fielding t ...
.
Ornamental grasses, such as perennial
A perennial plant or simply perennial is a plant that lives more than two years. The term ('' per-'' + '' -ennial'', "through the years") is often used to differentiate a plant from shorter-lived annuals and biennials. The term is also widel ...
bunch grass
Tussock grasses or bunch grasses are a group of grass species in the family Poaceae. They usually grow as singular plants in clumps, tufts, hummocks, or bunches, rather than forming a sod or lawn, in meadows, grasslands, and prairies. As perennia ...
es, are used in many styles of garden design for their foliage, inflorescences, seed heads. They are often used in natural landscaping
Natural landscaping, also called native gardening, is the use of native plants and adapted species, including trees, shrubs, groundcover, and grasses which are local to the geographic area of the garden.
Benefits
Maintenance
Natural land ...
, xeriscaping
Xeriscaping is the process of Garden design, landscaping, or gardening, that reduces or eliminates the need for irrigation. It is promoted in regions that do not have accessible, plentiful, or reliable supplies of fresh water and has gained accep ...
and slope and beach stabilization in contemporary landscaping, wildlife garden
A wildlife garden (or wild garden) is an environment created by a gardener that serves as a sustainable haven for surrounding wildlife. Wildlife gardens contain a variety of habitats that cater to native and local plants, birds, amphibians, ...
ing, and native plant gardening
Natural landscaping, also called native gardening, is the use of native plants and adapted species, including trees, shrubs, groundcover, and grasses which are local to the geographic area of the garden.
Benefits
Maintenance
Natural landsca ...
. They are used as screens and hedges.
Sports turf
Grass playing fields, courses and pitches are the traditional playing surfaces for many sports
Sport pertains to any form of competitive physical activity or game that aims to use, maintain, or improve physical ability and skills while providing enjoyment to participants and, in some cases, entertainment to spectators. Sports can, ...
, including American football
American football (referred to simply as football in the United States and Canada), also known as gridiron, is a team sport played by two teams of eleven players on a rectangular field with goalposts at each end. The offense, the team wi ...
, association football
Association football, more commonly known as football or soccer, is a team sport played between two teams of 11 players who primarily use their feet to propel the ball around a rectangular field called a pitch. The objective of the game is ...
, baseball
Baseball is a bat-and-ball sport played between two teams of nine players each, taking turns batting and fielding. The game occurs over the course of several plays, with each play generally beginning when a player on the fielding t ...
, cricket
Cricket is a bat-and-ball game played between two teams of eleven players on a field at the centre of which is a pitch with a wicket at each end, each comprising two bails balanced on three stumps. The batting side scores runs by st ...
, golf
Golf is a club-and-ball sport in which players use various clubs to hit balls into a series of holes on a course in as few strokes as possible.
Golf, unlike most ball games, cannot and does not use a standardized playing area, and coping wi ...
, and rugby
Rugby may refer to:
Sport
* Rugby football in many forms:
** Rugby league: 13 players per side
*** Masters Rugby League
*** Mod league
*** Rugby league nines
*** Rugby league sevens
*** Touch (sport)
*** Wheelchair rugby league
** Rugby union: 1 ...
. Grass surfaces are also sometimes used for horse racing
Horse racing is an equestrian performance sport, typically involving two or more horses ridden by jockeys (or sometimes driven without riders) over a set distance for competition. It is one of the most ancient of all sports, as its basic pr ...
and tennis
Tennis is a racket sport that is played either individually against a single opponent (singles) or between two teams of two players each (doubles). Each player uses a tennis racket that is strung with cord to strike a hollow rubber ball cov ...
. Type of maintenance and species of grass used may be important factors for some sports, less critical for others. In some sports facilities, including indoor domes and other places where maintenance of a grass field would be difficult, grass may be replaced with artificial turf
Artificial turf is a surface of synthetic fibers made to look like natural grass. It is most often used in arenas for sports that were originally or are normally played on grass. However, it is now being used on residential lawns and commerc ...
, a synthetic grass-like substitute.
Cricket
In cricket, the pitch is the strip of carefully mowed and rolled grass where the bowler bowls. In the days leading up to the match it is repeatedly mowed and rolled to produce a very hard, flat surface for the ball to bounce off.
Golf
Grass on golf courses is kept in three distinct conditions: that of the ''rough'', the ''fairway'', and the ''putting green''. Grass on the fairway is mown short and even, allowing the player to strike the ball cleanly. Playing from the rough is a disadvantage because the long grass may affect the flight of the ball. Grass on the putting green is the shortest and most even, ideally allowing the ball to roll smoothly over the surface. An entire industry revolves around the development and marketing of turf grass varieties.
Tennis
In tennis, grass is grown on very hard-packed soil, and the bounce of a tennis ball may vary depending on the grass's health, how recently it has been mowed, and the wear and tear of recent play. The surface is softer than hard courts
A hardcourt (or hard court) is a surface or floor on which a sport is played, most usually in reference to tennis courts. It is typically made of rigid materials such as asphalt or concrete, and covered with acrylic resins to seal the surface an ...
and clay
Clay is a type of fine-grained natural soil material containing clay minerals (hydrous aluminium phyllosilicates, e.g. kaolin, Al2 Si2 O5( OH)4).
Clays develop plasticity when wet, due to a molecular film of water surrounding the clay pa ...
(other tennis surfaces), so the ball bounces lower, and players must reach the ball faster resulting in a different style of play which may suit some players more than others. Among the world's most prestigious court for grass tennis is Centre Court at Wimbledon, London
Wimbledon () is a district and town of Southwest London, England, southwest of the centre of London at Charing Cross; it is the main commercial centre of the London Borough of Merton. Wimbledon had a population of 68,187 in 2011 which includes ...
which hosts the final of the annual Wimbledon Championships
The Wimbledon Championships, commonly known simply as Wimbledon, is the oldest tennis tournament in the world and is widely regarded as the most prestigious. It has been held at the All England Club in Wimbledon, London, since 1877 and is pla ...
in England, one of the four Grand Slam tournaments.
Economically important grasses
A number of grasses are invasive species that damage natural ecosystems, including forms of ''Phragmites australis'' which are native to Eurasia but has spread around the world.
Role in society
Grasses have long had significance in human society. They have been cultivated as feed for people and domesticated animals for thousands of years. The primary ingredient of beer
Beer is one of the oldest and the most widely consumed type of alcoholic drink in the world, and the third most popular drink overall after water and tea. It is produced by the brewing and fermentation of starches, mainly derived from ce ...
is usually barley or wheat, both of which have been used for this purpose for over 4,000 years.
In some places, particularly in suburb
A suburb (more broadly suburban area) is an area within a metropolitan area, which may include commercial and mixed-use, that is primarily a residential area. A suburb can exist either as part of a larger city/urban area or as a separ ...
an areas, the maintenance of a grass lawn is a sign of a homeowner's responsibility to the overall appearance of their neighborhood. One work credits lawn maintenance to:
In communities with drought
A drought is defined as drier than normal conditions.Douville, H., K. Raghavan, J. Renwick, R.P. Allan, P.A. Arias, M. Barlow, R. Cerezo-Mota, A. Cherchi, T.Y. Gan, J. Gergis, D. Jiang, A. Khan, W. Pokam Mba, D. Rosenfeld, J. Tierney, an ...
problems, watering of lawns may be restricted to certain times of day or days of the week. Many US municipalities and homeowners' associations have rules which require lawns to be maintained to certain specifications, sanctioning those who allow the grass to grow too long.
The smell of the freshly cut grass is produced mainly by cis-3-Hexenal.
Some common aphorism
An aphorism (from Greek ἀφορισμός: ''aphorismos'', denoting 'delimitation', 'distinction', and 'definition') is a concise, terse, laconic, or memorable expression of a general truth or principle. Aphorisms are often handed down by ...
s involve grass. For example:
* "The grass is always greener on the other side" suggests an alternate state of affairs will always seem preferable to one's own.
* "Don't let the grass grow under your feet" tells someone to get moving.
* "A snake
Snakes are elongated, limbless, carnivorous reptiles of the suborder Serpentes . Like all other squamates, snakes are ectothermic, amniote vertebrates covered in overlapping scales. Many species of snakes have skulls with several more ...
in the grass" means dangers that are hidden.
* "When elephant
Elephants are the largest existing land animals. Three living species are currently recognised: the African bush elephant, the African forest elephant, and the Asian elephant. They are the only surviving members of the family Elephantida ...
s fight, it is the grass which suffers" tells of bystanders caught in the crossfire.
A folk myth about grass is that it refuses to grow where any violent death has occurred.[Olmert, Michael (1996). ''Milton's Teeth and Ovid's Umbrella: Curiouser & Curiouser Adventures in History'', p. 208. Simon & Schuster, New York. .]
Image gallery
File:Ruwbeemdgras Poa trivialis ligula.jpg, Leaves of '' Poa trivialis'' showing the ligules A ligule (from "strap", variant of ''lingula'', from ''lingua'' "tongue") is a thin outgrowth at the junction of leaf and leafstalk of many grasses (Poaceae) and sedges. A ligule is also a strap-shaped extension of the corolla, such as that of a r ...
File:Bamboo DSCN2465.jpg, Bamboo stem and leaves, nodes are evident
File:Chasmanthium latifolium-spikelet.jpg, A ''Chasmanthium latifolium
''Chasmanthium latifolium'', known as northern wood-oats, inland sea oats, northern sea oats, and river oats is a species of grass native to the central and eastern United States, Manitoba, and northeastern Mexico; it grows as far north as Pennsy ...
'' spikelet
File:En Spica spiculae.png, Wheat spike and spikelet
File:En Aperta.png, Spikelet opened to show caryopsis
File:Harestail grass.jpg, Harestail grass
File:Grass.jpg, Grass
File:Saccharum-officinarum2.JPG, Sugarcane (''Saccharum officinarum
''Saccharum officinarum'' is a large, strong-growing species of grass in the genus '' Saccharum''. Its stout stalks are rich in sucrose, a simple sugar which accumulates in the stalk internodes. It originated in New Guinea, and is now cultivated ...
'')
File:Bromus hordeaceus unten.jpeg, Roots of ''Bromus hordeaceus
''Bromus hordeaceus'', the soft brome, is an annual or biennial species of grass in the true grass family ( Poaceae). It is also known in North America as bull grass, soft cheat, and soft chess.
It is the most common species of ''Bromus'' in Bri ...
''
File:Ohra.jpg, Barley mature spikes ('' Hordeum vulgare'')
File:Koeh-283.jpg, Illustration depicting both staminate and pistillate flowers of maize (''Zea mays
Maize ( ; ''Zea mays'' subsp. ''mays'', from es, maíz after tnq, mahiz), also known as corn (North American and Australian English), is a cereal grain first domesticated by indigenous peoples in southern Mexico about 10,000 years ago. The ...
'')
File:Flowering Grass.JPG, A grass flower head (meadow foxtail) showing the plain-coloured flowers with large anthers.
File:Grass Anthers.JPG, Anthers detached from a meadow foxtail flower
File:Setaria verticillata W IMG 1084.jpg, ''Setaria verticillata
''Setaria verticillata'' is a species of grass known by the common names hooked bristlegrass, rough bristle-grass and bristly foxtail. It is native to Europe, but it is known on most continents as an introduced species and often a noxious weed. I ...
'', bristly foxtail
File:Setaria verticillata W IMG 1083.jpg, ''Setaria verticillata
''Setaria verticillata'' is a species of grass known by the common names hooked bristlegrass, rough bristle-grass and bristly foxtail. It is native to Europe, but it is known on most continents as an introduced species and often a noxious weed. I ...
'', bristly foxtail
File:Oryza sativa in Kadavoor.jpg, '' Oryza sativa'', Kerala
Kerala ( ; ) is a state on the Malabar Coast of India. It was formed on 1 November 1956, following the passage of the States Reorganisation Act, by combining Malayalam-speaking regions of the erstwhile regions of Cochin, Malabar, South Ca ...
, India
India, officially the Republic of India (Hindi: ), is a country in South Asia. It is the List of countries and dependencies by area, seventh-largest country by area, the List of countries and dependencies by population, second-most populous ...
See also
* Agrostology
Agrostology (from Greek , ''agrōstis'', "type of grass"; and , ''-logia''), sometimes graminology, is the scientific study of the grasses (the family Poaceae, or Gramineae). The grasslike species of the sedge family (Cyperaceae), the rush family ...
* Forb
A forb or phorb is an herbaceous flowering plant that is not a graminoid (grass, sedge, or rush). The term is used in biology and in vegetation ecology, especially in relation to grasslands and understory. Typically these are dicots without woo ...
*GrassBase
GrassBase (or GrassBase – The Online World Grass Flora) is a web-based database of grasses, continually maintained and updated by the Royal Botanic Gardens, Kew.
As of 2015, GrassBase was one of the largest (along with GrassWorld) structured ...
* PACMAD clade
* Thinopyrum intermedium
''Thinopyrum intermedium'', known commonly as intermediate wheatgrass, is a sod-forming perennial grass in the Triticeae tribe of Pooideae native to Europe and Western Asia. It is part of a group of plants commonly called wheatgrasses because of ...
References
External links
*
Need a Definition of Grass?
Vegetative Key to Grasses
Poaceae
a
''The Plant List''
a
''The Families of Flowering Plants (DELTA)''
at th
''Angiosperm Phylogeny Website''
''Poaceae Classification''
from the onlin
''Catalogue of New World Grasses''
Poaceae
at the onlin
''Guide to the Flora of Mongolia''
Poaceae
at the onlin
''Flora of Taiwan''
Poaceae
at the onlin
''Flora of Pakistan''
Poaceae
at the onlin
''Flora of Zimbabwe''
Poaceae
at the onlin
''Flora of Western Australia''
* Grasses of Australia (AusGrass2) – http://ausgrass2.myspecies.info/
Gramineae
at the onlin
''Flora of New Zealand''
NZ Grass Key
An Interactive Key to New Zealand Grasses a
''Landcare Research''
The Grass Genera of the World
a
''DELTA intkey''
''GrassWorld''
{{Authority control
Poales families
Grasslands
Plant life-forms
Plants by habit
Extant Albian first appearances