Georgiacetus BW
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''Georgiacetus'' is an
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
genus Genus (; : genera ) is a taxonomic rank above species and below family (taxonomy), family as used in the biological classification of extant taxon, living and fossil organisms as well as Virus classification#ICTV classification, viruses. In bino ...
of ancient whale known from the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
period of the
United States The United States of America (USA), also known as the United States (U.S.) or America, is a country primarily located in North America. It is a federal republic of 50 U.S. state, states and a federal capital district, Washington, D.C. The 48 ...
. Fossils are known from Georgia, Alabama, and Mississippi and protocetid fossils from the right time frame, but not yet confirmed as ''Georgiacetus'', have been found in Texas () and South Carolina (). created a new clade, Pelagiceti, for the common ancestor of
Basilosauridae Basilosauridae is a family of extinct cetaceans that lived during the middle to late Eocene. Basilosaurids are known from all continents including Antarctica, and are probably the first fully aquatic cetaceans.Buono M, Fordyce R.E., Marx F.G., F ...
and all of its descendants, including Neoceti, the living cetaceans. He placed ''Georgiacetus'' near the base of this clade together with ''
Eocetus ''Eocetus'' is an extinct Protocetidae, protocetid Archaeoceti, early whale known from the early-late Eocene (Bartonian, ) Giushi Formation in Mokattam, Gebel Mokattam, (, paleocoordinates ) outside Cairo, Egypt. Fossil remains have also been di ...
'' and perhaps ''
Babiacetus ''Babiacetus'' is an Extinction, extinct genus of early cetacean that lived during the late Lutetian middle Eocene of India ().. Retrieved April 2013. It was named after its type locality, the Harudi Formation in the Babia Hills (: paleocoordin ...
'' because of the assumed presence of a fluke and very compressed posterior caudal vertebrae in these genera. ''Georgiacetus'' is an extinct protocetid (early whale) which lived about and hunted the rich, Suwannee Current powered coastal sea which once covered the Southeastern United States. This was during the earliest
Bartonian The Bartonian is, in the International Commission on Stratigraphy's (ICS) geologic time scale, a stage or age in the middle of the Eocene Epoch or Series. The Bartonian Age spans the time between . It is preceded by the Lutetian and is follow ...
Stage of the
Eocene The Eocene ( ) is a geological epoch (geology), epoch that lasted from about 56 to 33.9 million years ago (Ma). It is the second epoch of the Paleogene Period (geology), Period in the modern Cenozoic Era (geology), Era. The name ''Eocene'' comes ...
Epoch (). Current research puts ''Georgiacetus'' as the link between the protocetids and modern whales, making the Georgia whale a scientifically important ancestor to all modern whales. An articulated and completed cast of the find is currently on display at the
Georgia Southern University Georgia Southern University (informally known as Southern or Georgia Southern) is a public university, public research university in the U.S. state of Georgia (U.S. state), Georgia. The largest campus is in Statesboro, Georgia, Statesboro, with ...
Museum in
Statesboro, Georgia Statesboro is the most populous city in and the county seat of Bulloch County, Georgia, United States. Located in the southeastern part of the state, its population was 33,438 at the 2020 United States census, 2020 census. It is the principal city ...
; Georgia Southern University also houses the actual 1983 ''Georgiacetus'' fossils and has the animal’s fossilized hip bones on display.


Discovery

''Georgiacetus'' was discovered in 1983 during the construction of the nuclear power plant Plant Vogtle in
Burke County, Georgia Burke County is a county located along the eastern border of the U.S. state of Georgia in the Piedmont. As of the 2020 census, the population was 24,596. The county seat is Waynesboro. Burke County is part of the Augusta-Richmond County, G ...
(in the
Lutetian The Lutetian is, in the geologic timescale, a stage (stratigraphy), stage or age (geology), age in the Eocene. It spans the time between . The Lutetian is preceded by the Ypresian and is followed by the Bartonian. Together with the Bartonian it ...
Blue Bluff Formation, , paleocoordinates ). The find consisted of three individuals all identified as belonging to the same species; two of the individuals were represented by isolated bones recovered separately which included a few vertebrae, ribs and tooth crowns. The main find was discovered during the cutting of a drainage channel for the plant's River Water Intake Structure on the
Savannah River The Savannah River is a major river in the Southeastern United States, forming most of the border between the states of Georgia (U.S. state), Georgia and South Carolina. The river flows from the Appalachian Mountains to the Atlantic Ocean, ...
at a depth of about below the surface. It was a 75%-80% complete individual consisting of more than 60 bones and teeth; including a well-preserved skull and mandible. Fortunately, Georgia Power halted construction activities in the area and assisted in the recovery effort. The main mass was collected from an area about square, they were not articulated. There was evidence of light scavenging with scratches on one rib but only ten shark’s teeth were recovered with the find. There was no duplication of fossilized bones in the main mass so a single individual was represented. This was an adult animal with fully erupted and well-worn molars. There was no apparent cause of death. The main mass of bones were first reviewed in the field (1983) by geologist E. A. Shapiro of the Georgia Geologic Survey (which was "abolished" in 2004). On Shapiro’s recommendation, Georgia Southern University was called in and Richard Petkewich and Gale Bishop led the effort to recover the specimen.


Description


Cranium and dentition

Dr. Richard Hulbert, also with Georgia Southern University at that time, led the research team which described the find as a new genus and species in 1998. The skull is more than long, wide and about deep, fleshed out it would have been decidedly larger. The structure of its inner ear bones confirms ''Georgiacetus'' as belonging with the whales; throughout their long history, all whales and their terrestrial ancestors have possessed an identifiable and unique inner ear bone structure. It’s hard to know the total length of the animal as no legs or tail vertebrae were found, but researchers estimate a length of at considering the head would have been nearly three feet long for a living animal. ''Georgiacetus'' was equipped as an active predator capable of taking and processing large prey. There was a prominent crest at the top rear of the skull which would have anchored powerful jaw muscles. The front teeth were curved, banana-shaped (though in life most of the banana shape would have been deep in the jaw with just the tip exposed) and peg-like. These teeth were adapted to seize and hold struggling prey; moving back in the jaws there were deeply rooted triangular edged teeth for shearing, and further back were sharp molars for crushing. Large fish, birds or turtles could have been caught and processed into manageable chunks. The tooth and skull arrangement shows a clear, direct relationship between the
Basilosauridae Basilosauridae is a family of extinct cetaceans that lived during the middle to late Eocene. Basilosaurids are known from all continents including Antarctica, and are probably the first fully aquatic cetaceans.Buono M, Fordyce R.E., Marx F.G., F ...
family and ''Georgiacetus''. Basilosaurids possessed teeth and a skull remarkably similar in function and structure, both have
nostril A nostril (or naris , : nares ) is either of the two orifices of the nose. They enable the entry and exit of air and other gasses through the nasal cavities. In birds and mammals, they contain branched bones or cartilages called turbinates ...
s (blowholes) located halfway back on the snout, just in front of the eyes. The basilosaurid tooth arrangement appears as a more efficient and specialized version of the ''Georgiacetus'' arrangement. ''Georgiacetus'' predated the basilosaurids by nearly three million years; the Georgia Whale was very likely the source of the basilosaurids.


Postcranium

described new ''Georgiacetus'' material from Alabama and Mississippi. These finds included several tail vertebrae, teeth, and a partial mandible. The most important find was a single end-of-tail vertebra, one which would have occupied the very end of the vertebral column. This end-of-tail vertebra possessed two distinct, muscle supporting flanges. It did not show that simplified, compressed end-of-tail arrangement seen in both modern whales and ''Basilosaurus''. It did not show any fluke supporting adaptations. This single vertebra showed that ''Georgiacetus'' apparently lacked a fluke and likely swam with its hind legs. ''Georgiacetus'' had a tail and lacked the fluke present in slightly younger fossils. It probably swam using its hindlimbs by wiggling its hips and moving its trunk up and down, a locomotor behaviour abandoned by modern whales. Whales evolved in South Asia, and it was previously thought that the fluke helped early whales spread across Earth from there, so ''Georgiacetus''' presence in America and its legs and tail contradicts this hypothesis. also established the clade Pelagiceti to show the relationship between ''Georgiacetus'', the basilosaurids and modern whales. This clade begins just after ''Georgiacetus'', includes the basilosaurids, baleen and toothed whales. This shows that ''Georgiacetus'' led to the basilosaurids, which in turn led to modern whales. The right and left hip bones of ''Georgiacetus'' were recovered though no hind leg or tail material was preserved. The hip bones were relatively large and showed well developed sockets. This indicates that ''Georgiacetus'' had well developed hind legs. Because the hip bones were so much larger than those known from ''Basilosaurus'', and ''Basilosaurus'' was a dramatically larger animal than ''Georgiacetus'', it was cautiously assumed that the hind legs of ''Georgiacetus'' were much larger, relative to body size, than the tiny hind legs of ''Basilosaurus''. Yet the hip bones were not fused to the spine, (neither are those of ''Basilosaurus'') which suggests that the Georgia whale probably could not support its own weight out of the water.


Paleoenvironment

Global sea levels were about higher than modern levels when these whales lived. The fossils were discovered inland from the modern coastline. The invertebrate fossils recovered along with the main mass of bones showed that the whale died well off-shore in a shallow, open water environment approximately from the coastline (today Georgia's
Fall Line A fall line (or fall zone) is the area where an upland region and a coastal plain meet and is noticeable especially the place rivers cross it, with resulting rapids or waterfalls. The uplands are relatively hard crystalline basement rock, and the ...
). These invertebrate fossils also allowed accurate dating of the remains.


Dispersal into North America

''Georgiacetus'' was held as the earliest North American whale until published a paper on '' Carolinacetus'', a slightly older, more primitive whale than ''Georgiacetus'', based on a partial skull, mandible, several teeth, ribs and vertebra. An important find in itself, one of the interesting questions the authors asked was how did whales get to North America? Whales emerged about 52 million years ago in the
Tethys Ocean The Tethys Ocean ( ; ), also called the Tethys Sea or the Neo-Tethys, was a prehistoric ocean during much of the Mesozoic Era and early-mid Cenozoic Era. It was the predecessor to the modern Indian Ocean, the Mediterranean Sea, and the Eurasia ...
(between present-day India and Asia). It was a warm, fertile shallow sea very much like the one which covered southeastern United States 40 million years ago when whales (''Georgiacetus'' & ''Carolinacetus'') were already present in America. There are two schools of thought on how they got there: one involves a direct, deep water Atlantic crossing, the other involves a polar route northwards up the European coastline, across to Greenland, and southward along the North American coastline. This polar route would not have been a cold water trip as the climate was much warmer then and Greenland was very likely green. Both routes would require long deepwater crossings. Also, as sea levels were much higher at that time, the journey would be longer than it is today. Whichever route taken, the fact that the journey occurred shows that by the time ''Carolinacetus'' and ''Georgiacetus'' lived, whales (which did not yet possess flukes) were already capable of extended deep water activity.


Notes


References

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External links

* {{Taxonbar, from=Q2658298 Protocetidae Monotypic prehistoric cetacean genera Fossil taxa described in 1998 Eocene mammals of North America