A generalist species is able to thrive in a wide variety of environmental conditions and can make use of a variety of different
resources
Resource refers to all the materials available in our environment which are technologically accessible, economically feasible and culturally sustainable and help us to satisfy our needs and wants. Resources can broadly be classified upon their ...
(for example, a
heterotroph
A heterotroph (; ) is an organism that cannot produce its own food, instead taking nutrition from other sources of organic carbon, mainly plant or animal matter. In the food chain, heterotrophs are primary, secondary and tertiary consumers, but ...
with a varied
diet). A specialist species can thrive only in a narrow range of environmental conditions or has a limited diet. Most
organism
In biology, an organism () is any life, living system that functions as an individual entity. All organisms are composed of cells (cell theory). Organisms are classified by taxonomy (biology), taxonomy into groups such as Multicellular o ...
s do not all fit neatly into either group, however. Some species are highly specialized (the most extreme case being monophagous,
eating one specific type of food), others less so, and some can tolerate many different environments. In other words, there is a continuum from highly specialized to broadly generalist species.
Description
Omnivore
An omnivore () is an animal that has the ability to eat and survive on both plant and animal matter. Obtaining energy and nutrients from plant and animal matter, omnivores digest carbohydrates, protein, fat, and fiber, and metabolize the nut ...
s are usually generalists.
Herbivore
A herbivore is an animal anatomically and physiologically adapted to eating plant material, for example foliage or marine algae, for the main component of its diet. As a result of their plant diet, herbivorous animals typically have mouthp ...
s are often specialists, but those that eat a variety of plants may be considered generalists. A well-known example of a specialist animal is the monophagous
koala
The koala or, inaccurately, koala bear (''Phascolarctos cinereus''), is an arboreal herbivorous marsupial native to Australia. It is the only extant representative of the family Phascolarctidae and its closest living relatives are the ...
, which subsists almost entirely on
eucalyptus
''Eucalyptus'' () is a genus of over seven hundred species of Flowering plant, flowering trees, shrubs or Mallee (habit), mallees in the Myrtaceae, myrtle Family (biology), family, Myrtaceae. Along with several other genera in the Tribe (biology) ...
leaves. The
raccoon
The raccoon ( or , ''Procyon lotor''), sometimes called the common raccoon to distinguish it from other species, is a mammal native to North America. It is the largest of the procyonid family, having a body length of , and a body weight o ...
is a generalist, because it has a natural range that includes most of North and Central America, and it is omnivorous, eating
berries
A berry is a small, pulpy, and often edible fruit. Typically, berries are juicy, rounded, brightly colored, sweet, sour or tart, and do not have a stone or pit, although many pips or seeds may be present. Common examples are strawberries, rasp ...
, insects such as butterflies, eggs, and various small animals.
The distinction between generalists and specialists is not limited to animals. For example, some plants require a narrow range of temperatures, soil conditions and
precipitation
In meteorology, precipitation is any product of the condensation of atmospheric water vapor that falls under gravitational pull from clouds. The main forms of precipitation include drizzle, rain, sleet, snow, ice pellets, graupel and hai ...
to survive while others can tolerate a broader range of conditions. A
cactus
A cactus (, or less commonly, cactus) is a member of the plant family Cactaceae, a family comprising about 127 genera with some 1750 known species of the order Caryophyllales. The word ''cactus'' derives, through Latin, from the Ancient Gre ...
could be considered a specialist species. It will die during winters at high
latitude
In geography, latitude is a coordinate that specifies the north– south position of a point on the surface of the Earth or another celestial body. Latitude is given as an angle that ranges from –90° at the south pole to 90° at the north po ...
s or if it receives too much water.
When
body weight is controlled for, specialist feeders such as
insectivore
A robber fly eating a hoverfly
An insectivore is a carnivorous animal or plant that eats insects. An alternative term is entomophage, which can also refer to the human practice of eating insects.
The first vertebrate insectivores wer ...
s and
frugivore
A frugivore is an animal that thrives mostly on raw fruits or succulent fruit-like produce of plants such as roots, shoots, nuts and seeds. Approximately 20% of mammalian herbivores eat fruit. Frugivores are highly dependent on the abundance an ...
s have larger
home range
A home range is the area in which an animal lives and moves on a periodic basis. It is related to the concept of an animal's territory which is the area that is actively defended. The concept of a home range was introduced by W. H. Burt in 1943. H ...
s than generalists like some
folivores (leaf-eaters), whose food-source is less abundant; they need a bigger area for
foraging
Foraging is searching for wild food resources. It affects an animal's fitness because it plays an important role in an animal's ability to survive and reproduce. Foraging theory is a branch of behavioral ecology that studies the foraging behavi ...
.
An example comes from the research of
Tim Clutton-Brock, who found that the
black-and-white colobus, a folivore generalist, needs a home range of only 15
ha. On the other hand, the more specialized
red colobus monkey has a home range of 70 ha, which it requires to find patchy shoots, flowers and fruit.
When environmental conditions change, generalists are able to adapt, but specialists tend to fall victim to
extinction
Extinction is the termination of a kind of organism or of a group of kinds ( taxon), usually a species. The moment of extinction is generally considered to be the death of the last individual of the species, although the capacity to breed ...
much more easily. For example, if a species of fish were to go extinct, any specialist
parasite
Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
s would
also face extinction. On the other hand, a species with a highly specialized
ecological niche
In ecology, a niche is the match of a species to a specific environmental condition.
Three variants of ecological niche are described by
It describes how an organism or population responds to the distribution of resources and competitors (fo ...
is more effective at
competing with other organisms. For example, a fish and its
parasite
Parasitism is a Symbiosis, close relationship between species, where one organism, the parasite, lives on or inside another organism, the Host (biology), host, causing it some harm, and is Adaptation, adapted structurally to this way of lif ...
s are in an
evolutionary arms race
In evolutionary biology, an evolutionary arms race is an ongoing struggle between competing sets of co-evolving genes, phenotypic and behavioral traits that develop escalating adaptations and counter-adaptations against each other, resembling ...
, a form of
co-evolution
In biology, coevolution occurs when two or more species reciprocally affect each other's evolution through the process of natural selection. The term sometimes is used for two traits in the same species affecting each other's evolution, as well ...
, in which the fish constantly develops defenses against the parasite, while the parasite in turn evolves adaptations to cope with the specific defenses of its host. This tends to drive the
speciation
Speciation is the evolutionary process by which populations evolve to become distinct species. The biologist Orator F. Cook coined the term in 1906 for cladogenesis, the splitting of lineages, as opposed to anagenesis, phyletic evolution withi ...
of more specialized species provided conditions remain relatively stable. This involves
niche partitioning as new species are formed, and
biodiversity
Biodiversity or biological diversity is the variety and variability of life on Earth. Biodiversity is a measure of variation at the genetic ('' genetic variability''), species ('' species diversity''), and ecosystem ('' ecosystem diversity' ...
is increased.
A benefit of a specialist species is that because the species has a more clearly defined niche, this reduces competition from other species. On the other hand, generalist species, by their nature, cannot realize as much resources from one niche, but instead find resources from many. Because other species can also be generalists, there is more competition between species, reducing the amount of resources for all generalists in an ecosystem. Specialist herbivores can have morphological differences as compared to generalists that allow them to be more efficient at hunting a certain prey item, or able to eat a plant that generalists would be less tolerant of.
Summary
See also
*
Cosmopolitan distribution
*
Endemism
Endemism is the state of a species being found in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also found els ...
*
Fitness landscape
*
List of feeding behaviours
References
https://www.webpages.uidaho.edu/range556/appl_behave/projects/different_strokes.html
{{modelling ecosystems
Ecology