Guttulate Foliage-gleaner
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The guttulate foliage-gleaner (''Syndactyla guttulata'') is a species of
bird Birds are a group of warm-blooded vertebrates constituting the class (biology), class Aves (), characterised by feathers, toothless beaked jaws, the Oviparity, laying of Eggshell, hard-shelled eggs, a high Metabolism, metabolic rate, a fou ...
in the Furnariinae subfamily of the ovenbird
family Family (from ) is a Social group, group of people related either by consanguinity (by recognized birth) or Affinity (law), affinity (by marriage or other relationship). It forms the basis for social order. Ideally, families offer predictabili ...
Furnariidae. It is
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to Venezuela.


Taxonomy and systematics

The guttulate foliage-gleaner was originally described as ''Anabazenops guttulatus''. It early acquired the English name "guttulated foliage-gleaner", a name which persisted until it was pointed out that the word ''guttulated'' does not exist in English. "Webster's International indeed states that the English noun ''guttula'' is an obscure word for a drop-shaped spot, and the adjectival form is ''guttulate'', not "guttulated". The guttulate foliage-gleaner has two subspecies, the
nominate Nomination is part of the process of selecting a candidate for either election to a public office, or the bestowing of an honor or award. A collection of nominees narrowed from the full list of candidates is a short list. Political office In th ...
''S. g. guttulata'' ( Sclater, PL, 1858) and ''S. g. pallida'' ( Zimmer, JT & Phelps, WH, 1944).


Description

The guttulate foliage-gleaner is long and weighs . It is a largish furnariid with a more wedge-shaped bill than many others of its genus. The sexes have the same plumage. Adults of the nominate subspecies have a faint dull ochraceous
supercilium The supercilium is a plumage feature found on the heads of some bird species. It is a stripe which runs from the base of the bird's beak above its eye, finishing somewhere towards the rear of the bird's head.Dunn and Alderfer (2006), p. 10 Also k ...
that fades on its way to the nape, and dark brown ear
coverts A covert feather or tectrix on a bird is one of a set of feathers, called coverts (or ''tectrices''), which cover other feathers. The coverts help to smooth airflow over the wings and tail. Ear coverts The ear coverts are small feathers behind t ...
with narrow buff streaks on an otherwise pale-spotted dark brown face. Their crown is dark rufescent brown with thin rusty streaks, and paler rusty spots on the forehead. Their back is dark rufescent brown wide black-edged rusty to whitish buff streaks. Their rump, uppertail coverts, and tail are reddish chestnut. Their wings are dark rufescent brown with reddish chestnut tips on the flight feathers. Their throat is yellowish with dark brown at its lower edge, their breast and belly dark brown with buffy streaks that fade by the lower belly, and their undertail coverts more rufescent brown. Their iris is dark brown, their
maxilla In vertebrates, the maxilla (: maxillae ) is the upper fixed (not fixed in Neopterygii) bone of the jaw formed from the fusion of two maxillary bones. In humans, the upper jaw includes the hard palate in the front of the mouth. The two maxil ...
dark gray to blackish, their
mandible In jawed vertebrates, the mandible (from the Latin ''mandibula'', 'for chewing'), lower jaw, or jawbone is a bone that makes up the lowerand typically more mobilecomponent of the mouth (the upper jaw being known as the maxilla). The jawbone i ...
pale horn-gray with some dark at the base, and their legs and feet olive-gray. Subspecies ''S. g. pallida'' is similar to the nominate with paler edges on the upperparts' feathers and a paler rufous rump and tail.Remsen, Jr., J. V. (2020). Guttulate Foliage-gleaner (''Syndactyla guttulata''), version 1.0. In Birds of the World (J. del Hoyo, A. Elliott, J. Sargatal, D. A. Christie, and E. de Juana, Editors). Cornell Lab of Ornithology, Ithaca, NY, USA. https://doi.org/10.2173/bow.gufgle1.01 retrieved August 31, 2023


Distribution and habitat

The guttulate foliage-gleaner is found near the north coast of Venezuela. The nominate subspecies occurs in Sierra de San Luis in Falcón state and in the western part of the
Venezuelan Coastal Range The Venezuelan Coastal Range ( or ), also known as Venezuelan Caribbean Mountain System (), is a mountain range system and one of the eight natural regions of Venezuela, that runs along the central and eastern portions of Venezuela's northern coa ...
as far east as
Aragua Aragua State (, ) is one of the 23 states of Venezuela. It is located in the north-central region of Venezuela. It has plains, jungles and Caribbean beaches. The most popular beaches are Cata and Choroni. It has Venezuela's first national par ...
state. ''S. g. pallida'' is found disjunctly in the eastern part of the Coastal Range between
Anzoátegui Anzoátegui State (, ) is one of the 23 states of Venezuela, states of Venezuela, located in the northeastern region of the country. Anzoátegui is well known for its beaches that attract many visitors. Its coast consists of a single beach approx ...
and
Monagas Monagas State (, ) is one of the 23 states of Venezuela. Monagas State covers a total surface area of and, as of the 2011 census, had a population of 905,443. Monagas State is surrounded by Sucre State in the north, Anzoátegui State in the ...
states. The species inhabits
primary Primary or primaries may refer to: Arts, entertainment, and media Music Groups and labels * Primary (band), from Australia * Primary (musician), hip hop musician and record producer from South Korea * Primary Music, Israeli record label Work ...
montane
evergreen forest An evergreen forest is a forest made up of evergreen trees. They occur across a wide range of climatic zones, and include trees such as conifers and holly in cold climates, eucalyptus, live oak, acacias, magnolia, and banksia in more temperate zo ...
and
secondary forest A secondary forest (or second-growth forest) is a forest or woodland area which has regenerated through largely natural processes after human-caused Disturbance (ecology), disturbances, such as Logging, timber harvest or agriculture clearing, or ...
at elevations between .


Behavior


Movement

The guttulate foliage-gleaner is a year-round resident throughout its range.


Feeding

The guttulate foliage-gleaner feeds mostly on
arthropod Arthropods ( ) are invertebrates in the phylum Arthropoda. They possess an arthropod exoskeleton, exoskeleton with a cuticle made of chitin, often Mineralization (biology), mineralised with calcium carbonate, a body with differentiated (Metam ...
s. It typically forages singly or in pairs and readily joins
mixed-species feeding flock A mixed-species feeding flock, also termed a mixed-species foraging flock, mixed hunting party or informally bird wave, is a flock of usually insectivorous birds of different species that join each other and move together while foraging. These ar ...
s. It also occasionally follows
army ant The name army ant (or legionary ant or ''marabunta'') is applied to over 200 ant species in different lineages. Because of their aggressive predatory foraging groups, known as "raids", a huge number of ants forage simultaneously over a limited ...
swarms. It mostly forages in dense undergrowth in the forest's understory, but will rarely also feed up to the mid-storey. It gleans for its prey among dead leaves and
bromeliad The Bromeliaceae (the bromeliads) are a Family (biology), family of monocot flowering plants of about 80 genera and 3700 known species, native mainly to the Tropics, tropical Americas, with several species found in the American subtropics and on ...
s and also flakes bark and chisels into twigs. It often clings sideways and upside down on branches while foraging.


Breeding

The guttulate foliage-gleaner is assumed to be monogamous. What could have been its nest was in a twig-filled cavity in a wall. Nothing else is known about its breeding biology.


Vocalization

The guttulate foliage-gleaner's song is "a harsh, accelerating 'cjak, cjak, czak czak-zak-zak-zak-za-za-za', sometimes followed by a few more notes at end". Its call is "a harsh, rough 'chak' ".


Status

The
IUCN The International Union for Conservation of Nature (IUCN) is an international organization working in the field of nature conservation and sustainable use of natural resources. Founded in 1948, IUCN has become the global authority on the status ...
has assessed the guttulate foliage-gleaner as being of Least Concern. Though it has a small range and an unknown population size, the latter is believed to be stable. No immediate threats have been identified. It is considered uncommon to locally fairly common and occurs in
Henri Pittier National Park Henri Pittier National Park is the oldest national park in Venezuela, originally created in 1937 under the name of ''Rancho Grande'' by decree of President Eleazar López Contreras. In 1953 the park was renamed in honor of Henri Pittier, a distin ...
.


References

{{Taxonbar, from=Q1259097 guttulate foliage-gleaner Birds of the Venezuelan Coastal Range guttulate foliage-gleaner guttulate foliage-gleaner Endemic birds of Venezuela Taxonomy articles created by Polbot