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__NOTOC__ The Roque Chico de Salmor giant lizard (''Gallotia simonyi simonyi'') is the
extinct Extinction is the termination of an organism by the death of its Endling, last member. A taxon may become Functional extinction, functionally extinct before the death of its last member if it loses the capacity to Reproduction, reproduce and ...
nominate subspecies In biological classification, subspecies (: subspecies) is a rank below species, used for populations that live in different areas and vary in size, shape, or other physical characteristics ( morphology), but that can successfully interbreed. ...
of Simony's giant lizard(''Gallotia simonyi''). It was
endemic Endemism is the state of a species being found only in a single defined geographic location, such as an island, state, nation, country or other defined zone; organisms that are indigenous to a place are not endemic to it if they are also foun ...
to the small islet Roque Chico de Salmor (lit. "Salmor Small Rock") northwest of
El Hierro El Hierro (), nicknamed ''Isla del Meridiano'' (the "Meridian Island"), is the farthest south and west of the Canary Islands (an autonomous community of Spain), in the Atlantic Ocean off the coast of Africa, with a population of 11,659 (2023). ...
, the westernmost of the
Canary Islands The Canary Islands (; ) or Canaries are an archipelago in the Atlantic Ocean and the southernmost Autonomous communities of Spain, Autonomous Community of Spain. They are located in the northwest of Africa, with the closest point to the cont ...
.


Description

Preserved male specimens measure between 223 and 236 mm long, excluding the tail, and females between 174 and 197 mm. Compared to the extant subspecies from the mainland ('' G. s. machadoi''), the subspecies from Roque Chico de Salmor was larger and more robust, with the top of the head more triangular and less oval, more depressed head, and on average, more dorsal scales, fewer temporal scales, more femoral pores, and more scales on the sixth ring of the tail.
Mitochondrial DNA Mitochondrial DNA (mtDNA and mDNA) is the DNA located in the mitochondrion, mitochondria organelles in a eukaryotic cell that converts chemical energy from food into adenosine triphosphate (ATP). Mitochondrial DNA is a small portion of the D ...
studies suggest that the two subspecies separated recently, as a result of rising sea levels at the end of the
Pleistocene The Pleistocene ( ; referred to colloquially as the ''ice age, Ice Age'') is the geological epoch (geology), epoch that lasted from to 11,700 years ago, spanning the Earth's most recent period of repeated glaciations. Before a change was fin ...
.


Distribution and habitat

Roque Chico de Salmor is a small islet situated 830
meter The metre (or meter in US spelling; symbol: m) is the base unit of length in the International System of Units (SI). Since 2019, the metre has been defined as the length of the path travelled by light in vacuum during a time interval of of ...
s to the northwest of El Hierro and 340 meters from the larger Roque Grande de Salmor.Salvador, A. (2015
Lagarto gigante de El Hierro – Gallotia simonyi
In: ''Enciclopedia Virtual de los Vertebrados Españoles.'' Salvador, A., Marco, A. (Eds.). Museo Nacional de Ciencias Naturales, Madrid. http://www.vertebradosibericos.org/
Its extension is less than one
hectare The hectare (; SI symbol: ha) is a non-SI metric unit of area equal to a square with 100-metre sides (1 hm2), that is, square metres (), and is primarily used in the measurement of land. There are 100 hectares in one square kilometre. ...
and is surrounded by tall cliffs. Vegetation is sparse but it is home to breeding colonies of
seabird Seabirds (also known as marine birds) are birds that are adaptation, adapted to life within the marine ecosystem, marine environment. While seabirds vary greatly in lifestyle, behaviour and physiology, they often exhibit striking convergent ...
s. Plants present include '' Astydamia latifoliae'', '' Mesembrianthemum'', ''
Chenopodium ''Chenopodium'' is a genus of numerous species of perennial or annual herbaceous flowering plants known as the goosefoot, which occur almost anywhere in the world. It is placed in the family Amaranthaceae in the APG II system; older classifica ...
'', '' Chenoloides tomentosa'', ''
Beta Beta (, ; uppercase , lowercase , or cursive ; or ) is the second letter of the Greek alphabet. In the system of Greek numerals, it has a value of 2. In Ancient Greek, beta represented the voiced bilabial plosive . In Modern Greek, it represe ...
'', ''
Silene ''Silene'' is a genus of flowering plants in the family Caryophyllaceae. Containing nearly 900 species, it is the largest genus in the family. Common names include campion and catchfly. Many ''Silene'' species are widely distributed, particularl ...
'' and '' Rubia fruticosa''.


Discovery

Despite being hunted by the
Guanches The Guanche were the Indigenous peoples, indigenous inhabitants of the Spain, Spanish Canary Islands, located in the Atlantic Ocean some to the west of modern Morocco and the North African coast. The islanders spoke the Guanche language, which i ...
, who also started clearing the island's forests and introduced exotic predators like
dog The dog (''Canis familiaris'' or ''Canis lupus familiaris'') is a domesticated descendant of the gray wolf. Also called the domestic dog, it was selectively bred from a population of wolves during the Late Pleistocene by hunter-gatherers. ...
s and possibly
cat The cat (''Felis catus''), also referred to as the domestic cat or house cat, is a small domesticated carnivorous mammal. It is the only domesticated species of the family Felidae. Advances in archaeology and genetics have shown that the ...
s, giant lizards were still common in areas of El Hierro when Europeans arrived at the beginning of the 15th century. They severely declined with
colonization 475px, Map of the year each country achieved List of sovereign states by date of formation, independence. Colonization (British English: colonisation) is a process of establishing occupation of or control over foreign territories or peoples f ...
, however, and by the 18th century were forgotten by the inhabitants of the island. In 1799,
José de Viera y Clavijo José de Viera y Clavijo (28 December 1731 – 21 February 1813), was a Spanish, of Portuguese descent, Enlightenment ecclesiastic, poet, historian, botanist, ethnographer, and professor. He is best known for his exhaustive ''History of the Can ...
wrote in his ''Diccionario de Historia Natural de las Islas Canarias'' that rumors existed of a small islet near El Hierro that was inhabited by lizards so large that they could bend iron bars with their tails; he presumed these animals to be
caiman A caiman ( (also spelled cayman) from Taíno language, Taíno ''kaiman'') is an alligatorid belonging to the subfamily Caimaninae, one of two primary lineages within the Alligatoridae family (biology), family, the other being alligators. ...
s. Following the publication of Viera y Clavijo's book in 1866,
Franz Steindachner Franz Steindachner (11 November 1834 in Vienna – 10 December 1919 in Vienna) was an Austrian zoologist, ichthyologist, and herpetologist. He published over 200 papers on fishes and over 50 papers on reptiles and amphibians. Steindachner des ...
of the
Naturhistorisches Museum The Natural History Museum Vienna () is a large natural history museum located in Vienna, Austria. The NHM Vienna is one of the largest museums and non-university research institutions in Austria and an important center of excellence for all matt ...
of
Vienna Vienna ( ; ; ) is the capital city, capital, List of largest cities in Austria, most populous city, and one of Federal states of Austria, nine federal states of Austria. It is Austria's primate city, with just over two million inhabitants. ...
requested his colleague Oskar Simony to investigate the matter during a scheduled trip to the Canary Islands. On 28 August 1889, Simony arrived in El Hierro and collected four lizards in Roque Chico de Salmor with the help of grocer Eloy Díaz Casañas. Steindachner used three of these lizards to describe a new species, ''
Lacerta Lacerta is one of the 88 modern constellations defined by the International Astronomical Union. Its name is Latin for lizard. A small, faint constellation, it was defined in 1687 by the astronomer Johannes Hevelius. Its brightest stars form a "W" ...
simonyi''.


Extinction

News of the discovery instantly caught the attention of naturalists, adventurers, and collectionists, who traveled to Roque Chico de Salmor to catch their own specimens. The most damaging was
wildlife trade Wildlife trade refers to the exchange of products derived from non-domesticated animals or plants usually extracted from their natural environment or raised under controlled conditions. It can involve the trade of living or dead individuals, tis ...
r Thomas Castle (born Tomás Ximénez del Castillo in
Puerto de la Cruz Puerto de la Cruz is a city and municipality in the northern part of the island of Tenerife, Canary Islands, Spain. It was formerly known by its English translation, "Port of the Cross", although now it is known by its Spanish name in all langu ...
) who, after convincing Díaz Casañas to gift him the fourth lizard collected with Simony and selling it for a high price, returned to capture 21 lizards in 1896 and 27 more in 1908, selling them in
London London is the Capital city, capital and List of urban areas in the United Kingdom, largest city of both England and the United Kingdom, with a population of in . London metropolitan area, Its wider metropolitan area is the largest in Wester ...
and
Hamburg Hamburg (, ; ), officially the Free and Hanseatic City of Hamburg,. is the List of cities in Germany by population, second-largest city in Germany after Berlin and List of cities in the European Union by population within city limits, 7th-lar ...
. This trade spurred Díaz Casañas, his son, nephew, and local fishermen to capture more lizards for sale or exhibition. In 1931, English teacher Hugh B. Scott and local journalist José Padrón Machín collected two lizards in Roque Chico de Salmor, but they were already so rare that they spent the first three days without seeing one. No more lizards were seen after this date, and a local legend arose that a British or German trader had captured twenty lizards and poisoned the rest to keep the exclusivity of their sale. Ten specimens are preserved in the Naturhistorisches Museum of Vienna,
British Museum of Natural History The Natural History Museum in London is a museum that exhibits a vast range of specimens from various segments of natural history. It is one of three major museums on Exhibition Road in South Kensington, the others being the Science Museum (Lo ...
, Santa Cruz de la Palma, and
Santa Cruz de Tenerife Santa Cruz de Tenerife (; locally ), commonly abbreviated as Santa Cruz, is a city, the capital of the island of Tenerife, Province of Santa Cruz de Tenerife, and one of the capitals of the Canary Islands, along with Las Palmas. Santa Cruz has a ...
. The species ''Gallotia simonyi'', then considered a
monotypic taxon In biology, a monotypic taxon is a taxonomic group (taxon) that contains only one immediately subordinate taxon. A monotypic species is one that does not include subspecies or smaller, infraspecific taxa. In the case of Genus, genera, the term ...
, was believed extinct until the subspecies ''G. s. machadoi'' was discovered in the Crag of Tibataje in 1975. In 1999, 36 members of this subspecies were introduced successfully to Roque Chico de Salmor, exhibiting low mortality and signs of reproduction.Salvador, A. (2015
Lagarto gigante de El Hierro – Gallotia simonyi
In: ''Enciclopedia Virtual de los Vertebrados Españoles.'' Salvador, A., Marco, A. (Eds.). Museo Nacional de Ciencias Naturales, Madrid. http://www.vertebradosibericos.org/


See also

* List of Macaronesian animals extinct in the Holocene


References


Further bibliography

* Blanco, Juan Carlos & González, José Luis (eds.) (1992):
Libro rojo de los vertebrados de España
'. ICONA, Madrid. * Böhme, W. & Bings, Werner (1975): Zur Frage des Überlebens von ''Lacerta s. simonyi'' Steindachner. ''Salamandra'' 11(1): 39–46. rticle in German* Diaz, Carlos Naeslund & Bischoff, Wolfgang (1994): Studien am Roque Chico de Salmor bei El Hierro (Kanaren): 1. Mögliche Ursachen für das Aussterben von ''Gallotia simonyi'', 2. Die Artzugehörigkeit seiner Geckos (''Tarentola''). ''Salamandra'' 30(4): 246–253. [Article in German
HTML abstract
* European Environment Agency (2006)
European Nature Information System (EUNIS)Species Factsheet: ''Gallotia simonyi simonyi''
Downloaded on 24 Feb 2007. * Maas, Peter H.J. (2006)
The Extinction Website
Extinctions in Europe. Downloaded on 18 May 2006. * Database entry includes a range map and justification for why this species is critically endangered


External links

* lacerta.de

. Contains several good photos of habitat on Roque Chico de Salmor. Retrieved 2007-FEB-25. {{Taxonbar, from=Q7366788 Gallotia Extinct reptiles Reptile extinctions since 1500 Lizards of Europe Reptiles of the Canary Islands El Hierro Taxa named by Franz Steindachner