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The Galápagos Islands (
Spanish Spanish might refer to: * Items from or related to Spain: **Spaniards are a nation and ethnic group indigenous to Spain **Spanish language, spoken in Spain and many Latin American countries **Spanish cuisine Other places * Spanish, Ontario, Can ...
: , , ) are an
archipelago An archipelago ( ), sometimes called an island group or island chain, is a chain, cluster, or collection of islands, or sometimes a sea containing a small number of scattered islands. Examples of archipelagos include: the Indonesian Arc ...
of volcanic islands. They are distributed on each side of the equator in the
Pacific Ocean The Pacific Ocean is the largest and deepest of Earth's five oceanic divisions. It extends from the Arctic Ocean in the north to the Southern Ocean (or, depending on definition, to Antarctica) in the south, and is bounded by the contin ...
, surrounding the centre of the
Western Hemisphere The Western Hemisphere is the half of the planet Earth that lies west of the prime meridian (which crosses Greenwich, London, United Kingdom) and east of the antimeridian. The other half is called the Eastern Hemisphere. Politically, the te ...
, and are part of the Republic of Ecuador. Located west of continental Ecuador, the islands are known for their large number of endemic species that were studied by
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
during the second voyage of HMS ''Beagle''. His observations and collections contributed to the
inception of Darwin's theory The inception of Darwin's theory occurred during an intensively busy period which began when Charles Darwin returned from the survey voyage of the ''Beagle'', with his reputation as a fossil collector and geologist already established. He was gi ...
of
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
by means of
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Cha ...
. The Galápagos Islands and their surrounding waters form the
Galápagos Province Galápagos () is a province of Ecuador in the country's Insular region, located approximately off the western coast of the mainland. The capital is Puerto Baquerizo Moreno. The province administers the Galápagos Islands, a group of tiny vol ...
of Ecuador, the
Galápagos National Park Galápagos National Park ( es, Parque Nacional Galápagos) was established in 1959. It began operation in 1968, and it is Ecuador's first national park and a UNESCO World Heritage Site. Park history The government of Ecuador has designated ...
, and the
Galápagos Marine Reserve The Galápagos Marine Reserve (GMR) lies a thousand kilometres from the Ecuadorian mainland and covers an area of around . The Galápagos Islands and the surrounding waters represent one of the world’s most unusual ecosystems and are rich areas ...
. The principal language on the islands is Spanish. The islands have a population of slightly over 25,000. The first recorded visit to the islands happened by chance in 1535, when
Fray Tomás de Berlanga Fray or Frays or The Fray may refer to: Arts, entertainment, and media Fictional entities *Fray, a phenomenon in Terry Pratchett's '' The Carpet People'' *Fray, the main character in the video games: **''Fray in Magical Adventure'' **''Fray CD' ...
, the Bishop of Panamá, was surprised to find this undiscovered land on a voyage to
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
to arbitrate in a dispute between
Francisco Pizarro Francisco Pizarro González, Marquess of the Atabillos (; ;  – 26 June 1541) was a Spanish conquistador, best known for his expeditions that led to the Spanish conquest of Peru. Born in Trujillo, Spain to a poor family, Pizarro chose ...
and
Diego de Almagro Diego de Almagro (; – July 8, 1538), also known as El Adelantado and El Viejo, was a Spanish conquistador known for his exploits in western South America. He participated with Francisco Pizarro in the Spanish conquest of Peru. While subd ...
. Berlanga eventually returned to the Spanish Empire and described the conditions of the islands and the animals that inhabited them. The group of islands was shown and named "Insulae de los Galopegos" (Islands of the tortoises) in Abraham Ortelius's map "America Sive Novi Orbis" published in 1570. The first crude
map A map is a symbolic depiction emphasizing relationships between elements of some space, such as objects, regions, or themes. Many maps are static, fixed to paper or some other durable medium, while others are dynamic or interactive. Although ...
of the islands was made in 1684 by the buccaneer Ambrose Cowley, who named the individual islands after some of his fellow pirates or after English royalty and noblemen. These names were used in the authoritative navigation charts of the islands prepared during the ''Beagle'' survey under captain
Robert FitzRoy Vice-Admiral Robert FitzRoy (5 July 1805 – 30 April 1865) was an English officer of the Royal Navy and a scientist. He achieved lasting fame as the captain of during Charles Darwin's famous voyage, FitzRoy's second expedition to Tierra de ...
, and in Darwin's popular book ''
The Voyage of the Beagle ''The Voyage of the Beagle'' is the title most commonly given to the book written by Charles Darwin and published in 1839 as his ''Journal and Remarks'', bringing him considerable fame and respect. This was the third volume of ''The Narrative ...
''. The newly independent Republic of Ecuador took the islands from Spanish ownership in 1832, and subsequently gave them new Spanish names. The older names remained in use in English-language publications, including
Herman Melville Herman Melville ( born Melvill; August 1, 1819 – September 28, 1891) was an American novelist, short story writer, and poet of the American Renaissance period. Among his best-known works are ''Moby-Dick'' (1851); ''Typee'' (1846), a rom ...
's '' The Encantadas'' of 1854. Administratively, Galapagos constitutes one of the provinces of Ecuador, made up of three cantons that bear the names of its most populated islands, namely: San Cristóbal, Santa Cruz and Isabela.


Geology

Volcanism Volcanism, vulcanism or volcanicity is the phenomenon of eruption of molten rock (magma) onto the surface of the Earth or a solid-surface planet or moon, where lava, pyroclastics, and volcanic gases erupt through a break in the surface called a ...
has been continuous on the Galápagos Islands for at least 20 million years, and perhaps even longer. The
mantle plume A mantle plume is a proposed mechanism of convection within the Earth's mantle, hypothesized to explain anomalous volcanism. Because the plume head partially melts on reaching shallow depths, a plume is often invoked as the cause of volcanic hot ...
beneath the east-ward moving Nazca Plate (51 km/ myr) has given rise to a 3-kilometre-thick platform under the island chain and seamounts. Besides the Galápagos
Archipelago An archipelago ( ), sometimes called an island group or island chain, is a chain, cluster, or collection of islands, or sometimes a sea containing a small number of scattered islands. Examples of archipelagos include: the Indonesian Arc ...
, other key tectonic features in the region include the Northern Galápagos Volcanic Province between the archipelago and the Galápagos
Spreading Center A mid-ocean ridge (MOR) is a seafloor mountain system formed by plate tectonics. It typically has a depth of about and rises about above the deepest portion of an ocean basin. This feature is where seafloor spreading takes place along a diver ...
(GSC) 200 km to the north at the boundary of the Nazca Plate and the
Cocos Plate The Cocos Plate is a young oceanic tectonic plate beneath the Pacific Ocean off the west coast of Central America, named for Cocos Island, which rides upon it. The Cocos Plate was created approximately 23 million years ago when the Farallon Plate ...
. This spreading center truncates into the
East Pacific Rise The East Pacific Rise is a mid-ocean rise (termed an oceanic rise and not a mid-ocean ridge due to its higher rate of spreading that results in less elevation increase and more regular terrain), a divergent tectonic plate boundary located alon ...
on the west and is bounded by the Cocos Ridge and Carnegie Ridge in the east. Furthermore, the Galápagos Hotspot is at the northern boundary of the Pacific Large Low Shear Velocity Province while the Easter Hotspot is on the southern boundary. The Galápagos Archipelago is characterized by numerous contemporaneous volcanoes, some with plume magma sources, others from the asthenosphere, possibly due to the young and thin
oceanic crust Oceanic crust is the uppermost layer of the oceanic portion of the tectonic plates. It is composed of the upper oceanic crust, with pillow lavas and a dike complex, and the lower oceanic crust, composed of troctolite, gabbro and ultramafic ...
. The GSC caused structural weaknesses in this thin lithosphere leading to eruptions forming the Galápagos Platform. Fernandina and Isabela in particular are aligned along these weaknesses. Lacking a well-defined
rift zone A rift zone is a feature of some volcanoes, especially shield volcanoes, in which a set of linear cracks (or rifts) develops in a volcanic edifice, typically forming into two or three well-defined regions along the flanks of the vent. Believed t ...
, the islands have a high rate of
inflation In economics, inflation is an increase in the general price level of goods and services in an economy. When the general price level rises, each unit of currency buys fewer goods and services; consequently, inflation corresponds to a reduct ...
prior to eruption. Sierra Negra on Isabela Island experienced a 240 cm uplift between 1992 and 1998, most recent eruption in 2005, while Fernandina on Fernandina Island indicated an uplift of 90 cm, most recent eruption in 2009. Alcedo on Isabela Island had an uplift of greater than 90 cm, most recent eruption in 1993. Additional characteristics of the Galápagos Archipelago are closer volcano spacing, smaller volcano sizes, and larger
caldera A caldera ( ) is a large cauldron-like hollow that forms shortly after the emptying of a magma chamber in a volcano eruption. When large volumes of magma are erupted over a short time, structural support for the rock above the magma chamber is ...
s. For instance, Isabela Island includes 6 major volcanoes, Ecuador, Wolf, Darwin, Alcedo, Sierra Negraa and Cerro Azul, with most recent eruptions ranging from 1813 to 2008. The neighboring islands of Santiago and Fernandina last erupted in 1906 and 2009, respectively. Overall, the 9 active volcanoes in the archipelago have erupted 24 times between 1961 and 2011. The shape of these volcanoes is tall and rounded as opposed wide and smooth in the Hawaiian Islands. The Galápagos's shape is due to the pattern of radial and circumferential
fissure A fissure is a long, narrow crack opening along the surface of Earth. The term is derived from the Latin word , which means 'cleft' or 'crack'. Fissures emerge in Earth's crust, on ice sheets and glaciers, and on volcanoes. Ground fissure ...
, radial on the flanks, but circumferential near the caldera summits. It is the circumferential fissures which give rise to stacks of short lava flows. The volcanoes at the west end of the archipelago are in general, taller, younger, have well developed calderas, and are mostly composed of
tholeiitic basalt The tholeiitic magma series is one of two main magma series in subalkaline igneous rocks, the other being the calc-alkaline series. A magma series is a chemically distinct range of magma compositions that describes the evolution of a mafic magma ...
, while those on the east are shorter, older, lack calderas, and have a more diverse composition. The ages of the islands, from west to east are 0.05 Ma for Fernandina, 0.65 Ma for Isabela, 1.10 Ma for Santiago, 1.7 Ma for Santa Cruz, 2.90 Ma for Santa Fe, and 3.2 Ma for San Cristobal. The calderas on Sierra Negra and Alcedo have active fault systems. The Sierra Negra fault is associated with a sill 2 km below the caldera. The caldera on Fernandina experienced the largest basaltic volcano collapse in history, with the 1968
phreatomagmatic eruption Phreatomagmatic eruptions are volcanic eruptions resulting from interaction between magma and water. They differ from exclusively magmatic eruptions and phreatic eruptions. Unlike phreatic eruptions, the products of phreatomagmatic eruptions cont ...
. Fernandina has also been the most active volcano since 1790, with recent eruptions in 1991, 1995, 2005, and 2009, and the entire surface has been covered in numerous flows since 4.3 Ka. The western volcanoes have numerous
tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock ...
cones A cone is a three-dimensional geometric shape that tapers smoothly from a flat base (frequently, though not necessarily, circular) to a point called the apex or vertex. A cone is formed by a set of line segments, half-lines, or lines conn ...
.


Physical geography

The islands are located in the eastern Pacific Ocean, off the west coast of South America. The majority of islands are also more broadly part of the South Pacific. The closest land mass is that of mainland
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
, the country to which they belong, to the east. The islands are found at the coordinates 1°40'N–1°36'S, 89°16'–92°01'W. Straddling the equator, islands in the chain are located in both the northern and southern hemispheres, with Volcán Wolf and Volcán Ecuador on Isla Isabela being directly on the equator.
Española Island Española Island (Spanish: ''Isla Española'') is part of the Galápagos Islands. The English named it ''Hood Island'' after Viscount Samuel Hood. It is located in the extreme southeast of the archipelago and is considered, along with Sant ...
, the southernmost
islet An islet is a very small, often unnamed island. Most definitions are not precise, but some suggest that an islet has little or no vegetation and cannot support human habitation. It may be made of rock, sand and/or hard coral; may be permanen ...
of the archipelago, and
Darwin Island Darwin Island (Spanish: ''Isla Darwin'') is among the smallest in the Galápagos Archipelago with an area of just . It is named in honour of English scientist Charles Darwin. With no dry landing sites, Darwin Island's main attractions are fo ...
, the northernmost one, are spread out over a distance of . The
International Hydrographic Organization The International Hydrographic Organization (IHO) is an intergovernmental organisation representing hydrography. , the IHO comprised 98 Member States. A principal aim of the IHO is to ensure that the world's seas, oceans and navigable waters a ...
(IHO) considers them wholly within the South Pacific Ocean, however. The Galápagos Archipelago consists of of land spread over of ocean. The largest of the islands, Isabela, measures and makes up close to three-quarters of the total land area of the Galápagos. Volcán Wolf on Isabela is the highest point, with an elevation of above sea level. The group consists of 18 main islands, 3 smaller islands, and 107 rocks and
islet An islet is a very small, often unnamed island. Most definitions are not precise, but some suggest that an islet has little or no vegetation and cannot support human habitation. It may be made of rock, sand and/or hard coral; may be permanen ...
s. The islands are located at the Galapagos Triple Junction. The archipelago is located on the Nazca Plate (a tectonic plate), which is moving east/southeast, diving under the South American Plate at a rate of about per year. It is also atop the
Galápagos hotspot The Galápagos hotspot is a volcanic hotspot in the East Pacific Ocean responsible for the creation of the Galápagos Islands as well as three major aseismic ridge systems, Carnegie, Cocos and Malpelo which are on two tectonic plates. The hots ...
, a place where the Earth's crust is being melted from below by a
mantle plume A mantle plume is a proposed mechanism of convection within the Earth's mantle, hypothesized to explain anomalous volcanism. Because the plume head partially melts on reaching shallow depths, a plume is often invoked as the cause of volcanic hot ...
, creating volcanoes. The first islands formed here at least 8 million and possibly up to 90 million years ago. While the older islands have disappeared below the sea as they moved away from the mantle plume, the youngest islands, Isabela and Fernandina, are still being formed. In April 2009, lava from the volcanic island Fernandina started flowing both towards the island's shoreline and into the center caldera. In late June 2018, Sierra Negra, one of five volcanoes on Isabela and one of the most active in the Galapagos archipelago, began erupting for the first time since 2005. Lava flows made their way to the coastline, prompting the evacuation of about fifty nearby residents and restricting tourist access.


Main islands

The 18 main islands (each having a land area at least 1 km2) of the archipelago (with their English names) shown alphabetically: * Baltra (South Seymour) Island – Baltra is a small flat island located near the centre of the Galápagos. It was created by geological uplift. The island is very arid, and vegetation consists of salt bushes, prickly pear cacti and palo santo trees. Until 1986, Baltra (Seymour) Airport was the only airport serving the Galápagos. Now, there are two airports which receive flights from the continent; the other is located on San Cristóbal Island. Private planes flying to Galápagos must fly to Baltra, as it is the only airport with facilities for planes overnight. On arriving in Baltra, all visitors are immediately transported by bus to one of two docks. The first dock is located in a small bay, where the boats cruising Galápagos await passengers. The second is a ferry dock, which connects Baltra to the island of Santa Cruz. During the 1940s, scientists decided to move 70 of Baltra's land iguanas to the neighboring North Seymour Island as part of an experiment. This move proved unexpectedly useful when the native iguanas became extinct on Baltra as a result of the island's military occupation in
World War II World War II or the Second World War, often abbreviated as WWII or WW2, was a world war that lasted from 1939 to 1945. It involved the vast majority of the world's countries—including all of the great powers—forming two opposing ...
. During the 1980s, iguanas from North Seymour were brought to the
Charles Darwin Research Station Charles is a masculine given name predominantly found in English and French speaking countries. It is from the French form ''Charles'' of the Proto-Germanic name (in runic alphabet) or ''*karilaz'' (in Latin alphabet), whose meaning was ...
as part of a breeding and repopulation project, and in the 1990s, land iguanas were reintroduced to Baltra. As of 1997, scientists counted 97 iguanas living on Baltra; 13 of which had hatched on the islands. * Bartolomé (Bartholomew) Island – Bartolomé Island is a volcanic islet just off the east coast of Santiago Island in the Galápagos Islands group. it is one of the younger islands in the Galápagos archipelago. This island, and neighbouring Sulivan Bay on Santiago (James) island, are named after lifelong friend of Charles Darwin, Sir Bartholomew James Sulivan, who was a lieutenant aboard HMS ''Beagle''. Today Sulivan Bay is often misspelled Sullivan Bay. This island is one of the few that are home to the
Galápagos penguin The Galápagos penguin (''Spheniscus mendiculus'') is a penguin endemic to the Galápagos Islands, Ecuador. It is the only penguin found north of the equator. Most inhabit Fernandina Island and the west coast of Isabela Island. The cool wate ...
which is the only wild penguin species to live on the equator. The
green turtle The green sea turtle (''Chelonia mydas''), also known as the green turtle, black (sea) turtle or Pacific green turtle, is a species of large sea turtle of the family Cheloniidae. It is the only species in the genus ''Chelonia''. Its range exten ...
is another animal that resides on the island. * Darwin (Culpepper) Island – This island is named after Charles Darwin. It has an area of and a maximum altitude of . Here fur seals, frigates, marine iguanas, swallow-tailed gulls, sea lions, whales, marine turtles, and red-footed and Nazca boobies can be seen. Darwin's Arch, a natural rock arch which would at one time have been part of this larger structure, is located less than a kilometre from the main Darwin Island, and it is a landmark well known to the island's few visitors. It collapsed in May 2021. * Española (Hood) Island – Its name was given in honor of Spain. It also is known as ''Hood'', after Viscount Samuel Hood. It has an area of and a maximum altitude of . Española is the oldest island at around 3.5 million years, and the southernmost in the group. Due to its remote location, Española has a large number of endemic species. It has its own species of lava lizard, mockingbird, and
Galápagos tortoise The Galápagos tortoise or Galápagos giant tortoise (''Chelonoidis niger'') is a species of very large tortoise in the genus ''Chelonoidis'' (which also contains three smaller species from mainland South America). It comprises 15 subspecies ( ...
. Española's marine iguanas exhibit a distinctive red coloration change between the breeding season. Española is the only place where the waved albatross nests. Some of the birds have attempted to breed on Genovesa (Tower) Island, but unsuccessfully. Española's steep cliffs serve as the perfect runways for these birds, which take off for their ocean feeding grounds near the mainland of Ecuador and Peru. Española has two visitor sites. Gardner Bay is a swimming and snorkelling site, and offers a great beach. Punta Suarez has migrant, resident, and endemic wildlife, including brightly colored
marine iguana The marine iguana (''Amblyrhynchus cristatus''), also known as the sea iguana, saltwater iguana, or Galápagos marine iguana, is a species of iguana found only on the Galápagos Islands (Ecuador). Unique among modern lizards, it is a marine rept ...
s, Española lava lizards, hood mockingbirds,
swallow-tailed gull The swallow-tailed gull (''Creagrus furcatus'') is an equatorial seabird in the gull family, Laridae. It is the only species in the genus ''Creagrus'', which derives from the Latin ''Creagra'' and the Greek ''kreourgos'' which means butcher, al ...
s, blue-footed boobies, Nazca boobies,
red-billed tropicbird The red-billed tropicbird (''Phaethon aethereus'') is a tropicbird, one of three closely related species of seabird of tropical oceans. Superficially resembling a tern in appearance, it has mostly white plumage with some black markings on the wi ...
s, Galápagos hawks, three species of Darwin's finches, and the waved albatross. * Fernandina (Narborough) Island – The name was given in honor of King Ferdinand II of Aragon, who sponsored the voyage of Columbus. Fernandina has an area of and a maximum altitude of . This is the youngest and westernmost island. On 13 May 2005, a new, very eruptive process began on this island, when an ash and water vapor cloud rose to a height of and lava flows descended the slopes of the volcano on the way to the sea. Punta Espinosa is a narrow stretch of land where hundreds of marine iguanas gather, largely on black lava rocks. The famous
flightless cormorant The flightless cormorant (''Nannopterum harrisi''), also known as the Galapagos cormorant, is a cormorant endemic to the Galapagos Islands, and an example of the highly unusual fauna there. It is unique in that it is the only known cormorant th ...
s inhabit this island, as do
Galápagos penguin The Galápagos penguin (''Spheniscus mendiculus'') is a penguin endemic to the Galápagos Islands, Ecuador. It is the only penguin found north of the equator. Most inhabit Fernandina Island and the west coast of Isabela Island. The cool wate ...
s, pelicans,
Galápagos sea lion The Galápagos sea lion (''Zalophus wollebaeki'') is a species of sea lion that lives and breeds on the Galápagos Islands and, in smaller numbers, on Isla de la Plata (Ecuador). Being fairly social, they are often spotted sun-bathing on sandy ...
s and Galápagos fur seals. Different types of
lava Lava is molten or partially molten rock (magma) that has been expelled from the interior of a terrestrial planet (such as Earth) or a moon onto its surface. Lava may be erupted at a volcano or through a fracture in the crust, on land or un ...
flows can be compared, and the
mangrove A mangrove is a shrub or tree that grows in coastal saline or brackish water. The term is also used for tropical coastal vegetation consisting of such species. Mangroves are taxonomically diverse, as a result of convergent evolution in severa ...
forests can be observed. * Floreana (Charles or Santa María) Island – It was named after
Juan José Flores Juan José Flores y Aramburu (19 July 1800 – 1 October 1864) was a Venezuelan-born military general who became the first (in 1830), third (in 1839) and fourth (in 1843) President of the new Republic of Ecuador. He is often referred to as "The ...
, the first
President of Ecuador The president of Ecuador ( es, Presidente del Ecuador), officially called the Constitutional President of the Republic of Ecuador ( es, Presidente Constitucional de la República del Ecuador), serves as both the head of state and head of govern ...
, during whose administration the government of Ecuador took possession of the archipelago. It is also called Santa Maria, after one of the caravels of Columbus. It has an area of and a maximum elevation of . It is one of the islands with the most interesting human history, and one of the earliest to be inhabited.
Flamingo Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbea ...
s and green sea turtles nest (December to May) on this island. The ''patapegada'' or Galápagos petrel, a sea bird which spends most of its life away from land, is found here. At Post Office Bay, where 19th-century whalers kept a
wooden barrel A barrel or cask is a hollow cylindrical container with a bulging center, longer than it is wide. They are traditionally made of wooden staves and bound by wooden or metal hoops. The word vat is often used for large containers for liquids, u ...
that served as a post office, mail could be picked up and delivered to its destinations, mainly Europe and the United States, by ships on their way home. At the "Devil's Crown", an underwater
volcanic cone Volcanic cones are among the simplest volcanic landforms. They are built by ejecta from a volcanic vent, piling up around the vent in the shape of a cone with a central crater. Volcanic cones are of different types, depending upon the nature and ...
and
coral Corals are marine invertebrates within the class Anthozoa of the phylum Cnidaria. They typically form compact colonies of many identical individual polyps. Coral species include the important reef builders that inhabit tropical oceans and ...
formations are found. * Genovesa (Tower) Island – The name is derived from
Genoa Genoa ( ; it, Genova ; lij, Zêna ). is the capital of the Italian region of Liguria and the sixth-largest city in Italy. In 2015, 594,733 people lived within the city's administrative limits. As of the 2011 Italian census, the Province of ...
, Italy, the birthplace of Christopher Columbus. It has an area of and a maximum altitude of . This island is formed by the remaining edge of a large
caldera A caldera ( ) is a large cauldron-like hollow that forms shortly after the emptying of a magma chamber in a volcano eruption. When large volumes of magma are erupted over a short time, structural support for the rock above the magma chamber is ...
that is submerged. Its nickname of "the bird island" is clearly justified. At Darwin Bay,
frigatebird Frigatebirds are a family of seabirds called Fregatidae which are found across all tropical and subtropical oceans. The five extant species are classified in a single genus, ''Fregata''. All have predominantly black plumage, long, deeply forked ...
s and
swallow-tailed gull The swallow-tailed gull (''Creagrus furcatus'') is an equatorial seabird in the gull family, Laridae. It is the only species in the genus ''Creagrus'', which derives from the Latin ''Creagra'' and the Greek ''kreourgos'' which means butcher, al ...
s, the only nocturnal species of gull in the world, can be seen. Red-footed
boobies A booby is a seabird in the genus ''Sula'', part of the family Sulidae. Boobies are closely related to the gannets (''Morus''), which were formerly included in ''Sula''. Systematics and evolution The genus ''Sula'' was introduced by the Fren ...
, noddy
tern Terns are seabirds in the family Laridae that have a worldwide distribution and are normally found near the sea, rivers, or wetlands. Terns are treated as a subgroup of the family Laridae which includes gulls and skimmers and consists of e ...
s, lava gulls, tropic birds,
dove Columbidae () is a bird family consisting of doves and pigeons. It is the only family in the order Columbiformes. These are stout-bodied birds with short necks and short slender bills that in some species feature fleshy ceres. They primarily ...
s,
storm petrel Storm-petrel may refer to one of two bird families, both in the order Procellariiformes, once treated as the same family. The two families are: * Northern storm petrels (''Hydrobatidae'') are found in the Northern Hemisphere, although some speci ...
s and Darwin finches are also in sight. Prince Philip's Steps is a bird-watching
plateau In geology and physical geography, a plateau (; ; ), also called a high plain or a tableland, is an area of a highland consisting of flat terrain that is raised sharply above the surrounding area on at least one side. Often one or more sides ...
with Nazca and red-footed boobies. There is a large Palo Santo forest. * Isabela (Albemarle) Island – This island was named in honor of Queen Isabella I of Castile. With an area of , it is the largest island of the Galápagos. Its highest point is Volcán Wolf, with an altitude of . The island's
seahorse A seahorse (also written ''sea-horse'' and ''sea horse'') is any of 46 species of small marine fish in the genus ''Hippocampus''. "Hippocampus" comes from the Ancient Greek (), itself from () meaning "horse" and () meaning "sea monster" or " ...
shape is the product of the merging of six large volcanoes into a single land mass. On this island, Galápagos penguins, flightless cormorants, marine iguanas, pelicans and Sally Lightfoot crabs abound. At the skirts and calderas of the volcanoes of Isabela, land iguanas and Galápagos tortoises can be observed, as well as Darwin finches, Galápagos hawks, Galápagos doves and very interesting lowland vegetation. The third-largest human settlement of the archipelago,
Puerto Villamil Puerto Villamil is a small port village located on the southeastern edge of Isla Isabela Island (Ecuador), Isabela in the Galapagos Islands. Of the 2,200 people who live on Isabela, the majority live in Puerto Villamil. The harbor is frequentl ...
, is located at the southeastern tip of the island. * Marchena (Bindloe) Island – Named after Fray Antonio Marchena, it has an area of and a maximum altitude of .
Galapagos hawk The Galápagos hawk (''Buteo galapagoensis'') is a large hawk endemic to most of the Galápagos Islands. Description The Galapágos hawk is similar in size to the red-tailed hawk (''Buteo jamaicensis'') and the Swainson's hawk (''Buteo swain ...
s and sea lions inhabit this island, and it is home to the Marchena lava lizard, an endemic animal. * North Seymour Island – Its name was given after an English nobleman,
Lord Hugh Seymour Vice-Admiral Lord Hugh Seymour (29 April 1759 – 11 September 1801) was a senior British Royal Navy officer of the late 18th century who was the fifth son of Francis Seymour-Conway, 1st Marquess of Hertford, and became known for being both a ...
. It has an area of and a maximum altitude of . This island is home to a large population of blue-footed boobies and swallow-tailed gulls. It hosts one of the largest populations of frigate birds. It was formed from geological uplift. * Pinzón (Duncan) Island – Named after the Pinzón brothers, captains of the Pinta and Niña caravels, it has an area of and a maximum altitude of . * Pinta (Louis) Island – Named after the Pinta caravel, it has an area of and a maximum altitude of . Sea lions, Galápagos hawks, giant tortoises, marine iguanas, and dolphins can be seen here. Pinta Island was home to the last remaining Pinta tortoise, called Lonesome George. He was moved from Pinta Island to the
Charles Darwin Research Station Charles is a masculine given name predominantly found in English and French speaking countries. It is from the French form ''Charles'' of the Proto-Germanic name (in runic alphabet) or ''*karilaz'' (in Latin alphabet), whose meaning was ...
on Santa Cruz Island, where scientists attempted to breed from him. However, Lonesome George died in June 2012 without producing any offspring. * Rábida (Jervis) Island – It bears the name of the convent of Rábida, where Columbus left his son during his voyage to the Americas. It has an area of and a maximum altitude of . The high amount of iron contained in the lava at Rábida gives it a distinctive red colour. White-cheeked pintail ducks live in a saltwater lagoon close to the beach, where brown pelicans and boobies have built their nests. Until recently, flamingos were also found in the lagoon, but they have since moved on to other islands, likely due to a lack of food on Rábida. Nine species of finches have been reported in this island. * San Cristóbal (Chatham) Island – It bears the name of the patron saint of seafarers, "
St. Christopher Saint Christopher ( el, Ἅγιος Χριστόφορος, ''Ágios Christóphoros'') is venerated by several Christian denominations as a martyr killed in the reign of the 3rd-century Roman emperor Decius (reigned 249–251) or alternatively u ...
". Its English name was given after
William Pitt, 1st Earl of Chatham William Pitt, 1st Earl of Chatham, (15 November 170811 May 1778) was a British statesman of the Whig group who served as Prime Minister of Great Britain from 1766 to 1768. Historians call him Chatham or William Pitt the Elder to distinguish ...
. It has an area of and its highest point rises to . This is the first island in the Galápagos Archipelago Charles Darwin visited during his voyage on the ''Beagle''. This islands hosts frigate birds, sea lions, giant tortoises, blue- and red-footed
boobies A booby is a seabird in the genus ''Sula'', part of the family Sulidae. Boobies are closely related to the gannets (''Morus''), which were formerly included in ''Sula''. Systematics and evolution The genus ''Sula'' was introduced by the Fren ...
, tropicbirds,
marine iguana The marine iguana (''Amblyrhynchus cristatus''), also known as the sea iguana, saltwater iguana, or Galápagos marine iguana, is a species of iguana found only on the Galápagos Islands (Ecuador). Unique among modern lizards, it is a marine rept ...
s, dolphins and
swallow-tailed gull The swallow-tailed gull (''Creagrus furcatus'') is an equatorial seabird in the gull family, Laridae. It is the only species in the genus ''Creagrus'', which derives from the Latin ''Creagra'' and the Greek ''kreourgos'' which means butcher, al ...
s. Its vegetation includes ''Calandrinia galapagos'', ''Lecocarpus darwinii'', and trees such as ''Lignum vitae''. The largest freshwater lake in the archipelago, Laguna El Junco, is located in the highlands of San Cristóbal. The capital of the province of Galápagos is Puerto Baquerizo Moreno, which lies at the southern tip of the island, and is close to San Cristóbal Airport. * Santa Cruz (Indefatigable) Island – Given the name of the Holy Cross in Spanish. It was originally named Norfolk Island by Cowley, but renamed after the British frigate HMS ''Indefatigable'' after her visit there in 1812. It has an area of and a maximum altitude of . Santa Cruz hosts the largest human population in the archipelago, the town of Puerto Ayora. The
Charles Darwin Research Station Charles is a masculine given name predominantly found in English and French speaking countries. It is from the French form ''Charles'' of the Proto-Germanic name (in runic alphabet) or ''*karilaz'' (in Latin alphabet), whose meaning was ...
and the headquarters of the Galápagos National Park Service are located here. The GNPS and CDRS operate a tortoise breeding centre here, where young tortoises are hatched, reared, and prepared to be reintroduced to their natural
habitat In ecology, the term habitat summarises the array of resources, physical and biotic factors that are present in an area, such as to support the survival and reproduction of a particular species. A species habitat can be seen as the physical ...
. The Highlands of Santa Cruz offer exuberant flora, and are famous for the lava tunnels. Large tortoise populations are found here. Black Turtle Cove is a site surrounded by mangroves, which sea turtles, rays and small sharks sometimes use as a mating area. Cerro Dragón, known for its flamingo lagoon, is also located here, and along the trail one may see land iguanas foraging. * Santa Fe (Barrington) Island – Named after a city in Spain, it has an area of and a maximum altitude of . Santa Fe hosts a forest of Opuntia
cactus A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Gree ...
, which are the largest of the archipelago, and Palo Santo. Weathered cliffs provide a haven for swallow-tailed gulls, red-billed tropic birds and
shearwater Shearwaters are medium-sized long-winged seabirds in the petrel family Procellariidae. They have a global marine distribution, but are most common in temperate and cold waters, and are pelagic outside the breeding season. Description These tub ...
petrels. Santa Fe species of land iguanas are often seen, as well as lava lizards. * Santiago (San Salvador, James) Island – Its name is equivalent to Saint James in English; it is also known as San Salvador, after the first island discovered by Columbus in the Caribbean Sea. This island has an area of and a maximum altitude of . Marine iguanas, sea lions, fur seals, land and sea turtles,
flamingo Flamingos or flamingoes are a type of wading bird in the family Phoenicopteridae, which is the only extant family in the order Phoenicopteriformes. There are four flamingo species distributed throughout the Americas (including the Caribbea ...
s, dolphins and sharks are found here. Pigs and goats, which were introduced by humans to the islands and have caused great harm to the endemic species, have been eradicated (pigs by 2002; goats by the end of 2006). Darwin finches and Galápagos hawks are usually seen, as well as a colony of fur seals. At Sulivan Bay, a recent (around 100 years ago) pahoehoe
lava Lava is molten or partially molten rock (magma) that has been expelled from the interior of a terrestrial planet (such as Earth) or a moon onto its surface. Lava may be erupted at a volcano or through a fracture in the crust, on land or un ...
flow can be observed. * Wolf (Wenman) Island – This island was named after the German geologist Theodor Wolf. It has an area of and a maximum altitude of . Here,
fur seal Fur seals are any of nine species of pinnipeds belonging to the subfamily Arctocephalinae in the family '' Otariidae''. They are much more closely related to sea lions than true seals, and share with them external ears (pinnae), relatively l ...
s, frigatebirds, Nazca and red-footed boobies, marine iguanas, sharks, whales, dolphins and swallow-tailed gulls can be seen. The most famous resident is the vampire finch, which feeds partly on blood pecked from other birds, and is only found on this island.


Minor islands

*
Daphne Major Daphne Major is a volcanic island just north of Santa Cruz Island and just west of the Baltra Airport in the Archipelago of Colón, commonly known as the Galápagos Islands. It consists of a tuff crater, devoid of trees, whose rim rises above the ...
– A small island directly north of Santa Cruz and directly west of Baltra, this very inaccessible island appears, though unnamed, on Ambrose Cowley's 1684 chart. It is important as the location of multidecade finch population studies by Peter and Rosemary Grant. *
South Plaza Island South Plaza (Spanish language, Spanish: ''Isla Plaza Sur'') is a small island off the east coast of Santa Cruz Island (Galápagos), Santa Cruz in the Galápagos Islands. It has an area of 0.13 km2 and a maximum altitude of 23 metres. Sou ...
(Plaza Sur) – It is named in honor of a former president of Ecuador, General Leónidas Plaza. It has an area of and a maximum altitude of . The flora of South Plaza includes '' Opuntia'' cactus and '' Sesuvium'' plants, which form a reddish carpet on top of the lava formations. Iguanas (land, marine and some hybrids of both species) are abundant, and large numbers of birds can be observed from the cliffs at the southern part of the island, including tropic birds and swallow-tailed gulls. *
Nameless Island Nameless Island (Spanish: ''Isla Sin Nombre'') is an islet of the Galápagos Islands group, in Ecuador, between Pinzón Island and Santa Cruz Island (Galápagos). The island is most commonly used for scuba diving Scuba diving is a mode of ...
– A small islet used mostly for
scuba diving Scuba diving is a mode of underwater diving whereby divers use breathing equipment that is completely independent of a surface air supply. The name "scuba", an acronym for " Self-Contained Underwater Breathing Apparatus", was coined by Chr ...
. *
Roca Redonda Roca Redonda is a flat-topped, steep-sided islet located roughly northwest of the island of Isabela, in the Galápagos Islands of Ecuador. It measures long and wide with a maximum elevation of . Its isolation and inaccessibility coupled with it ...
– An islet approximately northwest of Isabela. Herman Melville devotes the third and fourth sketches of The Encantadas to describing this islet (which he calls "Rock Rodondo") and the view from it.


Climate

Although the islands are located on the equator, the Humboldt Current brings cold water to them, causing frequent drizzles during most of the year. The weather is periodically influenced by the
El Niño El Niño (; ; ) is the warm phase of the El Niño–Southern Oscillation (ENSO) and is associated with a band of warm ocean water that develops in the central and east-central equatorial Pacific (approximately between the International Date ...
events, which occur about every 3 to 7 years and are characterized by warm sea surface temperatures, a rise in sea level, greater wave action, and a depletion of nutrients in the water. During the season known as the ''garúa'' (June to November), the temperature by the sea is , a steady and cold wind blows from south and southeast, frequent drizzles (''garúas'') last most of the day, and dense fog conceals the islands. During the warm season (December to May), the average sea and air temperature rises to , there is no wind at all, there are sporadic, though strong, rains and the sun shines. Weather changes as altitude increases in the large islands. Temperature decreases gradually with altitude, while precipitation increases due to the condensation of moisture in clouds on the slopes. There is a large range in precipitation from one place to another, not only with altitude, but also depending on the location of the islands, and also with the seasons. The archipelago is mainly characterized by a mixture of a
tropical savanna climate Tropical savanna climate or tropical wet and dry climate is a tropical climate sub-type that corresponds to the Köppen climate classification categories ''Aw'' (for a dry winter) and ''As'' (for a dry summer). The driest month has less than of p ...
and a
semi-arid climate A semi-arid climate, semi-desert climate, or steppe climate is a dry climate sub-type. It is located on regions that receive precipitation below potential evapotranspiration, but not as low as a desert climate. There are different kinds of sem ...
. It also transits into a
tropical rainforest climate A tropical rainforest climate, humid tropical climate or equatorial climate is a tropical climate sub-type usually found within 10 to 15 degrees latitude of the equator. There are some other areas at higher latitudes, such as the coast of southea ...
in the northwest. The following table corresponding to the wet 1969 shows the variation of precipitation in different places of
Santa Cruz Island Santa Cruz Island ( Spanish: ''Isla Santa Cruz'', Chumash: ''Limuw'') is located off the southwestern coast of Ventura, California, United States. It is the largest island in California and largest of the eight islands in the Channel Islands ...
: The precipitation also depends on the geographical location. During March 1969, the precipitation over Charles Darwin Station, on the southern coast of Santa Cruz was , while on Baltra Island, the precipitation during the same month was only . This is because Baltra is located behind Santa Cruz with respect to the prevailing southerly winds, so most of the moisture gets precipitated in the Santa Cruz highlands. There are significant changes in precipitation from one year to another, too. At Charles Darwin Station, the precipitation during March 1969 was , but during March 1970, it was only . On the larger islands, the pattern of generally wet highlands and drier lowlands affects the flora. The vegetation in the highlands tends to be green and lush, with tropical woodland in places. The lowland areas tend to have arid and semi-arid vegetation, with many thorny shrubs and cacti, and almost bare volcanic rock elsewhere.


Ecology


Terrestrial

Most of the Galápagos is covered in semi-desert vegetation, including shrublands, grasslands, and dry forest. A few of the islands have high-elevation areas with cooler temperatures and higher rainfall, which are home to humid-climate forests and shrublands, and montane grasslands (''pampas'') at the highest elevations. There are about 500 species of native
vascular plant Vascular plants (), also called tracheophytes () or collectively Tracheophyta (), form a large group of land plants ( accepted known species) that have lignified tissues (the xylem) for conducting water and minerals throughout the plant. They ...
s on the islands, including 90 species of ferns. About 180 vascular plant species are endemic. The islands are well known for their distinctive
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
species, including
giant tortoises Giant tortoises are any of various large land tortoises Giant tortoise or giant tortoises may also refer to: * Galápagos giant tortoise, ''Chelonoidis nigra'' are a complex of the largest living species of tortoise. :: Pinta giant tortoise or ...
,
finches The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
,
flightless cormorant The flightless cormorant (''Nannopterum harrisi''), also known as the Galapagos cormorant, is a cormorant endemic to the Galapagos Islands, and an example of the highly unusual fauna there. It is unique in that it is the only known cormorant th ...
s, Galápagos lava lizards and
marine iguana The marine iguana (''Amblyrhynchus cristatus''), also known as the sea iguana, saltwater iguana, or Galápagos marine iguana, is a species of iguana found only on the Galápagos Islands (Ecuador). Unique among modern lizards, it is a marine rept ...
s, which evolved to adapt to islands' environments.


History


Pre-Columbian era

Whether the Incas ever made it to the islands is disputed.
Oceanic Oceanic may refer to: *Of or relating to the ocean *Of or relating to Oceania **Oceanic climate **Oceanic languages **Oceanic person or people, also called "Pacific Islander(s)" Places * Oceanic, British Columbia, a settlement on Smith Island, ...
Pacific islands in the same general area as Galápagos (including
Clipperton Island Clipperton Island ( or ; ) is an uninhabited, coral atoll in the eastern Pacific Ocean. It is from Paris, France, from Papeete, Tahiti, and from Mexico. It is an overseas state private property of France under direct authority of the Minis ...
,
Cocos Island Cocos Island ( es, Isla del Coco) is an island in the Pacific Ocean administered by Costa Rica, approximately southwest of the Costa Rican mainland. It constitutes the 11th of the 13 districts of Puntarenas Canton of the Province of Puntarena ...
,
Desventuradas Islands The Desventuradas Islands ( es, Islas Desventuradas, , "Unfortunate Islands" or ''Islas de los Desventurados'', "Islands of the Unfortunate Ones") is a group of four small oceanic islands located off the coast of Chile, northwest of Santiago in ...
,
Juan Fernández Islands The Juan Fernández Islands ( es, Archipiélago Juan Fernández) are a sparsely inhabited series of islands in the South Pacific Ocean reliant on tourism and fishing. Situated off the coast of Chile, they are composed of three main volcanic i ...
and the
Revillagigedo Islands The Revillagigedo Islands ( es, Islas Revillagigedo, ) or Revillagigedo Archipelago are a group of four volcanic islands in the Pacific Ocean, known for their unique ecosystem. They lie approximately from Socorro Island south and southwest of C ...
) were all uninhabited when discovered by Europeans, with nothing to indicate any prehistoric human activity. The easternmost oceanic island in the South Pacific that was discovered with a human population was
Easter Island Easter Island ( rap, Rapa Nui; es, Isla de Pascua) is an island and special territory of Chile in the southeastern Pacific Ocean, at the southeasternmost point of the Polynesian Triangle in Oceania. The island is most famous for its ne ...
. Through DNA studies and linguistics, its population, the
Rapa Nui people The Rapa Nui (Rapa Nui: , Spanish: ) are the Polynesian peoples indigenous to Easter Island. The easternmost Polynesian culture, the descendants of the original people of Easter Island make up about 60% of the current Easter Island population a ...
, are known to be Polynesian rather than Indigenous American. In 1572, Spanish chronicler
Pedro Sarmiento de Gamboa Pedro Sarmiento de Gamboa (1532–1592) was a Spanish explorer, author, historian, mathematician, and astronomer. His birthplace is not certain and may have been Pontevedra, in Galicia, where his paternal family originated, or Alcalá de Henare ...
claimed that
Topa Inca Yupanqui Topa Inca Yupanqui or Túpac Inca Yupanqui ( qu, 'Tupaq Inka Yupanki'), translated as "noble Inca accountant," (c. 1441–c. 1493) was the tenth Sapa Inca (1471–93) of the Inca Empire, fifth of the Hanan dynasty. His father was Pachacuti, and h ...
, the second Sapa Inca of the
Inca Empire The Inca Empire (also known as the Incan Empire and the Inka Empire), called ''Tawantinsuyu'' by its subjects, ( Quechua for the "Realm of the Four Parts",  "four parts together" ) was the largest empire in pre-Columbian America. The adm ...
had visited the archipelago, but there is little evidence for this, and many experts consider it a far-fetched legend, especially since the Incas were not seafaring people. According to a 1952 archaeological survey by Thor Heyerdahl and Arne Skjølsvold, potsherds and other artifacts from several sites on the islands suggest visitation by South American peoples in
pre-Columbian era In the history of the Americas, the pre-Columbian era spans from the original settlement of North and South America in the Upper Paleolithic period through European colonization, which began with Christopher Columbus's voyage of 1492. Usually, ...
.Heyerdahl, Thor; & Skjolsvold, Arne (1956). "Archaeological Evidence of Pre-Spanish Visits to the Galápagos Islands", Memoirs 12, Society for American Archaeology. The group located an
Inca The Inca Empire (also known as the Incan Empire and the Inka Empire), called ''Tawantinsuyu'' by its subjects, (Quechua for the "Realm of the Four Parts",  "four parts together" ) was the largest empire in pre-Columbian America. The admin ...
flute and shards from more than 130 pieces of ceramics, which were later identified as pre-Incan. However, no remains of graves, ceremonial vessels or constructions have ever been found, suggesting no permanent settlement occurred before the Spanish arrived in the 16th century. A 2016 reanalysis of Heyerdahl and Skjølsvold's archaeological sites rejected their conclusions. They found that at all locations, artifacts of Indian and European origin were interspersed without the distinct spatial or stratigraphic arrangement that would be expected from independent sequential deposition (indeed, Heyerdahl and Skjølsvold had reported the intermixing of European and American artifacts in their original report). Radiocarbon dates from the sites placed them in the historical (post-Spanish-arrival) era, and preliminary paleoenvironmental analysis showed no disturbance older than 500 years before present, suggesting the islands were probably not visited prior to their Spanish discovery in 1535. The authors suggested that native artifacts found by Heyerdahl and Skjølsvold had probably been brought as mementos or souvenirs at the time of Spanish occupation. A 2008 report by archeologists from the
Australian National University The Australian National University (ANU) is a public research university located in Canberra, the capital of Australia. Its main campus in Acton encompasses seven teaching and research colleges, in addition to several national academies an ...
stated that certain Asia-Pacific taxa may have been growing in the Galápagos prior to 1535, which, "would constitute a strong line of evidence for accidental or deliberate landfall in the Galápagos by a Polynesian vessel." Their report also claimed there is a growing body of evidence to suggest that Pacific islands beyond Easter Island "did not play a ‘stepping stone’ role in the interaction between Amerindians or Polynesians in prehistory." Whatever their identity, the first visitors to the islands were likely unimpressed by the lack of fresh water on the islands.


European voyages

European discovery of the Galápagos Islands occurred when Spaniard
Fray Tomás de Berlanga Fray or Frays or The Fray may refer to: Arts, entertainment, and media Fictional entities *Fray, a phenomenon in Terry Pratchett's '' The Carpet People'' *Fray, the main character in the video games: **''Fray in Magical Adventure'' **''Fray CD' ...
, the fourth Bishop of Panama, sailed to
Peru , image_flag = Flag of Peru.svg , image_coat = Escudo nacional del Perú.svg , other_symbol = Great Seal of the State , other_symbol_type = National seal , national_motto = "Firm and Happy f ...
to settle a dispute between
Francisco Pizarro Francisco Pizarro González, Marquess of the Atabillos (; ;  – 26 June 1541) was a Spanish conquistador, best known for his expeditions that led to the Spanish conquest of Peru. Born in Trujillo, Spain to a poor family, Pizarro chose ...
and his lieutenants. Berlanga's vessel drifted off course when the winds diminished, and his party reached the islands by mere chance on 10 March 1535. They were uninhabited, and continued to remain unclaimed once the men left. The Galápagos Islands first appeared on the maps of
Gerardus Mercator Gerardus Mercator (; 5 March 1512 – 2 December 1594) was a 16th-century geographer, cosmographer and Cartography, cartographer from the County of Flanders. He is most renowned for creating the Mercator 1569 world map, 1569 world map based on ...
and Abraham Ortelius, in about 1570. The islands were named "Insulae de los Galopegos" (Islands of the Tortoises) in reference to the
giant tortoise Giant tortoises are any of several species of various large land tortoises, which include a number of extinct species, as well as two extant species with multiple subspecies formerly common on the islands of the western Indian Ocean and on the ...
s found there. The first English captain to visit the Galápagos Islands was
Richard Hawkins Admiral Sir Richard Hawkins (or Hawkyns) (c. 1562 – 17 April 1622) was a 17th-century English seaman, explorer and privateer. He was the son of Admiral Sir John Hawkins. Biography He was from his earlier days familiar with ships and the s ...
, in 1593. Until the early 19th century, the archipelago was often used as a hideout by mostly English pirates who attacked a
Spanish treasure fleet The Spanish treasure fleet, or West Indies Fleet ( es, Flota de Indias, also called silver fleet or plate fleet; from the es, label=Spanish, plata meaning "silver"), was a convoy system of sea routes organized by the Spanish Empire from 1566 to ...
carrying gold and silver from South America to Spain. In 1793, James Colnett described the flora and fauna of Galápagos, and suggested the islands could be used as base for the whalers operating in the Pacific Ocean. He drew the first accurate navigation charts of the islands. Whalers and maritime fur traders killed and captured thousands of the Galápagos tortoises to extract their fat. The tortoises could be kept on board ship as a means of providing of fresh protein, as these animals could survive for several months on board without any food or water. The hunting of the tortoises was responsible for greatly diminishing, and in some cases eliminating, certain species. Along with whalers came the fur-seal hunters, who brought the population of this animal close to extinction. The first known permanent human resident on Galápagos was Patrick Watkins, an Irish sailor who was marooned on the Island Floreana from 1807 to 1809. According to later accounts, Watkins managed to survive by hunting, growing vegetables and trading with visiting whalers, before finally stealing an open boat and navigating to
Guayaquil , motto = Por Guayaquil Independiente en, For Independent Guayaquil , image_map = , map_caption = , pushpin_map = Ecuador#South America , pushpin_re ...
. In 1818 the Nantucket whaleship ''Globe'', under Captain George Washington Gardner, discovered a "mother lode" of sperm whales some thousand miles west of the South American coast approximately at the equator. He returned to Nantucket in 1820 with more than 2,000 barrels of sperm whale oil and the news of his discovery. This led to an influx of whaleships to exploit the new whaling ground and the Galápagos Islands became a frequent stop for the whalers both before and after visiting what came to be known as the Offshore Grounds. This led to the establishment in the Galápagos Islands of a kind of unofficial "post office" where whaleships stopped to pick up and drop off letters as well as for provisioning and repairing. In October 1820, the whaleship ''Essex'', out of Nantucket, stopped at the Galápagos for these purposes on its way to the Offshore Grounds. On what was then known as Charles Island, while most of the crew were hunting tortoises one crewmember, English boatsteerer Thomas Chappel, for reasons still unclear, lit a fire which quickly burned out of control. Some of the tortoise hunters had a narrow escape and had to run a gauntlet of fire to get back to the ship. Soon almost the entire island was in flames. Crewmembers reported that after a day of sailing away they could still see the flames against the horizon. One crewmember who returned to the Galápagos several years afterward described the entire island as still a blackened wasteland.


Ecuadorian Galápagos


Scientific expeditions and first settlers

Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
annexed the Galápagos Islands on 12 February 1832, naming them the Archipelago of Ecuador. This new name added to several names that had been, and are still, used to refer to the archipelago. The first governor of Galápagos, General José de Villamil, brought a group of convicts to populate the island of Floreana, and in October 1832, some artisans and farmers joined them. The voyage of the ''Beagle'' brought the survey ship HMS ''Beagle'', under captain
Robert FitzRoy Vice-Admiral Robert FitzRoy (5 July 1805 – 30 April 1865) was an English officer of the Royal Navy and a scientist. He achieved lasting fame as the captain of during Charles Darwin's famous voyage, FitzRoy's second expedition to Tierra de ...
, to the Galápagos on 15 September 1835 to survey approaches to harbours. The captain and others on board, including his companion, the young naturalist
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
, made observations on the geology and biology on Chatham, Charles, Albemarle and James islands before they left on 20 October to continue on their round-the-world expedition. Primarily a geologist at the time, Darwin was impressed by the quantity of volcanic craters they saw, later referring to the archipelago as "that land of craters". His study of several volcanic formations over the five weeks he stayed in the islands led to several important geological discoveries, including the first, correct explanation for how volcanic
tuff Tuff is a type of rock made of volcanic ash ejected from a vent during a volcanic eruption. Following ejection and deposition, the ash is lithified into a solid rock. Rock that contains greater than 75% ash is considered tuff, while rock ...
is formed. Darwin noticed the
mockingbird Mockingbirds are a group of New World passerine birds from the family Mimidae. They are best known for the habit of some species mimicking the songs of other birds and the sounds of insects and amphibians, often loudly and in rapid succession. ...
s differed between islands, though he thought the birds now known as
Darwin's finches Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or t ...
were unrelated to each other, and did not bother labelling them by island. Nicholas Lawson, acting Governor of Galápagos for the Republic of Equator, met them on ''Charles Island'', and as they walked to the prison colony, Lawson told Darwin the tortoises differed from island to island. Towards the end of the voyage, Darwin speculated that the distribution of the mockingbirds and the tortoises might "undermine the stability of Species". When specimens of birds were analyzed on his return to England, it was found that many apparently different kinds of birds were species of
finch The true finches are small to medium-sized passerine birds in the family Fringillidae. Finches have stout conical bills adapted for eating seeds and nuts and often have colourful plumage. They occupy a great range of habitats where they are usua ...
es, which were unique to islands. These facts were crucial in Darwin's development of his theory of
natural selection Natural selection is the differential survival and reproduction of individuals due to differences in phenotype. It is a key mechanism of evolution, the change in the heritable traits characteristic of a population over generations. Cha ...
explaining
evolution Evolution is change in the heritable characteristics of biological populations over successive generations. These characteristics are the expressions of genes, which are passed on from parent to offspring during reproduction. Variation ...
, which was presented in ''
On the Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
''. In April 1888 , a Navy-crewed research vessel assigned to the
United States Fish Commission The United States Fish Commission, formally known as the United States Commission of Fish and Fisheries, was an agency of the United States government created in 1871 to investigate, promote, and preserve the fisheries of the United States. In 1 ...
, briefly touched eight islands in the Galapagos group for specimens; this included Wreck Bay on Chatham Island (now San Cristóbal Island) on 4 April and Charles Island (now Floreana Island) on 8 April.
José Valdizán José is a predominantly Spanish and Portuguese form of the given name Joseph. While spelled alike, this name is pronounced differently in each language: Spanish ; Portuguese (or ). In French, the name ''José'', pronounced , is an old vernacu ...
and
Manuel Julián Cobos Manuel may refer to: People * Manuel (name) * Manuel (Fawlty Towers), a fictional character from the sitcom ''Fawlty Towers'' * Charlie Manuel, manager of the Philadelphia Phillies * Manuel I Komnenos, emperor of the Byzantine Empire * Manu ...
tried a new colonization, beginning the exploitation of a type of lichen found in the islands (''Roccella portentosa'') used as a coloring agent. After the assassination of Valdizán by some of his workers, Cobos brought from the continent to San Cristóbal Island a group of more than a hundred workers, and tried his luck at planting sugar cane. He ruled his plantation with an iron hand, which led to his assassination in 1904. In 1897, Antonio Gil began another plantation on Isabela Island. Over the course of a whole year, from September 1904, an expedition of the Academy of Sciences of California, led by Rollo Beck, stayed in the Galápagos collecting scientific material on geology, entomology,
ornithology Ornithology is a branch of zoology that concerns the "methodological study and consequent knowledge of birds with all that relates to them." Several aspects of ornithology differ from related disciplines, due partly to the high visibility and th ...
,
botany Botany, also called , plant biology or phytology, is the science of plant life and a branch of biology. A botanist, plant scientist or phytologist is a scientist who specialises in this field. The term "botany" comes from the Ancient Greek w ...
,
zoology Zoology ()The pronunciation of zoology as is usually regarded as nonstandard, though it is not uncommon. is the branch of biology that studies the animal kingdom, including the structure, embryology, evolution, classification, habits, and ...
, and herpetology. Another expedition from that Academy was done in 1932 (Templeton Crocker Expedition) to collect insects, fish, shells,
fossil A fossil (from Classical Latin , ) is any preserved remains, impression, or trace of any once-living thing from a past geological age. Examples include bones, shells, exoskeletons, stone imprints of animals or microbes, objects preserved ...
s, birds, and plants.


From geopolitical hotspot to national park

For a long time during the early 1900s and at least through 1929, a cash-strapped Ecuador had reached out for potential buyers of the islands to alleviate financial troubles at home. The US had repeatedly expressed its interest in buying the islands for military use as they were positioned strategically guarding the
Panama Canal The Panama Canal ( es, Canal de Panamá, link=no) is an artificial waterway in Panama that connects the Atlantic Ocean with the Pacific Ocean and divides North and South America. The canal cuts across the Isthmus of Panama and is a condui ...
. Besides the United States, Japan,
Germany Germany,, officially the Federal Republic of Germany, is a country in Central Europe. It is the second most populous country in Europe after Russia, and the most populous member state of the European Union. Germany is situated betwe ...
and Chile also expressed interest in establishing bases in the islands at the turn of the century. Chile had previously acquired the
Straits of Magellan The Strait of Magellan (), also called the Straits of Magellan, is a navigable sea route in southern Chile separating mainland South America to the north and Tierra del Fuego to the south. The strait is considered the most important natural pas ...
and
Easter Island Easter Island ( rap, Rapa Nui; es, Isla de Pascua) is an island and special territory of Chile in the southeastern Pacific Ocean, at the southeasternmost point of the Polynesian Triangle in Oceania. The island is most famous for its ne ...
for strategic reasons and lieutenant Gregorio Santa Cruz argued in 1903 that possessing an island in equatorial waters, like the Galápagos, would be of great benefit since the
geopolitical Geopolitics (from Greek γῆ ''gê'' "earth, land" and πολιτική ''politikḗ'' "politics") is the study of the effects of Earth's geography (human and physical) on politics and international relations. While geopolitics usually refers to ...
situation of Chile was expected to drastically change when the Panama Canal opened. Another benefit would be to widen the security radius of Chile. Chile was alarmed by the United States plans to establish a Guantanamo-like base in the Galápagos Islands since it would mean that Chile's nitrate-rich
northern provinces The Northern Provinces of South Africa is a biogeographical area used in the World Geographical Scheme for Recording Plant Distributions (WGSRPD). It is part of the WGSRPD region 27 Southern Africa. The area has the code "TVL". It includes the So ...
would be within the range of
United States Navy The United States Navy (USN) is the maritime service branch of the United States Armed Forces and one of the eight uniformed services of the United States. It is the largest and most powerful navy in the world, with the estimated tonnage ...
. Ecuador's staunch resistance to a US purchase or bases in the islands can be credited to Chilean diplomacy, which in turn was informally backed on this issue by Great Britain and Germany. In the 1920s and 1930s, a small wave of European settlers arrived in the islands. There occurred a series of unsolved disappearances on the island of Floreana in the 1930s among the largely European expatriate residents at the time. ''The Galápagos Affair: Satan Came to Eden'' is a 2013 feature-length documentary film about the same. Ecuadorian laws provided all colonists with the possibility of receiving twenty hectares each of free land, the right to maintain their citizenship, freedom from taxation for the first ten years in Galápagos, and the right to hunt and fish freely on all uninhabited islands where they might settle. The first European colonists to arrive were Norwegians who settled briefly on Floreana, before moving on to San Cristobal and Santa Cruz. A few years later, other colonists from Europe, America and Ecuador started arriving on the islands, seeking a simpler life. Descendants of the Norwegian Kastdalen family and the German Angermeyer still live on the islands. During World War II, Ecuador authorized the United States to establish a naval base in Baltra Island, and radar stations in other strategic locations. Baltra was established as a United States Army Air Force base. Baltra was given the name of "Beta Base" along with "Alpha Base" in Nicaragua and "Gamma Base" in Salinas (continental Ecuador). The Crews stationed at Baltra and the aforementioned locations established a geographic triangle of protection in charge of patrolling the Pacific for enemy submarines, and also provided protection for the
Panama Canal The Panama Canal ( es, Canal de Panamá, link=no) is an artificial waterway in Panama that connects the Atlantic Ocean with the Pacific Ocean and divides North and South America. The canal cuts across the Isthmus of Panama and is a condui ...
. After the war, the facilities were given to the government of Ecuador. Today, the island continues as an official Ecuadorian military base. The foundations and other remains of the US base can still be seen as one crosses the island. In 1946, a penal colony was established on Isabela Island, but it was suspended in 1959. The Galápagos became a national park in 1959, and tourism started in the 1960s, imposing several restrictions upon the human population already living on the island. However, opportunities in the tourism, fishing, and farming industries attracted a mass of poor fishermen and farmers from mainland Ecuador. In the 1990s and 2000s, violent confrontations between parts of the local population and the Galápagos National Park Service occurred, including capturing and killing giant tortoises and holding staff of the Galápagos National Park Service hostage to obtain higher annual sea cucumber quotas.


Politics

The islands are administered by a provincial government. It was made a province by presidential decree by President Guillermo Rodríguez Lara on 18 February 1973. The province is divided into cantons, each covering certain islands. The capital is Puerto Baquerizo Moreno.


Demographics

The largest ethnic group is composed of Ecuadorian Mestizos, the mixed descendants of Spanish colonists and indigenous Native Americans, who arrived mainly in the last century from the
continent A continent is any of several large landmasses. Generally identified by convention rather than any strict criteria, up to seven geographical regions In geography, regions, otherwise referred to as zones, lands or territories, are areas t ...
al part of
Ecuador Ecuador ( ; ; Quechua: ''Ikwayur''; Shuar: ''Ecuador'' or ''Ekuatur''), officially the Republic of Ecuador ( es, República del Ecuador, which literally translates as "Republic of the Equator"; Quechua: ''Ikwadur Ripuwlika''; Shuar: ' ...
. Some descendants of the early European and American colonists on the islands also still remain on the islands. In 1959, approximately 1,000 to 2,000 people called the islands their home. In 1972 a census in the archipelago recorded a population of 3,488. By the 1980s, this number had risen to more than 15,000 people, and in 2010 there were 25,124 people in the Galápagos. 2021 projected population was 40,685. Five of the islands are inhabited: Baltra, Floreana, Isabela, San Cristóbal, and Santa Cruz.


Travel

Options for air travel to the Galápagos are limited to two islands: San Cristobal ( San Cristóbal Airport) and Baltra (
Seymour Airport Seymour Galapagos Ecological Airport (Spanish: ''Aeropuerto Ecológico Galápagos Seymour'') is an airport serving the island of Baltra, one of the Galápagos Islands in Ecuador. Facilities The airport became the world's first "green" airp ...
). Private aircraft must use Baltra as it is the airport equipped with overnight plane accommodations. Seymour Airport on Baltra was recently renovated (2012–2013) to accommodate larger planes. Until 1969 the only way to visit was on a private or chartered vessel. There was no regular air service until Forrest Nelson's Hotel Galápagos began the first organized tours in April 1969. Soon other travel companies brought in tour ships and yachts, and local fishermen began converting their wooden boats for rudimentary cruising with guests. These vessels were the main source of overnight accommodations in the Galápagos. Today there are about 85 yachts and ships equipped for overnight guests. In 2006 the Baltra military governed island, was opened up to limited overnight camping. Baltra also requires permits by the military government for overnight stays on the beach. Other inhabited islands also allow camping on the beaches designated as "recreational" use to the locals. All of these camping permits are limited to number of people and nights, with most nights not to exceed three. Land based hotels are opening on the inhabited islands of San Cristobal, Santa Cruz, Floreana and Isabela. By 2012, more than half the visitors to Galápagos made their tours using day boats and these small hotels. Restaurants, easy access and economy make this an attractive travel option. There are only 116 visitor sites in the Galápagos: 54 land sites and 62 scuba-diving or snorkeling sites. Small groups are allowed to visit in 2- to 4-hour shifts only, to limit impact on the area. All groups are accompanied by licensed guides.


Environmental protection policy

Though the first protective legislation for the Galápagos was enacted in 1930 and supplemented in 1936, it was not until the late 1950s that positive action was taken to control what was happening to the native flora and fauna. In 1955, the International Union for the Conservation of Nature organized a fact-finding mission to the Galápagos. Two years later, in 1957,
UNESCO The United Nations Educational, Scientific and Cultural Organization is a specialized agency of the United Nations (UN) aimed at promoting world peace and security through international cooperation in education, arts, sciences and culture. It ...
, in cooperation with the government of Ecuador, sent another expedition to study the conservation situation and choose a site for a research station. In 1959, the centenary year of
Charles Darwin Charles Robert Darwin ( ; 12 February 1809 – 19 April 1882) was an English naturalist, geologist, and biologist, widely known for his contributions to evolutionary biology. His proposition that all species of life have descended ...
's publication of ''
The Origin of Species ''On the Origin of Species'' (or, more completely, ''On the Origin of Species by Means of Natural Selection, or the Preservation of Favoured Races in the Struggle for Life''),The book's full original title was ''On the Origin of Species by Me ...
'', the Ecuadorian government declared 97.5% of the archipelago's land area a national park, excepting areas already colonised. The
Charles Darwin Foundation The Charles Darwin Foundation ( Spanish: ''Fundación Charles Darwin'') was founded in 1959, under the auspices of UNESCO and the World Conservation Union. The Charles Darwin Research Station serves as headquarters for The Foundation, and is use ...
(CDF) was founded the same year. The core responsibility of CDF, an international nongovernmental organization (NGO) constituted in Belgium, is to conduct research and provide the research findings to the government for effective management of Galápagos. CDF's research efforts began with the establishment of the
Charles Darwin Research Station Charles is a masculine given name predominantly found in English and French speaking countries. It is from the French form ''Charles'' of the Proto-Germanic name (in runic alphabet) or ''*karilaz'' (in Latin alphabet), whose meaning was ...
on Santa Cruz Island in 1964. During the early years, conservation programs, such as eradication of introduced species and protection of native species, were carried out by research station personnel. Now much of that work is accomplished by the
Galápagos National Park Galápagos National Park ( es, Parque Nacional Galápagos) was established in 1959. It began operation in 1968, and it is Ecuador's first national park and a UNESCO World Heritage Site. Park history The government of Ecuador has designated ...
Service using the research findings and methodologies developed by CDF. In 1986, the 70,000 square kilometers (27,000 sq mi) of ocean surrounding the islands was declared a
marine reserve A marine reserve is a type of marine protected area (MPA). An MPA is a section of the ocean where a government has placed limits on human activity. A marine reserve is a marine protected area in which removing or destroying natural or cultural ...
, second in size only to Australia's Great Barrier Reef. In 1990, the archipelago became a whale sanctuary.
UNESCO The United Nations Educational, Scientific and Cultural Organization is a specialized agency of the United Nations (UN) aimed at promoting world peace and security through international cooperation in education, arts, sciences and culture. It ...
recognized the islands in 1978 as a
World Heritage Site A World Heritage Site is a landmark or area with legal protection by an international convention administered by the United Nations Educational, Scientific and Cultural Organization (UNESCO). World Heritage Sites are designated by UNESCO for h ...
and in 1985, as a
biosphere reserve A nature reserve (also known as a wildlife refuge, wildlife sanctuary, biosphere reserve or bioreserve, natural or nature preserve, or nature conservation area) is a protected area of importance for flora, fauna, or features of geological or ...
. This was later extended in December 2001 to include the marine reserve. In July 2010, the World Heritage Committee agreed to remove the Galápagos Islands from its list of precious sites endangered by environmental threats or overuse. Noteworthy species include: *
Galápagos dove The Galápagos dove (''Zenaida galapagoensis'') is a species of bird in the family Columbidae. It is endemic to the Galápagos, off Ecuador. It is fairly common and is found in a wide range of open and semi-open habitats, especially in the arid ...
, ''Zenaida galapagoensis'', two subspecies * Galápagos flycatcher, ''Myiarchus magnirostris'' * Galápagos land iguanas, ''
Conolophus The Galápagos land iguanas comprise the genus ''Conolophus'' of the Galápagos Islands (Ecuador). The number of species In biology, a species is the basic unit of classification and a taxonomic rank of an organism, as well as a unit o ...
'' spp. *
Marine iguana The marine iguana (''Amblyrhynchus cristatus''), also known as the sea iguana, saltwater iguana, or Galápagos marine iguana, is a species of iguana found only on the Galápagos Islands (Ecuador). Unique among modern lizards, it is a marine rept ...
, ''Amblyrhynchus cristatus'', the only iguana feeding in the sea *
Galápagos tortoise The Galápagos tortoise or Galápagos giant tortoise (''Chelonoidis niger'') is a species of very large tortoise in the genus ''Chelonoidis'' (which also contains three smaller species from mainland South America). It comprises 15 subspecies ( ...
(Galápagos giant
tortoise Tortoises () are reptiles of the family Testudinidae of the order Testudines (Latin: ''tortoise''). Like other turtles, tortoises have a shell to protect from predation and other threats. The shell in tortoises is generally hard, and like oth ...
), ''Chelonoidis nigra'', known as ''galápago'' in Spanish, it gave the name to the islands * Galápagos green turtle, ''Chelonia mydas agassisi'', a subspecies of the
green turtle The green sea turtle (''Chelonia mydas''), also known as the green turtle, black (sea) turtle or Pacific green turtle, is a species of large sea turtle of the family Cheloniidae. It is the only species in the genus ''Chelonia''. Its range exten ...
* Galápagos racer, ''Pseudalsophis biserialis'', an endemic species of snake with two subspecies * Sea cucumbers, the cause of environmental battles with fishermen over
quota Quota may refer to: Economics * Import quota, a trade restriction on the quantity of goods imported into a country * Market Sharing Quota, an economic system used in Canadian agriculture * Milk quota, a quota on milk production in Europe * Indi ...
s of this expensive Asian delicacy *
Flightless cormorant The flightless cormorant (''Nannopterum harrisi''), also known as the Galapagos cormorant, is a cormorant endemic to the Galapagos Islands, and an example of the highly unusual fauna there. It is unique in that it is the only known cormorant th ...
, ''Phalacrocorax harrisi'' * Great frigatebird and
magnificent frigatebird The magnificent frigatebird (''Fregata magnificens'') is a seabird of the frigatebird family Fregatidae. With a length of and wingspan of it is the largest species of frigatebird. It occurs over tropical and subtropical waters off America, bet ...
* Blue-footed booby, ''Sula nebouxii'', popular among visitors for their large blue feet which they show off in courtship *
Galápagos penguin The Galápagos penguin (''Spheniscus mendiculus'') is a penguin endemic to the Galápagos Islands, Ecuador. It is the only penguin found north of the equator. Most inhabit Fernandina Island and the west coast of Isabela Island. The cool wate ...
, ''Spheniscus mendiculus'', the only living tropical penguin * Waved albatross, ''Phoebastria irrorata'', the only living tropical albatross * Galápagos hawk, ''Buteo galapagoensis'', the islands' main scavenger (at the top of the food chain) and "environmental police" * Four
endemic Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found else ...
species of Galápagos mockingbirds, the first species Darwin noticed to vary from island to island * Thirteen endemic species of
tanager The tanagers (singular ) comprise the bird family Thraupidae, in the order Passeriformes. The family has a Neotropical distribution and is the second-largest family of birds. It represents about 4% of all avian species and 12% of the Neotropica ...
s, popularly called
Darwin's finches Darwin's finches (also known as the Galápagos finches) are a group of about 18 species of passerine birds. They are well known for their remarkable diversity in beak form and function. They are often classified as the subfamily Geospizinae or t ...
. Among them is the
sharp-beaked ground finch The sharp-beaked ground finch (''Geospiza difficilis'') is a species of bird in the Darwin's finch group of the tanager family Thraupidae. It is classified as a least-concern species by the International Union for Conservation of Nature and it is ...
''Geospiza difficilis septentrionalis'' which is sometimes called the "vampire finch" for its blood-sucking habits, and the tool-using woodpecker finch, ''Camarhynchus pallidus'' *
Galápagos sea lion The Galápagos sea lion (''Zalophus wollebaeki'') is a species of sea lion that lives and breeds on the Galápagos Islands and, in smaller numbers, on Isla de la Plata (Ecuador). Being fairly social, they are often spotted sun-bathing on sandy ...
s, ''Zalophus wollebaeki'', closely related to the California sea lion, but smaller * Two endemic genera of cacti, each with a single species: '' Jasminocereus thouarsii'', the candelabra cactus, and '' Brachycereus nesioticus'', the lava cactus


Environmental threats

Introduced plants and animals, such as feral goats, cats, and cattle, brought accidentally or willingly to the islands by humans, represent the main threat to Galápagos. Quick to reproduce and with no natural predators, these alien species decimated the habitats of native species. The native animals, lacking natural predators on the islands, are defenseless to introduced predators. There are over 700 introduced plant species today. There are only 500 native and endemic species. This difference is creating a major problem for the islands and the natural species that inhabit them. These plants have invaded large areas and eliminated endemic species in the humid zones of San Cristobal, Floreana, Isabela and Santa Cruz. Some of the most harmful introduced plants are the guayaba or guava (''Psidium guajava''),
avocado The avocado (''Persea americana'') is a medium-sized, evergreen tree in the laurel family ( Lauraceae). It is native to the Americas and was first domesticated by Mesoamerican tribes more than 5,000 years ago. Then as now it was prized for ...
(''Persea americana''),
cascarilla ''Croton eluteria'', known as cascarilla, is a plant species of the genus '' Croton'' that is native to the Caribbean. It has been naturalized in other tropical regions of the Americas. It grows to be a small tree or tall shrub, rarely reaching ...
(''Cinchona pubescens''),
balsa ''Ochroma pyramidale'', commonly known as the balsa tree, is a large, fast-growing tree native to the Americas. It is the sole member of the genus ''Ochroma''. The tree is famous for its wide usage in woodworking, with the name ''balsa'' being ...
(''Ochroma pyramidale''),
hill raspberry ''Rubus niveus'' (Mysore raspberry, Ceylon raspberry, hill raspberry; , hi, काला हिसालू, translit=kala hinsalu) is a species of ''Rubus'' native to southern Asia, from Afghanistan east through India and China to Taiwan and ...
(''Rubus niveus''), various
citrus ''Citrus'' is a genus of flowering trees and shrubs in the rue family, Rutaceae. Plants in the genus produce citrus fruits, including important crops such as oranges, lemons, grapefruits, pomelos, and limes. The genus ''Citrus'' is native to ...
(orange, grapefruit, lemon), floripondio, higuerilla (''
Ricinus communis ''Ricinus communis'', the castor bean or castor oil plant, is a species of perennial flowering plant in the spurge family, Euphorbiaceae. It is the sole species in the monotypic genus, ''Ricinus'', and subtribe, Ricininae. The evolution of cas ...
'') trees and the elephant grass, ''
Pennisetum purpureum ''Cenchrus purpureus'', synonym ''Pennisetum purpureum'', also known as Napier grass, elephant grass or Uganda grass, is a species of perennial tropical grass native to the African grasslands. arrell, G., Simons, S. A., & Hillocks, R. J. (2002). ...
''. Many species were introduced to the Galápagos by pirates. Thor Heyerdahl quoted documents that mention the
Viceroy of Peru The viceroys of Peru ruled the Viceroyalty of Peru from 1544 to 1824 in the name of the monarch of Spain. The territories under ''de jure'' rule by the viceroys included in the 16th and 17th century almost all of South America except eastern Braz ...
, knowing that British pirates ate the goats that they themselves had released in the islands, ordered dogs to be freed there to eliminate the goats. Also, when colonization of Floreana by José de Villamil failed, he ordered the goats, donkeys, cattle and other animals from the farms in Floreana be transferred to other islands for the purpose of later colonization. Non-native goats, pigs, dogs, rats, cats, mice, sheep, horses, donkeys, cows, poultry, ants, cockroaches, and some parasites inhabit the islands today. Dogs and cats attack the tame birds and destroy the nests of birds, land tortoises, and marine turtles. They sometimes kill small Galápagos tortoises and iguanas. Pigs are even more harmful, covering larger areas and destroying the nests of tortoises, turtles and iguanas, as well as eating the animals' native food. Pigs also knock down vegetation in their search for roots and insects. This problem abounds in Cerro Azul volcano and Isabela, and in Santiago, pigs may be the cause of the disappearance of the land iguanas that were so abundant when Darwin visited. The black rat (''Rattus rattus'') attacks small Galápagos tortoises when they leave the nest, so in Pinzón they stopped the reproduction for a period of more than 50 years; only adults were found on that island. Also, where the black rat is found, the endemic rat has disappeared. Cattle and donkeys eat all the available vegetation and compete with native species for scarce water. In 1959, fishermen introduced one male and two female goats to Pinta island; by 1973, the National Park service estimated the population of goats to be over 30,000 individuals. Goats were also introduced to Marchena in 1967 and to Rabida in 1971. A goat eradication program, however, cleared the goats from Pinta and Santiago and most of the goat population from Isabela, and, by 2006, all
feral A feral () animal or plant is one that lives in the wild but is descended from domesticated individuals. As with an introduced species, the introduction of feral animals or plants to non-native regions may disrupt ecosystems and has, in some ...
pigs, donkeys and non-sterile goats had been eliminated from Santiago and Isabela, the largest islands with the worst problems due to non-native mammals. Six species of small non-native vertebrates have established self-sufficient populations in Galápagos and may become invasive: Fowler's snouted tree frog ''
Scinax quinquefasciatus ''Scinax quinquefasciatus'' is a species of frog in the family Hylidae. It is found in Colombia and Ecuador. Its natural habitats are subtropical or tropical dry forests, subtropical or tropical moist lowland forests, subtropical or tropical moi ...
'', common house gecko '' Hemidactylus frenatus'', mourning gecko ''
Lepidodactylus lugubris ''Lepidodactylus lugubris'', known as the mourning gecko or common smooth-scaled gecko, is a species of lizard, a gecko of the family Gekkonidae. Description ''Lepidodactylus lugubris'' measure 8.5-10 cm in length including tail (4-4.4 cm snou ...
'', dwarf gecko '' Gonatodes caudiscutatus'', Peters' leaf-toed gecko ''
Phyllodactylus reissii ''Phyllodactylus reissii'', also known commonly as Peters' leaf-toed gecko or the coastal leaf-toed gecko, is a species of lizard in the family Phyllodactylidae. The species is endemic to northwestern South America. Etymology The specific nam ...
'', and smooth-billed ani '' Crotophaga ani''. Domestic fowl ''
Gallus gallus The red junglefowl (''Gallus gallus'') is a tropical bird in the family Phasianidae. It ranges across much of Southeast Asia and parts of South Asia. It was formerly known as the Bankiva or Bankiva Fowl. It is the species that gave rise to the ...
'' holds feral populations, which may have self-sufficient populations, but evidence is unclear. The fast-growing poultry industry on the inhabited islands has been cause for concern from local conservationists, who fear domestic birds could introduce disease into the endemic wild bird populations. The Galápagos marine sanctuary is under threat from a host of illegal fishing activities, in addition to other problems of development. The most pressing threat to the Marine Reserve comes from local, mainland and foreign fishing targeting marine life illegally within the Reserve, such as sharks (hammerheads and other species) for their fins, and the harvest of sea cucumbers out of season. Development threatens both land and sea species. The growth of both the tourism industry and local populations fuelled by high birth rates and illegal immigration threaten the wildlife of the Archipelago. The grounding of the oil tanker '' Jessica'' in 2001 and the subsequent oil spill brought this threat to world attention. In 2007,
UNESCO The United Nations Educational, Scientific and Cultural Organization is a specialized agency of the United Nations (UN) aimed at promoting world peace and security through international cooperation in education, arts, sciences and culture. It ...
put the Galápagos Islands on their
List of World Heritage in Danger The List of World Heritage in Danger is compiled by the United Nations Educational, Scientific and Cultural Organization (UNESCO) through the World Heritage Committee according to Article 11.4 of the World Heritage Convention,Full title: ''Conv ...
because of threats posed by invasive species, unbridled tourism and overfishing. On 29 July 2010, the
World Heritage Committee The World Heritage Committee selects the sites to be listed as UNESCO World Heritage Sites, including the World Heritage List and the List of World Heritage in Danger, defines the use of the World Heritage Fund and allocates financial assistance ...
decided to remove the Galápagos Islands from the list because the Committee found significant progress had been made by Ecuador in addressing these problems. On 28 January 2008,
Galápagos National Park Galápagos National Park ( es, Parque Nacional Galápagos) was established in 1959. It began operation in 1968, and it is Ecuador's first national park and a UNESCO World Heritage Site. Park history The government of Ecuador has designated ...
official Victor Carrion announced 53 sea lions (13 pups, 25 youngsters, 9 males and 6 females) were killed at the Galápagos Islands nature reserve on Pinta, with their heads caved in. In 2001,
poachers Poaching has been defined as the illegal hunting or capturing of wild animals, usually associated with land use rights. Poaching was once performed by impoverished peasants for subsistence purposes and to supplement meager diets. It was set a ...
killed 35 male sea lions. The Galápagos Islands were short-listed as a candidate to be one of the New7Wonders of Nature by the New7Wonders of Nature Foundation. As of February 2009, the archipelago was ranked first in Group B, the category for islands.New 7 Wonders of the World: Live Ranking
The islands' biodiversity is under threat from several sources. The human population is growing at a rate of 8% per year (1995). Introduced species have caused damage, and in 1996 a US$5 million, five-year eradication plan commenced in an attempt to rid the islands of introduced species such as goats, rats, deer, and donkeys. Except for the rats, the project was essentially completed in 2006. Rats have only been eliminated from the smaller Galápagos Islands of Rábida and Pinzón.
El Niño El Niño (; ; ) is the warm phase of the El Niño–Southern Oscillation (ENSO) and is associated with a band of warm ocean water that develops in the central and east-central equatorial Pacific (approximately between the International Date ...
has adversely affected the marine ecosystem. In January 2001, an oil slick from a stranded tanker threatened the islands, but winds and shifting ocean currents helped disperse the oil before much damage was done. The devastating El Niño of 1982–83 saw almost six times as much rain as normal in the Galapagos and created a wildlife catastrophe. The 1997–98 El Niño adversely affected wildlife in the waters surrounding the islands, as the waters were warmer than normal. Corals and barnacles suffered, hammerhead sharks were driven away, and most of the island's seabirds failed to breed in 1997–98. The mortality rate of marine iguanas rose as the green algae they feed on was replaced by inedible red algae. During the 1982–83 El Niño, 70% of the marine iguanas starved to death because of this."Galápagos Islands" in ''The Hutchinson Unabridged Encyclopedia with Atlas and Weather guide'' (2010). Abington: Helicon.


See also

*
List of birds of the Galápagos Islands This list of birds recorded in the Galápagos Islands includes species documents from the Galápagos Islands of Ecuador, where 185 species have been documented as of February 2019. Of them, 31 are endemic, three nest only in the Galápagos, and ...
*
List of animals in the Galápagos Islands This is a list of animals that live in the Galápagos Islands. The fauna of the Galápagos Islands include a total of 9,000 confirmed species. Of them, none have been introduced by humans, and seventeen are endemic. Mammals Rodents *Galápagos ...
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Galápagos National Park Galápagos National Park ( es, Parque Nacional Galápagos) was established in 1959. It began operation in 1968, and it is Ecuador's first national park and a UNESCO World Heritage Site. Park history The government of Ecuador has designated ...
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Galápagos hotspot The Galápagos hotspot is a volcanic hotspot in the East Pacific Ocean responsible for the creation of the Galápagos Islands as well as three major aseismic ridge systems, Carnegie, Cocos and Malpelo which are on two tectonic plates. The hots ...
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Galápagos Islands xeric scrub The Galápagos Islands xeric scrub, also known as the Galápagos Islands scrubland mosaic, is a terrestrial deserts and xeric shrublands ecoregion that covers the Galápagos Islands. The Galápagos Islands are volcanic in origin, and remote from c ...
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Galápagos Province Galápagos () is a province of Ecuador in the country's Insular region, located approximately off the western coast of the mainland. The capital is Puerto Baquerizo Moreno. The province administers the Galápagos Islands, a group of tiny vol ...
* Darwin's Arch *
Johanna Angermeyer Johanna Angermeyer, American-born author and artist, (born 1948) is the author of the critically acclaimed ''My Father's Island: A Galapagos Quest'' (now in its 9 reprint) and children's picture books, (illustrated by the author): "Is Your Mama An ...
, author of various books featuring the Galápagos Islands


References

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Further reading

* Black, Juan (1973). ''Galápagos, Archipiélago del Ecuador''. (Quito, Ecuador). Comprehensive monograph by a former officer of Galápagos National Park, financed by the World Wildlife Fund and the Charles Darwin Foundation for the Galápagos Islands * Grant, K. Thalia and Estes, Gregory B. (2009). ''Darwin in Galapagos: Footsteps to a New World''. Princeton University Press, Princeton

* Thor Heyerdahl, Heyerdahl, Thor; & Skjolsvold, Arne (1956). ''Archaeological Evidence of Pre-Spanish Visits to the Galápagos Islands'', Memoirs 12,
Society for American Archaeology The Society for American Archaeology (SAA) is a professional association for the archaeology of the Americas. It was founded in 1934 and its headquarters are in based in Washington, D.C. , it has 7,500 members. Its current president is Deborah L. ...
. * Müller, Bodo; & Stolt, Matthias (2003). ''Galápagos Die verwunschenen Inseln''. (BLV). . * Quammen, David (1996). ''The Song of the Dodo''. Touchstone, New York. * Romero, Simon (4 October 2009)
"To Protect Galápagos, Ecuador Limits a Two-Legged Species"
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The New York Times ''The New York Times'' (''the Times'', ''NYT'', or the Gray Lady) is a daily newspaper based in New York City with a worldwide readership reported in 2020 to comprise a declining 840,000 paid print subscribers, and a growing 6 million paid d ...
''. * Perry, Roger (1972). ''The Galapagos Islands''. (Dodd, Mead & Company). Short history illustrated with photographs and a map by a former director of Charles Darwin Research Station. * Wittmer, Margret (2013). ''Floreana: A Woman's Pilgrimage to the Galápagos''. (Moyer Bell Limited). Memoir written by the matriarch of the first family to successfully inhabit Floreana, after settling there in 1932. 2013 paperback edition (originally published by the Travel Book Club in 1961)


External links

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Galápagos geology, with general information on the Galápagos Islands
{{DEFAULTSORT:Galapagos Islands *Islands Biosphere reserves of Ecuador Surfing locations in Ecuador Ecoregions of Ecuador Pacific islands of Ecuador Seabird colonies States and territories established in 1973 World Heritage Sites in Ecuador Tourist attractions in Galápagos Province Articles containing video clips Tropical Eastern Pacific